ANTARCTIC AmericanG eophysicaUl nion RESEARCH SERIES Antarctic Research Series Volumes 28 Biology of the Antarctic SeasV III D. L. Pawson, L. S. Korniker (Eds.) 29 Upper AtmosphereR esearchi n Antarctica L.d. 1 Biology of the Antarctic SeasI Milton O. Lee Lanzerotti,C . G Park (Eds.) 30 Terrestrial BiologyI I B. Parker (Ed.) 2 Antarctic Snow and Ice StudiesM . Mellor (Ed.) 31 Biology of the Antarctic SeasI X L. S. Kornicker 3 Polychaeta Errantia of Antarctica O. Hartman 32 Biology of the Antarctic SeasX L. S. Kornicker 4 Geomagnetisma nd Aeronomy A. H. Waynick 33 Dry Valley Drilling Project L. D. McGinnis( Ed.) 5 Biologyo f the Antarctic SeasI I GA. Llano (Ed.) 34 Biology of the Antarctic SeasX 1 L. S. Korniker 6 Geologya nd Paleontologyo f the Antarctic J.B. Hadley (Ed.) 35 Biology of the Antarctic SeasX II D. Pawson 7 Polychaeta Myzostomidae and Sedentaria of Antarctica O. Hartman (Ed.) 36 Geology of the Central Transantarctic 8 Antarctic Soils and Soil Forming Processes (cid:127)/. Mountains M.D. Turnerd, . E Splettstoesse(Er ds.) C. E Tedrow( Ed.) 37 Terrestrial BiologyI II B. Parker (Ed.) 9 Studiesi n Antarctic Meteorology M.d. Rubin 38 Biology of the Antarctic SeasX III [crinoids, hydrozoa,c opepodsa, mphipoda] L. S. Korniker 10 Entomologyo f Antarctica d. L. Gressit( Ed.) ((cid:127)a.) 11 Biology of the Antarctic SeasI II GA. Llano, W. 39 Biology of the Antarctic SeasX IV L. S. L. Schmitt( Eds.) Kornicker( Ed.) 12 Antarctic Bird StudiesO . L. Austin,J r. (Ed.) 40 Biologyo f the Antarctic SeasX V L. S. Korniker 13 Antarctic Ascidiacea P Kott (Ed.) 14 Antarctic Cirripedia W.A . Newman,A . Ross 41 Biologyo f the Antarctic SeasX VI L. S. Korniker 15 Antarctic OceanologyI L. Reid (Ed.) 42 The RossI ce Shelf: Glaciologya nd Geophysics 16 Antarctic Snow and Ice StudiesI I A. P Crary C. R. Bentley,D . E. Hayes (Eds.) 43 Oceanologyo f the Antarctic Continental Shelf S. 17 Biology of the Antarctic SeasI V GA. Llano, L Jacobs( Ed.) E. Wall en (Eds.) 44 Biology of the Antarctic SeasX VII [benthic sati- 18 Antarctic Pinnipedia W.H . Butt (Ed.) ation, brittle star feeding, pelagic shrimps, 19 Antarctic OceanologyI I: The Australian-New marine birds] L. S. Korniker (Ed.) Zealand Sector D. E. Hayes (Ed.) 45 Biology of the Antarctic SeasX VIII, Crustacea 20 Antarctic Terrestrial Biology GA. Llano (Ed.) Tanaidacea of the Antarctic and the 21 Recent Antarctic and SubantarcticB rachiopods Subantarctic 1. On Material Collected at Tierra M. W.F oster (Ed.) del Fuego, Isla de los Estados, and the West 22 Human Adaptability to Antarctic Conditions E. Coast of the Antarctic Peninsula L. S. Korniker K. Eric Gunderson(E d.) 23 Biology of the Antarctic SeasV D. L. Pawson 46 GeologicalI nvestigationsi n Northern Victoria Land E. Stump( Ed.) 24 Birds of the Antarctic and Sub-Antarctic G E. 47 Biology of the Antarctic SeasX IX [copepods, Watson( Ed.) teleosts] L. S. Korniker (Ed.) 25 Meteorological Studiesa t Plateau Station, 48 Volcanoes of the Antarctic Plate and Southern Antarctica (cid:127) Businger( Ed.) Oceans W.E . LeMasurier(cid:127)1, . W. Thomson(E ds.) 26 Biology of the Antarctic SeasV I D. L. Pawson 49 Biology of the Antarctic SeasX X, Antarctic SiphonophoresF rom Plankton Sampleso f the 27 Biology of the Antarctic SeasV II D. L. Pawson United StatesA ntarctic Research Program L. S. Kornicker( Ed.) 5O Contributions to Antarctic Research I D.H. 64 Atmospheric Halos W. Tape Elliot (Ed.) 65 Fossil Scleractinian Corals From James Ross 51 Mineral Resources Potential of Antarctica J.F. Basin, Antarctica H. E Filkorn SplettstoesseGr, ,4. M. Dreschhoff( Eds.) 66 Volcanologicala nd Environmental Studieso f 52 Biology of the Antarctic SeasX XI [annelids, Mt. Erebus P. R. Kyle (Ed.) mites, leechesl L. S. Korniker (Ed.) 67 Contributions to Antarctic Research IV D. H. 53 Contributions to Antarctic Research II D.H. Elliot, G L. Blaisdell( Eds.) Elliot (Ed.) 68 Geologya nd SeismicS tratigraphy of the 54 Marine Geological and GeophysicalA tlas of the Antarctic Margin ,4. K. Cooper,P E Barker, G Circum-Antarctic to 30ES D. E. Hayes (Ed.) Brancolini (Eds.) 55 Molluscan Systematicsa nd Biostratigraphy 69 Biology of the Antarctic SeasX XIV, Antarctic Lower Tertiary La Meseta Formation, Seymour and Subantarctic Pycnogonida' Nymphonidae, Island, Antarctic Peninsula J. D. Stilwell, W.J . Colossendeidae,R hynchothoraxida, Zinsmeister Pycnogonidae,P hoxichilidiidae, Endeididae, 56 The Antarctic Paleoenvironment:A Perspective and Callipallenidae S. D. Cairns (Ed.) on Global Change, Part One J. P Kennett,D .A. 70 Foundations for EcologicalR esearchW est of the Warnke( Eds.) Antarctic Peninsula R. M. Ross,E . E. Hofmann, 57 Contributions to Antarctic Research III D. H. L. B. Quetin( Eds.) Elliot (Ed.) 71 Geology and SeismicS tratigraphy of the 58 Biology of the Antarctic Seasx (cid:127)Xll 3. V. Cairns Antarctic Margin, Part 2 F. p. t(cid:127)'arkerA, . K. Cooper( Eds.) 59 Physicaal nd BiogeochemicParl ocesseins 72 EcosystemD ynamics in a Polar Desert: The Antarctic Lakes W.J. Green, E. L Friedmann McMurdo Dry Valleys,A ntarctica John C. (Eds.) Priscu (Ed) 60 TheA ntarcticP aleoenvironmenAt :P erspective 73 Antarctic Sea Ice: Biological Processes, on Global Change, Part Two J. P Kennett,D . ,4. Interactions and Variability Michael P Lizotte, Warnke( Eds.) Kevin R. ,4frigo (Eds.) . 61 AntarcticM eteorologya nd ClimatologyS: tudies 74 Antarctic Sea Ice: PhysicalP rocesses, Based on Automatic Weather Stations D.H. Interactions and Variability Martin O. Jeffries Bromwich,C . R. Steams( Eds.) ((cid:127)a.) 62 Ultraviolet Radiation in Antarctica: 75 Ocean, Ice and Atmosphere: Interactions at the Measurements and Biological Effects C.S. Continental Margin StanleyS . Jacobs,R ay E WeilerP, ,4. Penhale( Eds.) Weiss(E ds.) 63 Biology of the Antarctic SeasX XIV, Antarctic and Subantarctic Pycnogonida:A mmotheidae and Austrodecidae S. D. Cairns (Ed.) THE ANTARCTIC RESEARCH SERIES The Antarctic Research Series, published since 1963 by the American GeophysicaUl nion, now comprisesm ore than 70 volumeso f authoritativeo rigi- nal resultso f scientificw ork in the high latitudeso f the southernh emisphere. Series volumes are typically thematic, concentratingo n a particulart opic or region,a nd may containm apsa nd lengthyp apersw ith large volumeso f data in tabular or digital format.A rearctic studiesa re often imerdisciplinaryo r interna- tional, and build upon earlier observationsto addressis sueso f naturalv ariability and global change.T he standardso f scientifice xcellencee xpectedf or the Series are maintainedb y editorsf ollowingr eview criteriae stablishedfo r the AGU pub- licationsp rogram.P riorities for publicationa re set by the Board of Associate Editors. Inquiries aboutp ublishedv olumes,w ork in progresso r new proposals may be sent to Antarctic Research Series, AGU, 2000 Florida Avenue NW, WashingtonD, C 20009 (http://wwwa. gu.org),o r to a membero f the Board. BOARD OF ASSOCIATE EDITORS RodneyM . Feldmann,C hairman,P aleontology RobertA . BindschadlerG, laciology David H. Bromwich,M eteorologya nd UpperA tmosphereP hysics Nelia W. Dunbar,G eology StanleyS . Jacobs,O ceanography Jerry D. Kudenov,M arine/PolychaeteB iology JohnC . Priscu,T errestrialB iology Mount Discovery,M cMurdo Sound,m orained epositsw hich containt he most important record,p reservedin glaciale rratics,o f Paleogeneli fe and paleoenvironmenotsf this inter- val from East Antarctica. The rich suite of fossiliferous erratics and various lithofacies recoveredp rovidest he groundworkfo r reconstructinEgo ceneh igh-latituden earshoree nvi- ronmentsa nd planktic/benthicc ommunitiesp reservedin the erratics.T he implicationsf or EastA ntarcticp aleoclimatea ndp aleoceanographpyri ort o the onseto f significangt laciation in Antarctica,a re far reachinga nd are of major interestt o the global scientificc ommunity. Photographta kenb y Dr. JeffreyD . Stilwell,J anuary3 , 1993. ANTARCTIC l/olume 76 RESEARCH SERIES Paleobiologya nd Paleoenvironmentosf EoceneR ocks, McMurdo Sound, East Antarctica JeffreyD . Stilwell and RodneyM . Feldmann Editors c(cid:127) AmericGane ophysUicnaiol n WashingtonD, .C. 2000 PALEOBIOLOGY AND PALEOENVIRONMENTS OF EOCENE ROCKS, MCMURDO SOUND, EAST ANTARCTICA JeffreyD . Stilwell and RodneyM . Feldmann,E ditors Publishedu ndert he aegiso f the Board of AssociateD irectors,A ntarcticR esearchS eries Library of CongressC ataloging-in-Publication Data Paleobiologya nd paleoenvironmentosf Eocener ocks,M cMurdo Sound,E astA ntarctica / JeffreyD . Stilwell and RodneyM . Feldmann,e ditors p.cm -- (Antarcticr esearchs eries; v. 76) Includesb ibliographicarle ferences. ISBN 0-87590-947-7 1. Paleontology--Eocene2.. Fossils--Antarctica--McMurdoS oundR egion. 3. Boulders--Antarctica--McMurdSoo undR egion.4 . Geology,S tratigraphic--Eocen5e.. Geology--Antarctica--McMurdSo oundR egion. I. Stilwell, JeffreyD . II. Feldmann, RodneyM . III. Series QE737.P35 2000 560'. 1784--dc21 00-20726 CIP ISBN 0-87590-947-7 ISSN 0066-4634 Cover Coastalg lacialm orained epositsa t Mount DiscoveryM, cMurdo Sound,w herea wealtho f fossiliferouse rraticso f predomi- nantlyE ocenea geh aveb eenr ecovered.M any importantr ock and fossils pecimenws ere discoveredin the depositss hown here.N ote Ice Pinnaclesa ndB lack Islandi n backgroundP. hotographta kenb y JeffreyD . Stilwell,N ovember1 1, 1995. Copyright2 000 by the AmericanG eophysicaUl nion 2000 Florida Avenue, N.W. Washington,D C 20009 Figures,t ables,a nd shorte xcerptsm ay be reprintedin scientificb ooksa ndj ournalsi f the sourceis properlyc ited. Authorizationt o photocopyit emsf or internalo r personalu se,o r the internalo r personaul seo f specificc lients,i s grantedb y theA mericanG eophysicaUl nion for librariesa ndo theru sersr egisteredw ith the CopyrightC learanceC enter( CCC) Transactional ReportingS ervicep, rovidedt hatt he basef ee of $01.50p er copyp lus$ 0.50 per pagei s paidd irectlyt o CCC, 222 RosewoodD r., Danvers, MA 01923. 0066-4634/00/$01.50+0.50. This consentd oesn ot extendt o otherk indso f copying,s ucha s copyingf or creatingn ew collectivew orkso r for resale.T he reproductiono f multiplec opiesa ndt he useo f full articleso r the useo f extractsi,n cludingf iguresa ndt ables,f or commerciapl ur- posesr equiresp ermissionfr om the AmericanG eophysicaUl nion. Publishedb y AmericanG eophysicaUl nion 2000 Florida Avenue, N.W. WashingtonD, .C. 20009 With the aid of grantO PP-9414962 from the National Science Foundation Printed in the United States of America. CONTENTS Preface JeffreyD . Stilwell and RodneyM . Feldmann xi The McMurdo Erratics: Introduction and Overview David M. Harwood and RichardH . Levy Glacial Geologya nd Origin of Fossiliferous-Erratic-BearinMgo raines,S outhernM cMurdo Sound,A ntarctica: An AlternativeI ce SheetH ypothesis Gary S. Wilson 19 SedimentaryL ithofacieso f the McMurdo SoundE rratics RichardH . Levy and David M. Harwood 39 Clay Mineral Compositiono f Glacial Erratics,M cMurdo Sound Mary AnneH olmes 63 Marine DiatomA ssemblagefsr om Eocenea ndY oungerE rratics,M cMurdo Sound,A ntarctica David M. Harwood and StevenM . Bohaty 73 Ebridiana nd SilicoflagellateB iostratigraphyfr om EoceneM cMurdo Erraticsa nd the SouthernO cean StevenM . Bohaty and David M. Harwood 99 Sporesa nd Pollen from the McMurdo SoundE rratics,A ntarctica RosemaryA . Askin 161 TertiaryM arine Palynomorphsfr om the McMurdo SoundE rratics,A ntarctica RichardH . Levy and David M. Harwood 183 Eocene Plant Macrofossils from Erratics, McMurdo Sound,A ntarctica Mike Pole, Bob Hill and David M. Harwood 243 FossilW ood from EoceneH igh LatitudeF orestsM cMurdo Sound,A ntarctica Jane E. Francis 253 EoceneM ollusca( Bivalvia, Gastropodaa nd Scaphopodaf)r om McMurdo Sound:S ystematicasn d PaleoecologicS ignificance JeffreyD . Stilwell 261 BryozoanF ragmentsf rom EoceneG lacial Erraticso f McMurdo Sound,E astA ntarctica Urszula Hara 321 RhynchonellideB rachiopodsfr om Eocenet o EarliestO ligoceneE rraticsi n the McMurdo SoundR egion,A ntarctica DaphneE . Lee and JeffreyD . St#well 325 A New Specieso f Austrobalanus(C irripedia,T horacicia)f rom EoceneE rratics,M ount Discovery,M cMurdo Sound,E ast Antarctica JohnS t. J. S. Buckeridge 329 Callichirus?s ymmetricu(sD ecapoda:T halassinoideaa)n dA ssociatedB urrows,E ocene,A ntarctica Carrie E. Schweitzera nd RodneyM . Feldmann 335 Fish Remainsf rom the Eoceneo f Mount Discovery,E astA ntarctica DouglasH . Long and JeffreyD . Stilwell 349 A ProbableP iscivorousC rocodilef rom EoceneD epositso f McMurdo Sound,E astA ntarctica Paul M.A. Willis and JeffreyD . Stilwell 355 The First Record of a Fossil Bird from East Antarctica Craig M. Jones 359 PaleobiogeographiScy nthesiso f the EoceneM acrofaunaf rom McMurdo Sound,A ntarctica JeffreyD . Stilwella nd WilliamJ . Zinsmeister 365 PREFACE Michael K. Brett-Surman, George Washington sions drawn about the climate of the continent from University,o bservedth at, "beinga paleontologisits like researchc onductedo n SeymourI sland. As a result, the beinga coronere xcepta ll the witnessesa re deada nd all studyo f thesee rraticsh as strengthenedo ur understand- the evidence has been left out in the rain for 65 million ing of the conditionsth at prevailedi n the high southern years." In the study of paleontologyi n Antarcticai t latitudesju st prior to final separationo f Antarcticaf rom could also be added that, if not left out in the rain, most the Australian continent and establishment of the isola- of the evidence remains buried beneath several thousand tion of the southernmost continent. feet of ice. Elucidating the geologic history of the Becauseo f the wide range of fossilsk nown from the Antarctic continentw ill always be plagued with this erratics,n umerouss pecialistsw ere recruitedt o studyt he problem. Nonetheless,n umerousc lever means have specimensT. he resulti s a collectiono f highly authorita- beenu sedt o extracta s muchi nformationa s is possible, tive articlesp rovidinga benchmarkf or further work in and as presentedin this volume. the area.A s with many suchs tudies,t his work may be In this light, one of the most intriguingt ime intervals regardeda s preliminary,w ith a next step undoubtedly in Antarctic history is the Eocene Epoch. During this requiringt he serendipitoudsi scoveryo f a new site. time, the climatic conditionsd eterioratedr apidly from We thank all the contributors to the volume for their the so-called "Greenhouse" conditions that dominated effortsi n bringingt he work to completionI.n additiont o Earth'sc onditionsfr om mid-Mesozoict ime throught he the authors,a largec adreo f reviewersr ead the contribu- early Cenozoic to the "Icehouse"c onditionst hat have tionsa nd providedv aluables uggestionsT.h e contento f dominatedt he climate sincet hat time. Unfortunately,t he thep apersi s the responsibilitoy f the authorsa nde ditors; recordo f Eocener ockso n the continenits sparseO. n the the quality of the final productw as much enhancedb y AntarcticP eninsula,s pecificallyo n SeymourI sland, a the reviewers and we thank them. Those who chose to be robustr ecordo f Eocener ocks and fossilsh asp rovided identified are acknowledgedin the individual articles. virtually all the informationw e possessa boutt his time Finally, the transformationf rom our idea of "camera- interval.T hus the discoverya nd descriptiono f Eocene ready" copy to the finishedp roduct was the task of erratic bouldersi n morainal depositsi n the McMurdo Karen Smith, Department of Geology, Kent State Soundr egionp rovideso nly the seconds iteo n the entire University,w ho did a magnificenjto b. Financials upport continentw here we can studyt he paleontologyo f this for the work was largely throught he National Science time interval.I n all likelihood,t he descriptiono f erratics FoundationA. s with all projectso f this type, the volume containingf ossils from any other place in the world of work necessaryto completet he taski s alwaysu nder- would warrant little study and would attract even less estimated.W e beg your indulgence.H owever, as with attention. However, when most of the vast area of fine wine,.... Antarctica lies beneath ice and when clues to the nature of the crusto f that part of the continentc anb e extracted JeffreyD . Stilwell only from studyo f erratics,t he discoveryc arriesw ith it JamesC ook University some excitement. Townsville,Q ueenslandA, ustralia The studyo f fossiliferouse rraticsg rowsm ore signifi- cant when they are found to containa diversea rray of RodneyM . Feldmann vertebratei,n vertebratea, ndp lantm aterial,w hichm akes Kent StateU niversity it possiblet o interprett he Eocene climatic and paleo- Kent, Ohio ceanographics ettingo f the McMurdo Soundr egion in some detail. Furthermore, the work reinforces conclu- Editors
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