Table Of Content

1991. TheJournal ofArachnology 19:102-104 ON THE SPIDER GENUS FEDOTOVIA (ARANEAE, GNAPHOSIDAE) Vladimir 1. Ovtsharenko; Zoological Institute, Academy of Sciences, Universitetskaja emb. Leningrad 199034 USSR 1, NormanI.Platnick: DepartmentofEntomology, AmericanMuseum ofNaturalHistory, Central Park West at 79th Street, New York, New York 10024 USA Abstract. Fedotovia Charitonov is removed from the synonymy ofthe laroniinegenus Eilica and placed as a valid member of the subfamily Gnaphosinae. Males of the type species, F. uzbekistanica Charitonov, are described for the first time, and the species is newly recorded from Kazakhstan and Mongolia. ThespidergenusFedotoviawasestablishedby only Baeriella Simon, Caridrassus Bryant, Gna- Charitonov (1946) for a single female from Ish- phosoides Hogg, Gytha Keyserling, and Laronia kent, Uzbekistan, describedasF. uzbekistanica. Simon) returns the distribution of Eilica to a In hisoriginaldescription, Charitonovprovided morenormalGondwananpattern, includingonly no comments on the possible relationships or speciesfrom Africa, India,Australia, and south- subfamilial placement of Fedotovia, but later ern parts ofthe Americas. (Charitonov 1969) he placed the genus in the The format ofthe descriptions and abbrevia- “Gnaphoseae,” comparing it with Asemesthes tions used follow those ofPlatnick and Shadab Simonand GnaphosoidesHogg. Thelattername (1975); all measurements are in mm. was synonymized with the laroniine gnaphosid genusEilicaKeyserlingbyPlatnick(1975); Plat- Fedotovia Charitonov nickandShadab(1981)subsequentlyplacedFe- dotoviaasasynonymofEilicaaswell. Thatsyn- Fedotovia Charitonov, 1946:24 (type species by orig- onymy, however, was not based on an inal designation Fedotovia uzbekistanica Charito- examination ofthe type specimen, but only on nov, 1946); Charitonov, 1969:98. Eilica: Platnick and Shadab, 1981:184 (in part). figures of the eye pattern and epigynum of F. uzbekistanica provided by Charitonov (1946). Diagnosis.—SpecimensofFedotoviaresemble We have recently had the opportunity to ex- those of the African gnaphosine genus Ase- amine Charitonov’s type, as well as newly col- mesthes in having a strongly recurved posterior lectedspecimens,whichindicatethatdespitethe eye row, andmedian eyesthatare much smaller numerous strikingepigynal similarities with Ei- than the lateral pairs, and will therefore key out lica (including the shape ofthe lateral epigynal toAsemesthesinthekeybyDalmas(1921).Males margins,thetwistedposteriorepigynalducts,and can be distinguished from those ofAsemesthes the coiled median epigynal ducts), Fedotovia is by the shorter tibial apophysis and the greatly neitherasynonymofEilicanoramemberofthe elongated embolus (Figs. 1-3), females by the Laroniinae. It lacks the retromarginal cheliceral median epigynal scape (Fig. 4). — laminae characteristic ofthat subfamilyand has Description.—Total length 5 7. Carapaceal- instead the retromarginal serrated keel charac- most triangular in dorsal view, widest between teristic of the subfamily Gnaphosinae. Males, coxae II and III, flattened, smoothly narrowed newly described here, have an elaborate palpal opposite palpal insertions, light brown with embolus unlike those of previously described darkenedposteriormarginandrowsofstiffsetae gnaphosinegenera,andwethereforeconsider radiatingfromthoracicgroove.Cephalicareanot dotovia a valid genus ofthe Gnaphosinae. elevated, thoracic groove short, straight, longi- The removal ofFedotovia from the list ofge- tudinal. From front, anterior eye row straight, neric synonyms ofEilica (which thus includes posteriorrowrecurved;fromabove,anteriorrow 102 OVTSHARENKO& PLATNICK-F££><9rOF/^ (GNAPHOSIDAE) 103 Figures 1-5.—Fedotovia uzbekistanica Charitonov; 1, left male palp, prolateral view; 2 same, ventral view; 3, same, retrolateralview; 4, epigynum, ventral view; 5, same, dorsal view. slightly recurved, posterior row strongly re- outpreeningcombs;trochantersunnotched.Ab- curved. PME flattened, irregularly triangular, domen grayish brown, males with small, shiny, AME circular, ALE and PLE oval; PME and triangular anterior scutum. Six spinnerets; an- AME subequal, much smaller than lateral eyes; terior laterals with six or seven long piriform PME separated by less than their diameter, by gland spigots set posteriorly from majorampul- more than their diameter from PLE; AME sep- late gland spigots; posterior medians short, tu- aratedbyabouttheirdiameter,aboutasfarfrom bular in males but widened and reflexed ante- MOQ ALE; slightlywiderin backthan in front, riorly in females. Male palp with short tibial longer than wide. Clypeal height roughly equal apophysisshifteddorsally, strong, distallyhook- to ALE width. Chelicerae with serrated keel on shaped median apophysis, and elaborately coil- retromarginbearingfourorfiveteeth;promargin ingembolus. Epigynumwithpairoflongitudinal with two closely setteeth. Endites obliquely de- lateralmarginsandmedianscape;epigynalducts pressed, distally rounded but not as convergent twisted, transversely oriented along posterior as in Gnaphosa, with anterolateral serrula. La- epigynal margin, in large circularcoilsmedially. biumlong,extendingtwo-thirdsofenditelength. Distribution.—Knownonlyfromaridhabitats Sternum lightbrown with darkened, rebordered in the USSR (Uzbekistan, Kazakhstan) and margins and tiny sclerotized extensions to and Mongolia. between coxae. Leg formula 4123. Legs light brown,bearingnumerousspines. Tarsiwithtwo Fedotovia uzbekistanica Charitonov elongateclawsdentateonlyattheirbase,without Figs. 1-5 claw tufts; tarsi and distal portions ofmetatarsi FedotoviauzbekistanicaChanionov, 1946:24,figs. 31, IandIIwiththick,darkscopula;metatarsiwith- 32(femaleholotypefromIshkent,Uzbekistan,USSR, 104 THEJOURNALOFARACHNOLOGY inUniversityofPerm,examined);Charitonov, 1969: tion, 15-28 May 1989 (A. A. Raikhanov, S. I. 98. Ibraev), 2 males, 6 females. Kzil-Orda: Dzhu- Diagnosis.—With the characters ofthe genus samly, Karmakchinskii region, clay desert, 19 and genitalia as in Figures 1-5. June 1989 (A. A. Zyuzin), 1 female. Uzbekistan; Male.—Totallength6.11.Carapace2.81 long, Yakkabag: Ishkent, elev. 1100-1300 m, 26 June i2n.t1e8rdwiisdtean.ceFse:mAuMrEII0.20.51,0AlLoEng.0.1E4y,ePsMizEes0.a1n2d, v1e9r4s2it(yDo.fMP.erFme)d.oMtoOvN),GO1 LfeImAal:eB(haoylaontyKpeh.onUgnio-r P1-LE 0.17; AME-AME 0.12, AME-ALE 0.05, Aimak; Ekhingol, 30 Aug. 1977 (K. Monkhbay- PMMOEQ-PME0.06,PME-PLE0.11,ALE-PLE0.13; ar), 1 female. length 0.33, front width 0.22, back width ACKNOWLEDGMENTS 0.30. Legspination(onlysurfacesbearingspines listed): femora: I, II dl-1-0, pO-M; III, IVdl- Theworkreportedonherewasdonewhilethe 1-0, pO-M, rO-1-1; patellae III, IV pO-1-0, rO- second author was a guest ofthe Zoological In- stitute, Akademia Nauk, Leningrad, and that 0-00,; tpi2b-ila-e:l,IVv22--22--22,;rI2I-lp-Ol-;0-I1V, vdllr--02--12,;pI2I-Ild-ll-, support is greatly appreciated. We thank A. S. Utochkin and N. M. Pakhorukov of the Uni- v2-2-2, r2-l-l; metatarsi: I, II v2-0-0; III dl-0- 0, pl-2-2, V2-2-2, rl-1-2; IVdl-0-0, pl-2-2, v2- vMe.rsUi.tySohfaPdaebrmofftohreaAccmeesrsitcoantyMpeusmaetuermiaolf,Naantd- lp-2, r1-2-2. Palpalembolusoriginatingbasally, e2-xtending over retrolateral surface ofcymbium, ural History for assistance with illustrations. curling back to ventral surface ofbulb distally LITERATURECITED (Figs. 1-3). Charitonov,D. E. 1946. Newformsofspidersofthe Female.—Total length 5.67. Carapace 2.44 USSR. Izv. Est.-Nauchn. Inst. Molotovsk. Univ., long, 2.03 wide. Femur II 1.75 long. Eye sizes 12:19-32. and interdistances: AME 0.08, ALE 0.17, PME Charitonov, D. E. 1969. Material’i k faune paukov 0.12, PLE 0.13; AME-AME 0.10, AME-ALE SSR.Uchel.Zap.Permsk.Ord.Trudy.ICrasn.Znam. 0.04,PME-PME0.03,PME-PLE0.13,ALE-PLE Gos. Univ. Ime A. M. Gorkogo, 179:59-133. 0.09; MOQ length 0.30, front width 0.26, back Dalmas, R. de. 1921. Monographicdesaraigneesde width 0.27. Leg spination as in male, except as lasectiondesPterotricha(Aran.Gnaphosidae).Ann. InIoItepdi:-1p-a1t;elIlVaedlI-II0,-0I;VmpeOt-a0t-a0r;sutsibIiIaIe:dOI-I0p-O0-,0p-O0-; PlaStonci.ck,EnNt.omIo.l.19F7r5a.nceA, r8e9v:i2s3i3o-n32o8f.the spidergenus Eilica(Araneae, Gnaphosidae). Amer. Mus. Novi- 2. Medianepigynalscapesituatedbetweenlat- tates, 2578:1-19. eral epigynal margins (Fig. 4); epigynal ducts ir- Platnick, N. 1. & M. U. Shadab. 1975. Arevision of regularlytwistedposteriorly,coiledmedially(Fig. thespidergenusGnaphosa(Araneae,Gnaphosidae) 5). inAmerica. Bull. Amer. Mus.Nat. Hist., 155:1-66. Specimens examined (in Zoological Institute, Platnick,N. I. & M. U. Shadab. 1981. Onthespider Leningrad,unlessotherwiseindicated).—USSR: genus Eilica (Araneae, Gnaphosidae). Bull. Amer. Kazakhstan: Chimkentskaya: Aksumbe, Kara- Mus. Nat. Hist., 170:183-188. tau Mountains, Suzakskii region, 16 June 1989 ManuscriptreceivedAugust1990, revisedOctober1990. (A. A. Zyuzin), 5 females. Gurievskaya: Baskar- gan Steppe, Ustyurt Plateau, Ustyurt Reserva-