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Paneroa, a New Genus of Eupatorieae (Asteraceae) from Mexico Edward E. Schilling Department of Ecology and Evolutionary Biology, University of Tennessee, 569 Dabney Hall, Knoxville, Tennessee 37996, U.S.A. [email protected] Abstract. The new genus Paneroa E. E. Schilling Arts 36: 476. 1901. TYPE: Mexico. Oaxaca: vie. (Asteraceae. Eupatorieae) from the Mix teen Alta of La Parada, 19 Aug. 1894, E. W. Nelson 991 region in the slate of Oaxaca, Mexico, is described (holotype, GH; isotypes, F not seen, K not seen, and illustrated, and a new combination, P. stachyo- US). Figure 1. folia (B. L. Robinson) E. E. Schilling, is made. The 4cgl-earnviubasbt eei ds scthpyahlerya lclaaterpirpeizes,en dds ahbgoye rsti. t s cMaolrtooelnrenicfaoutrlema,r seppsahspyilpleou gslee, anveaetnisdc, sintrciraetea,s inrge dddiisshta llgyr,e enL,r icsheormiceeso uosn, trsitcehmo maen dd eontshiteyr Aangaelryasteusm sLh.o, wi n thwaht ichth et heg esnpuesc ieiss wdaisst inocritg infraollmy vsiengueotautsi,v e0 .2p-a1r tsm ms. imLpelaev, es maultletircnealtleu latrh,r ousgohmoeuwt hoart described, and is closely related to other genera sometimes opposite proximally and alternate distally, iSisnc chclhuuladtrzian cBgt eiprioCzneotdinn oubcsyl.i nnIituu mmoce cruoDrusCs i.e nn ada enfmldo ircissFt. ilce irsecghimona ntnhiaat sswemisdaselilllyee r adocirus ttnaele latyor, l yero llusipnot,d iecbd, l,ba damseeasr go3ibn0t u-ss5oe0m toe Xwc huan1te1 ar-et2ev5,o almuptmeex,. IUCKNey Rweodr dLsi:s t, AMgeexraictuom, M, iAxtsetcear aAceltaae,, PaEnuepraotao.r ieae, scseuresrfsnaialcteee-s,g laapndadalexu ilaaglrr eslerunicr,f haodcmeenses se glyrne uemwn,he irtoseeu rsis ceearonicudes o:s ucsaa,bt taesxuriebad-l A survey of molecular variation for the ITS and ETS throughout the surface. Capitulescences terminal, of 7 rBsep.l aaLcti.e vrRe lroyeb gdiinoissnotsan n rtwe fvareosa mlne odtth etoh nraleyts tAd ogisfe trtihantecu tms ab musttp awlcehadys ospfpolealcicuieemds cblruasctteero; latpee; dhuenacdlse sd is1c-o7id5, mhmem, isdpehnesriecl,y erseecrti,c e8o-u1s1, oCfr aAzgieecrro,a ntuudnmupc uteLbd.l i s(hEtoe. d E o.d baSttaac)ihn. i lTlihnoegu ,t gsJruo.r uvLpe.y Poafn Aergoe r&at uBm. liXinn e28a -cr1 l0otos emlsymo m;o vreeewcrhleaapptt pasicnplgea tuseeplaraitleees,a , t4e5-.- r6iPb.h5by elXdla, r0iae.p5se- 03x 0.7a tcomu t3me5., comprehensive analysis of Conoclinium DC., and the SUrfaCe whlte pilose’ Wllh scaltered ^bsessilc glam result prompted a more detailed investigation of the dular lrichomes; flowers ca. 100 per head, corollas pchlalocerompelanstt of DAN. sAta chsyeqoufoelniucmes .u siTnhge a brerosuadltesr aorfr ayt hoef f1u.5n nmelmfo, rms,p acrsreelaym yp ilwoshei tew, it3h.2 —sc3a.5tt emremd, stuubbeses sscilae. tnrimsieov sotdeuelneeldtsec scsu,frs lri aaobo rmen fpd e Ahw m.hy oelsMotrraegpeciehxnhnoi.yce looatingfco i cglsiaetulunm dua,is ne wasP lhaycinosceenhsrf .oih ramaB dEea d.sa e Eltdrh.e eaSo dncdy his ittblhilneeinecs¬nge 0r10 o..77u nmmd1 emmd,,, wpagb ahopoiLui lcitls aoh.as , se1 awmonintmdh e ew mra indsae epi l oapntes gndu;dir sfastagatcylee lssee :ns dcaac, .nal o.t hb0e1e.1r1ss 0mm.cmm5a-.,, creamy white, glabrous, the branches ca. 5 mm, Paneroa E. E. Schilling, gen. nov. Basionym: papillose, conspicuously clavate. Cypselae black, Ageratum sect. Stachyofolium M. F. Johnson, 2.2-3 mm, prismatic, tapering to base, with 5 ribs, Ann. Missouri Bot. Gard. 58: 79. 1971. TYPE: glabrous, carpopodium not strongly differentiated; Paneroa stachyofolia (B. L. Robinson) E. E. pappus coroniform, whitish tan, 0.3-0.5 mm, margin Schilling. entire to obscurely dentate. Distribution, habitat, and IUCN Red List category. Paneroa stachyofolia is a rare species that occurs in open pine-oak woodlands in the Mixteca Alta region in the state of Oaxaca, Mexico, This area harbors Paneroa stachyofolia (B. L. Robinson) numerous endemics, including one endemic genus Schilling, comb. nov. Basionym: Ageratu (Ainea Ravenna), several near endemic genera chyofolium B. L Robinson, Proc. Amer. (Fosteria Molseed, Gibasoides D. R. Hunt, and Novon 18: 520-523. Published on 16 December 2008. doi: 10.3417/2007173 522 Hofmeisteria fasciculata Tamaulipa azurea Eupatorium hyssopifolium Eutrochium maculatum Carphephorus corymbosus Koanophyllon ayapanoides Chromolaena haenkeana Ageratum houstonianum >— Conoclinium coelestinum Fleischmannia incamata Critonia sexangularis Paneroa stachyofolia (cid:9668)— Volume 18, Number 4 Schilling 523 Paneroa (Asteraceae) from Mexico Gland, Switzerland, and Cambridge, United ^Schilling 2036, EU3^7049/EU3T7037/AY727174; Chro- Johnson, M. L. 1971. A monograph of the genus Agemtum L. EMUex3i3c7o0, 2J9a;l iscCoo, nJo. clLin iPuman ercoo e8le8s4li1n,u mEU 3(L3.7),0 5D2/CE.:U 3U37.S0.4A0.,/ Arkansas, E. E. Schilling 04-58, EU337056/EU337044/ EU337033; Critonia sexangularis (Klatt) R. M. King & H. Robinson: Mexico, Oaxaca, J. L. Panero 2970 (TEX), Bot Gard. 22: 1-581. f § 7 EU337051/EU337039/EU337028; Eupatorium hyssopifolium -PanAsoesurtoteh,r earc&Jne. aM-eL e.f xroi&c.mo . sJ Bo. u1rit9Ltht9.eo 7rn.nV i aNMi l4leea8wxs: ei c5nto6oa.xr 6.Ba- 5r7i1ot93tfo 9. n6\is.a | e4Ni8ao:c ve7ea9let-i e9s0f r.o imn ELEL.aUU: m33Uo33.n77St00.:A 34.58,U //AEF.SUlYo.A37ri32.d,7 7a01, N37K62e./;w E C EU. u3YSt3rioor7ric0kph,2u i6nu; mE 0.F 2l-meEEiasU.cc hPuml-Sa1cta5uhn7minl,l i iaEn( UgEin .3)c 39aE75m0,- 14aE67ta,./ (Walter) R. M. King & H. Robinson: U.S.A., Tennessee, E. E. Lundellia 1: 72-74. Schilling 95-21, EU337055/EU337043/EU337032; Hofineis- teria fasciculata (Bentham) Walpers: Mexico, Baja California Sur, J. L. Panero 2817, EU337046/AF384731/EU337025; K [l II / d (G ebach) R. M. King & H. Appendix 1. List of vouchers for chloroplast DNA sequence Robinson, Cuba, Holguin: Bennet 7582, EU337050/ postbhAer-wtrinsHe . noted. Gen Bank numbers listed as matK/ndhFI EsEoUUn)33 33E77.00 35E87. //EESUUch33i33ll77in00g24:75 ;/M EPUeax3ni3ceo7r,o0 3aO 4as; txaaTccahary,r omJfu.o lliiLpa.a (PBaa.zn ueLrre. oaR 4o(D4b5iCn3.¬), R. M. King & H. Robinson: U.S.A., Texas, E. E. Schilling 1146, EU337053/EU337041/EU337030.