On spiders from the tundra zone of the Kola Peninsula, Russia (Arachnida: Aranei)

2010·0.2 MB·

by  A V Tanasevitch

Arthropoda Selecta 19(1): 41–56 © ARTHROPODA SELECTA, 2010
(cid:1)(cid:2)(cid:3)(cid:4)(cid:5)(cid:6)(cid:7)(cid:8)(cid:9)(cid:4)(cid:3)(cid:10)(cid:9)(cid:11)(cid:12)(cid:3)(cid:13)(cid:14)(cid:8)(cid:3)(cid:13)(cid:15)(cid:2)(cid:7)(cid:9)(cid:16)(cid:3)(cid:17)(cid:11)(cid:2)(cid:8)(cid:3)(cid:11)(cid:10)(cid:3)(cid:13)(cid:14)(cid:8)(cid:3)(cid:18)(cid:11)(cid:19)(cid:16)(cid:3)(cid:20)(cid:8)(cid:2)(cid:6)(cid:2)(cid:4)(cid:15)(cid:19)(cid:16)(cid:21)(cid:3)(cid:22)(cid:15)(cid:4)(cid:4)(cid:6)(cid:16)
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(cid:28)(cid:3)(cid:29)(cid:30)(cid:31) (cid:30)!(cid:3)"(cid:31)#$%&’&((cid:3))&#*(cid:3)+&,-. &/&(cid:3)(cid:29)&,(cid:31)&."%&’(cid:30)
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;1<1(cid:3)=(cid:30)#(cid:30).>’?@4(cid:21)(cid:3)A1B1(cid:3)C*D(cid:30),&’:
1Centre for Forest Ecology and Production, Russian Academy of Sciences, Profsoyuznaya Str. 84/32, Moscow 117997, Russia. E-mail:
[email protected].
H(cid:12)(cid:14)(cid:17)(cid:9)(cid:7)(cid:21)(cid:16)(cid:7)(cid:21)(cid:9)(cid:16)#(cid:29)(cid:12)/(cid:15)/(cid:7)I(cid:22)(cid:16)(cid:29)(cid:16)(cid:31)(cid:10)(cid:10)(cid:7)(cid:10)(cid:7)(cid:21)(cid:9)(cid:16)(cid:13)(cid:26)(cid:22)(cid:17)(cid:10)(cid:11)(cid:14)(cid:16)(cid:20)(cid:17)(cid:10)(cid:7)(cid:29)(cid:12)(cid:20)(cid:16)(cid:11)(cid:7)(cid:1)JK!(cid:7)(cid:8)(cid:9)(cid:16)2(cid:20)(cid:16)L(cid:27)(cid:14)(cid:15)1(cid:7)(cid:26)(cid:29)(cid:6)(cid:7)CMN7O!(cid:7)(cid:5)(cid:16)(cid:20)(cid:22)(cid:11)(cid:15)(cid:7)(cid:23)(cid:23)P--P!(cid:7)(cid:1)(cid:16)(cid:20)(cid:20)(cid:10)1(cid:6)
2Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky Prospect 33, Moscow 119071, Russia. E-mail:
[email protected].
D(cid:14)(cid:20)(cid:17)(cid:10)(cid:17)(cid:26)(cid:17)(cid:7)(cid:21)(cid:9)(cid:16)#(cid:29)(cid:12)/(cid:7)I(cid:22)(cid:16)(cid:29)(cid:16)(cid:31)(cid:10)(cid:10)(cid:7)(cid:10)(cid:7)I(cid:11)(cid:16)(cid:29)LQ(cid:10)(cid:10)(cid:7)(cid:1)JK!(cid:7)R(cid:12)(cid:14)(cid:10)(cid:14)(cid:20)(cid:22)(cid:10)(cid:19)(cid:7)(cid:21)(cid:9)(cid:16)(cid:20)(cid:21)(cid:12)(cid:22)(cid:17)(cid:7)77!(cid:7)(cid:5)(cid:16)(cid:20)(cid:22)(cid:11)(cid:15)(cid:7)(cid:23)(cid:23)-(cid:24)P(cid:23)!(cid:7)(cid:1)(cid:16)(cid:20)(cid:20)(cid:10)1(cid:6)
KEY WORDS: Spiders, Russia, Kola Peninsula, tundra, chorology, new synonym.
(cid:28)R(cid:4)S(cid:2)$T(cid:2)(cid:7)URV$JW(cid:7)(cid:8)(cid:15)(cid:26)(cid:22)(cid:10)!(cid:7)(cid:1)(cid:16)(cid:20)(cid:20)(cid:10)1!(cid:7)(cid:28)(cid:16)(cid:29)(cid:30)(cid:20)(cid:22)(cid:10)(cid:19)(cid:7)(cid:21) (cid:16)(cid:11)!(cid:7)(cid:17)(cid:26)(cid:14)(cid:13)(cid:9)(cid:15)!(cid:7)"(cid:16)(cid:9)(cid:16)(cid:29)(cid:16)(cid:31)(cid:10)1!(cid:7)(cid:14)(cid:16)(cid:11)(cid:18)(cid:19)(cid:7)(cid:20)(cid:10)(cid:14)(cid:16)(cid:14)(cid:10)/(cid:6)
ABSTRACT. A checklist of 105 spiders from the (cid:21)(cid:16)#(cid:12)(cid:9)(cid:12)G(cid:30)(cid:12)(cid:7)(cid:28)(cid:16)(cid:29)(cid:30)(cid:20)(cid:22)(cid:16)(cid:31)(cid:16)(cid:7)(cid:21) (cid:16)(cid:11)(cid:15)(cid:7)1(cid:11)(cid:29)1(cid:12)(cid:17)(cid:20)1(cid:7)(cid:20)(cid:15)/(cid:16)(cid:19)(cid:7)(cid:20)(cid:12)(cid:11)(cid:12)(cid:9)
tundra zone of the Kola Peninsula is based and com- (cid:14)(cid:16)(cid:19)(cid:7)(cid:17)(cid:16)0(cid:22)(cid:16)(cid:19)(cid:7)(cid:9)(cid:15)(cid:20)(cid:21)(cid:9)(cid:16)(cid:20)(cid:17)(cid:9)(cid:15)(cid:14)(cid:12)(cid:14)(cid:10)1(cid:6)(cid:7)X(cid:15)(cid:26)(cid:14)(cid:15)(cid:7)(cid:17)(cid:26)(cid:14)(cid:13)(cid:9)(cid:7)(cid:28)(cid:16)(cid:29)(cid:30)(cid:20)
piled on the original collections and on available litera- (cid:22)(cid:16)(cid:31)(cid:16)(cid:7)(cid:21) (cid:16)(cid:11)(cid:15)(cid:7)"(cid:15)(cid:9)(cid:15)(cid:22)(cid:17)(cid:12)(cid:9)(cid:10)(cid:27)(cid:26)(cid:12)(cid:17)(cid:20)1(cid:7)(cid:16)(cid:17)(cid:20)(cid:26)(cid:17)(cid:20)(cid:17)(cid:11)(cid:10)(cid:12)/(cid:7)(cid:20)(cid:16)#(cid:20)(cid:17)(cid:11)(cid:12)(cid:14)
ture data. Oreoneta sinuosa (Tullgren, 1955) is report- (cid:14)(cid:16)(cid:7)(cid:15)(cid:9)(cid:22)(cid:17)(cid:10)0(cid:12)(cid:20)(cid:22)(cid:10)"(cid:7)I(cid:29)(cid:12)/(cid:12)(cid:14)(cid:17)(cid:16)(cid:11)!(cid:7)(cid:10)/(cid:12)(cid:12)(cid:17)(cid:7)(cid:17)(cid:10)(cid:21)(cid:10)0(cid:14)(cid:18)(cid:19)(cid:7)#(cid:16)(cid:9)(cid:12)
ed as a new to the Russian fauna. A new synonym is (cid:15)(cid:29)(cid:30)(cid:14)(cid:18)(cid:19)(cid:7)(cid:16)#(cid:29)(cid:10)(cid:22)(cid:7)(cid:10)(cid:7)(cid:20)(cid:16)(cid:20)(cid:17)(cid:15)(cid:11)(cid:29)(cid:12)(cid:14)(cid:15)(cid:7):(cid:10)(cid:9)(cid:16)(cid:22)(cid:16)(cid:15)(cid:9)(cid:12)(cid:15)(cid:29)(cid:30)(cid:14)(cid:18)/(cid:10)(cid:7)#(cid:16)
proposed: Lepthyphantes murmanicola Strand, 1913, (cid:9)(cid:12)(cid:15)(cid:29)(cid:30)(cid:14)(cid:18)/(cid:10)(cid:7)(cid:10)(cid:7)(cid:21)(cid:16)(cid:29)(cid:10)(cid:27)(cid:16)(cid:14)(cid:15)(cid:29)(cid:30)(cid:14)(cid:18)/(cid:10)(cid:7)(cid:11)(cid:10)(cid:13)(cid:15)/(cid:10)(cid:6)(cid:7)(cid:3)(cid:16)(cid:14)(cid:15)(cid:29)(cid:30)(cid:14)(cid:18)(cid:12)
syn.n. = Oryphantes angulatus (O. Pickard-Cambridge, (cid:17)(cid:26)(cid:14)(cid:13)(cid:9)(cid:16)(cid:11)(cid:18)(cid:12)(cid:7)(cid:20)(cid:16)(cid:16)#E(cid:12)(cid:20)(cid:17)(cid:11)(cid:15)(cid:7)(cid:14)(cid:15)(cid:10)#(cid:16)(cid:29)(cid:12)(cid:12)(cid:7)#(cid:12)(cid:13)(cid:14)(cid:18)(cid:7)(cid:11)(cid:7)(cid:17)(cid:15)(cid:22)(cid:20)(cid:16)(cid:14)(cid:16)
1881). Some previous misidentifications are rectified. /(cid:10)0(cid:12)(cid:20)(cid:22)(cid:16)/(cid:7)(cid:21)(cid:29)(cid:15)(cid:14)(cid:12)(cid:7)(cid:10)(cid:7)"(cid:15)(cid:9)(cid:15)(cid:22)(cid:17)(cid:12)(cid:9)(cid:10)(cid:27)(cid:26)L(cid:17)(cid:20)1(cid:7)(cid:16)(cid:17)(cid:20)(cid:26)(cid:17)(cid:20)(cid:17)(cid:11)(cid:10)(cid:12)/
Faunistic data and the distribution pattern are indicated (cid:20)(cid:21)(cid:12)Q(cid:10)2(cid:10)0(cid:14)(cid:18)"(cid:7)(cid:11)(cid:10)(cid:13)(cid:16)(cid:11)(cid:6)(cid:7)X(cid:15)(cid:26)(cid:14)(cid:15)(cid:7)(cid:10)(cid:14)(cid:17)(cid:9)(cid:15)(cid:27)(cid:16)(cid:14)(cid:15)(cid:29)(cid:30)(cid:14)(cid:18)"!(cid:7)(cid:15)(cid:27)(cid:16)
for each species. For the majority of the recorded spi- (cid:14)(cid:15)(cid:29)(cid:30)(cid:14)(cid:18)"(cid:7)(cid:10)(cid:7)(cid:15)(cid:14)(cid:17)(cid:9)(cid:16)(cid:21)(cid:16)(cid:31)(cid:12)(cid:14)(cid:14)(cid:16)(cid:7)/(cid:16)(cid:13)(cid:10)2(cid:10)Q(cid:10)(cid:9)(cid:16)(cid:11)(cid:15)(cid:14)(cid:14)(cid:18)"(cid:7)(cid:20)(cid:16)(cid:16)#
ders, the northern coast of the Kola Peninsula is the E(cid:12)(cid:20)(cid:17)(cid:11)(cid:7)(cid:27)(cid:14)(cid:15)0(cid:10)(cid:17)(cid:12)(cid:29)(cid:30)(cid:14)(cid:16)(cid:7)#(cid:16)(cid:31)(cid:15)0(cid:12)Y(cid:7)(cid:10)/(cid:12)(cid:14)(cid:14)(cid:16)(cid:7)I(cid:17)(cid:10)(cid:7)(cid:17)(cid:10)(cid:21)(cid:18)(cid:7)(cid:20)(cid:16)(cid:16)#
northernmost point of their known distributions. The E(cid:12)(cid:20)(cid:17)(cid:11)(cid:7)(cid:20)(cid:29)(cid:26)G(cid:15)(cid:17)(cid:7)(cid:16)(cid:20)(cid:14)(cid:16)(cid:11)(cid:14)(cid:18)/(cid:7)(cid:21)(cid:9)(cid:10)#(cid:12)G(cid:10)E(cid:12)/(cid:7)(cid:14)(cid:12)(cid:7)(cid:17)(cid:16)(cid:29)(cid:30)(cid:22)(cid:16)(cid:7)(cid:13)(cid:29)1
spider fauna of the tundra zone of the Kola Peninsula LG(cid:14)(cid:18)"(cid:7)2(cid:15)(cid:26)(cid:14)(cid:10)(cid:20)(cid:17)(cid:10)0(cid:12)(cid:20)(cid:22)(cid:10)"(cid:7)I(cid:29)(cid:12)/(cid:12)(cid:14)(cid:17)(cid:16)(cid:11)!(cid:7)(cid:14)(cid:16)(cid:7)(cid:10)(cid:7)(cid:13)(cid:29)1(cid:7)(cid:14)(cid:12)
has a typical boreal face, and is composed of wide- /(cid:14)(cid:16)(cid:31)(cid:16)0(cid:10)(cid:20)(cid:29)(cid:12)(cid:14)(cid:14)(cid:18)"(cid:7)(cid:15)(cid:9)(cid:22)(cid:17)(cid:16)(cid:15)(cid:29)(cid:30)(cid:21)(cid:10)(cid:19)(cid:20)(cid:22)(cid:10)"(cid:7)(cid:11)(cid:10)(cid:13)(cid:16)(cid:11)(cid:6)
spread or European boreal/polyzonal species. There
are no real Arctic elements found here. The zonal plant Introduction
communities have the fewest spider species, lack of the
specific elements, and are comprised of boreal or ubiq- The tundra landscape zone of the Kola Peninsula
uitous species. Intrazonal and anthropogenic commu- is a narrow belt ca 30–60 km wide (see Map). Owing
nities are much richer, and together with azonal com- to a strong vertical partitioning of the territory, nu-
munities serve as a shelter not only for the southern merous lakes and rivers, the tundra landscape is strong-
elements, but also for the arcto-alpine species. ly fragmented. The zonal plant communities are rep-
resented by different types of the undershrub-moss-
(cid:1)(cid:2)(cid:3)(cid:4)(cid:5)(cid:2)(cid:6)(cid:7)(cid:8)(cid:9)(cid:10)(cid:11)(cid:12)(cid:13)(cid:12)(cid:14)(cid:7)(cid:15)(cid:14)(cid:14)(cid:16)(cid:17)(cid:10)(cid:9)(cid:16)(cid:11)(cid:15)(cid:14)(cid:14)(cid:18)(cid:19)(cid:7)(cid:20)(cid:21)(cid:10)(cid:20)(cid:16)(cid:22) lichen tundra vegetation which alternates with open
(cid:23)(cid:24)(cid:25)(cid:7)(cid:11)(cid:10)(cid:13)(cid:16)(cid:11)(cid:7)(cid:21)(cid:15)(cid:26)(cid:22)(cid:16)(cid:11)(cid:7)(cid:17)(cid:26)(cid:14)(cid:13)(cid:9)(cid:16)(cid:11)(cid:16)(cid:19)(cid:7)(cid:27)(cid:16)(cid:14)(cid:18)(cid:7)(cid:28)(cid:16)(cid:29)(cid:30)(cid:20)(cid:22)(cid:16)(cid:31)(cid:16)(cid:7)(cid:21) (cid:16)(cid:11)(cid:15)! water surfaces, sedge-moss bogs, peatbogs, Betula
(cid:20)(cid:16)(cid:20)(cid:17)(cid:15)(cid:11)(cid:29)(cid:12)(cid:14)(cid:14)(cid:18)(cid:19)(cid:7)(cid:14)(cid:15)(cid:7)(cid:16)(cid:20)(cid:14)(cid:16)(cid:11)(cid:15)(cid:14)(cid:10)(cid:10)(cid:7)(cid:14)(cid:12)(cid:13)(cid:15)(cid:11)(cid:14)(cid:10)"(cid:7)(cid:20)#(cid:16)(cid:9)(cid:16)(cid:11)(cid:7)(cid:10)(cid:7)(cid:29)(cid:10) and Salix stands in depressions. The tops of hills and
(cid:17)(cid:12)(cid:9)(cid:15)(cid:17)(cid:26)(cid:9)(cid:14)(cid:18)"(cid:7)(cid:13)(cid:15)(cid:14)(cid:14)(cid:18)"(cid:6)(cid:7)$(cid:10)(cid:13)(cid:7)(cid:1)(cid:2)(cid:3)(cid:4)(cid:5)(cid:3)(cid:6)(cid:7)(cid:8)(cid:9)(cid:10)(cid:5)(cid:11)(cid:4)(cid:9)(cid:7)(cid:7)%&’(()*+,! abrupt stony slopes are occupied by crustose lichen
(cid:23)-(cid:25)(cid:25).(cid:7)(cid:11)(cid:21)(cid:12)(cid:9)(cid:11)(cid:18)(cid:12)(cid:7)(cid:16)(cid:17)/(cid:12)0(cid:12)(cid:14)(cid:7)(cid:13)(cid:29)1(cid:7)2(cid:15)(cid:26)(cid:14)(cid:18)(cid:7)(cid:1)(cid:16)(cid:20)(cid:20)(cid:10)(cid:10)(cid:6)(cid:7)(cid:12)(cid:3)(cid:13)(cid:6)(cid:14)(cid:15)(cid:16) communities. The seashore is usual narrow, with salt
(cid:13)(cid:14)(cid:7)(cid:5)(cid:6)(cid:3)(cid:9)(cid:8)(cid:17)(cid:11)(cid:2)(cid:17)(cid:7)(cid:5)(cid:10)(cid:18)(cid:4)(cid:19)(cid:7)(cid:7)34*5,6!(cid:7)(cid:23)-(cid:23)7!(cid:7)89,(cid:6),(cid:6)(cid:7)(cid:21)(cid:9)(cid:10)(cid:27)(cid:14)(cid:15)(cid:14) marshes and meadow-like vegetation. Data on 55 spi-
/(cid:29)(cid:15)(cid:13):(cid:10)/(cid:7)(cid:20)(cid:10)(cid:14)(cid:16)(cid:14)(cid:10)/(cid:16)/(cid:8)(cid:1)(cid:2)(cid:15)(cid:13)(cid:14)(cid:7)(cid:5)(cid:6)(cid:3)(cid:9)(cid:8)(cid:7)(cid:5)(cid:20)(cid:11)(cid:19)(cid:7)(cid:6)(cid:11)(cid:9)(cid:7)%;(cid:6)(cid:7)<= ders known from the tundra zone of Kola Peninsula
>?5*6 @5AB*=6)+!(cid:7)(cid:23)CC(cid:23).(cid:6)(cid:7)D(cid:20)(cid:21)(cid:9)(cid:15)(cid:11)(cid:29)(cid:12)(cid:14)(cid:18)(cid:7)(cid:14)(cid:12)(cid:22)(cid:16)(cid:17)(cid:16)(cid:9)(cid:18)(cid:12) are contained in two papers (Fedotov [1912] and
(cid:16):(cid:10)#(cid:16)0(cid:14)(cid:18)(cid:12)(cid:7)(cid:16)(cid:21)(cid:9)(cid:12)(cid:13)(cid:12)(cid:29)(cid:12)(cid:14)(cid:10)1(cid:7)(cid:21)(cid:9)(cid:12)(cid:13)(cid:18)(cid:13)(cid:26)E(cid:10)"(cid:7)(cid:15)(cid:11)(cid:17)(cid:16)(cid:9)(cid:16)(cid:11)(cid:6)(cid:7)F(cid:29)1 Shokhin et al. [2004]). New material from Teriberka
(cid:22)(cid:15)G(cid:13)(cid:16)(cid:31)(cid:16)(cid:7)(cid:11)(cid:10)(cid:13)(cid:15)(cid:7)(cid:21)(cid:9)(cid:10)(cid:11)(cid:12)(cid:13)(cid:12)(cid:14)(cid:18)(cid:7)/(cid:12)(cid:20)(cid:17)(cid:15)(cid:7)(cid:14)(cid:15)"(cid:16)(cid:13)(cid:16)(cid:22)(cid:7)(cid:10)(cid:7)(cid:17)(cid:10)(cid:21)(cid:7)(cid:15)(cid:9)(cid:12)(cid:15) and Dal’niye Zelentsy has doubled the number of
(cid:29)(cid:15)(cid:6)(cid:7)F(cid:29)1(cid:7)#(cid:16)(cid:29)(cid:30):(cid:10)(cid:14)(cid:20)(cid:17)(cid:11)(cid:15)(cid:7)(cid:16)(cid:17)/(cid:12)0(cid:12)(cid:14)(cid:14)(cid:18)"(cid:7)(cid:11)(cid:10)(cid:13)(cid:16)(cid:11)(cid:7)(cid:20)(cid:12)(cid:11)(cid:12)(cid:9)(cid:14)(cid:16)(cid:12) species in this list.
42 A.V. Tanasevitch & L.B. Rybalov
Map. Localities in Kola Peninsula: 1 — Bolshoy Aynovskiy Island; 2 — Rybachiy Peninsula; 3 — Shalim Island; 4 — Sedlovaty
Island, Yekaterininskiy Island, Pala Bay, Olen’ya Bay, Murmansk Field Station; 5 — Srednyaya Bay; 6 — Tyuva Bay; 7 — Kildin Island;
8 — Teriberka, Lodeynoye, Logeynaya Bay, Korabelnaya Bay; 9 — Dal’niye Zelentsy; circle — literature data [Fedotov, 1912], square —
collecting by L. Rybalov, V. Semenov & V. Piryugin; rhomb — collecting by A. Babenko; asterisk — Murmansk City; dotted line —
southern border of tundra landscape zone.
(cid:28)(cid:15)(cid:9)(cid:17)(cid:15)(cid:6)(cid:7)Z(cid:16)0(cid:22)(cid:10)(cid:7)(cid:20)#(cid:16)(cid:9)(cid:16)(cid:11)(cid:7)(cid:14)(cid:15)(cid:7)(cid:28)(cid:16)(cid:29)(cid:30)(cid:20)(cid:22)(cid:16)/(cid:7)(cid:21)(cid:16)(cid:29)(cid:26)(cid:16)(cid:20)(cid:17)(cid:9)(cid:16)(cid:11)(cid:12)W(cid:7)(cid:23)(cid:7)[(cid:7)\(cid:16)(cid:29)(cid:30):(cid:16)(cid:19)(cid:7)J(cid:19)(cid:14)(cid:16)(cid:11)(cid:20)(cid:22)(cid:10)(cid:19)(cid:7)(cid:16) (cid:11)Y(cid:7)O(cid:7)[(cid:7)(cid:1)(cid:18)#(cid:15)0(cid:10)(cid:19)(cid:7)(cid:21) (cid:16)(cid:11)Y(cid:7)7(cid:7)[(cid:7)](cid:15)(cid:29)(cid:10)/(cid:7)(cid:16) (cid:11)Y(cid:7)M(cid:7)[
U(cid:12)(cid:13)(cid:29)(cid:16)(cid:11)(cid:15)(cid:17)(cid:18)(cid:19)(cid:7)(cid:16) (cid:11)!(cid:7)(cid:2)(cid:22)(cid:15)(cid:17)(cid:12)(cid:9)(cid:10)(cid:14)(cid:10)(cid:14)(cid:20)(cid:22)(cid:10)(cid:19)(cid:7)(cid:16) (cid:11)!(cid:7)(cid:8)(cid:15)(cid:29)(cid:15)(cid:7)(cid:31)(cid:26)#(cid:15)!(cid:7)V(cid:29)(cid:12)(cid:14)(cid:30)1(cid:7)(cid:31)(cid:26)#(cid:15)!(cid:7)(cid:5)(cid:26)(cid:9)/(cid:15)(cid:14)(cid:20)(cid:22)(cid:15)1(cid:7)#(cid:10)(cid:16)(cid:29)(cid:16)(cid:31)(cid:10)0(cid:12)(cid:20)(cid:22)(cid:15)1(cid:7)(cid:20)(cid:17)(cid:15)(cid:14)Q(cid:10)1Y(cid:7)(cid:25)(cid:7)[(cid:7)U(cid:9)(cid:12)(cid:13)(cid:14)11(cid:7)(cid:31)(cid:26)#(cid:15)Y(cid:7)^(cid:7)[(cid:7)ZL(cid:11)(cid:15)
(cid:31)(cid:26)#(cid:15)Y(cid:7)P(cid:7)[(cid:7)(cid:28)(cid:10)(cid:29)(cid:30)(cid:13)(cid:10)(cid:14)(cid:7)(cid:16) (cid:11)Y(cid:7)C(cid:7)[(cid:7)Z(cid:12)(cid:9)(cid:10)#(cid:12)(cid:9)(cid:22)(cid:15)!(cid:7)R(cid:16)(cid:13)(cid:12)(cid:19)(cid:14)(cid:16)(cid:12)!(cid:7)R(cid:16)(cid:13)(cid:12)(cid:19)(cid:14)(cid:15)1(cid:7)(cid:31)(cid:26)#(cid:15)!(cid:7)(cid:28)(cid:16)(cid:9)(cid:15)#(cid:12)(cid:29)(cid:30)(cid:14)(cid:15)1(cid:7)(cid:31)(cid:26)#(cid:15)Y(cid:7)-(cid:7)[(cid:7)F(cid:15)(cid:29)(cid:30)(cid:14)(cid:10)(cid:12)(cid:7)(cid:3)(cid:12)(cid:29)(cid:12)(cid:14)Q(cid:18)Y(cid:7)(cid:22)(cid:9)(cid:26)(cid:31)(cid:7)[(cid:7)(cid:29)(cid:10)(cid:17)(cid:12)(cid:9)(cid:15)(cid:17)(cid:26)(cid:9)(cid:14)(cid:18)(cid:12)
(cid:13)(cid:15)(cid:14)(cid:14)(cid:18)(cid:12)(cid:7)_X(cid:12)(cid:13)(cid:16)(cid:17)(cid:16)(cid:11)!(cid:7)(cid:23)-(cid:23)O‘!(cid:7)(cid:22)(cid:11)(cid:15)(cid:13)(cid:9)(cid:15)(cid:17)(cid:7)[(cid:7)(cid:20)#(cid:16)(cid:9)(cid:18)(cid:7)R(cid:6)(cid:7)(cid:1)(cid:18)#(cid:15)(cid:29)(cid:16)(cid:11)(cid:15)!(cid:7)$(cid:6)(cid:7)U(cid:12)/(cid:12)(cid:14)(cid:16)(cid:11)(cid:15)(cid:7)(cid:10)(cid:7)$(cid:6)(cid:7)(cid:8)(cid:10)(cid:9)L(cid:31)(cid:10)(cid:14)(cid:15)Y(cid:7)(cid:9)(cid:16)/#(cid:7)[(cid:7)(cid:20)#(cid:16)(cid:9)(cid:18)(cid:7)J(cid:6)(cid:7)\(cid:15)#(cid:12)(cid:14)(cid:22)(cid:16)Y(cid:7)(cid:27)(cid:11)(cid:12)(cid:27)(cid:13)(cid:16)0(cid:22)(cid:15)(cid:7)[(cid:7)(cid:31)(cid:6)
(cid:5)(cid:26)(cid:9)/(cid:15)(cid:14)(cid:20)(cid:22)Y(cid:7)(cid:21)(cid:26)(cid:14)(cid:22)(cid:17)(cid:10)(cid:9)(cid:14)(cid:15)1(cid:7)(cid:29)(cid:10)(cid:14)(cid:10)1(cid:7)[(cid:7)LG(cid:14)(cid:15)1(cid:7)(cid:31)(cid:9)(cid:15)(cid:14)(cid:10)Q(cid:15)(cid:7)(cid:17)(cid:26)(cid:14)(cid:13)(cid:9)(cid:16)(cid:11)(cid:16)(cid:19)(cid:7)(cid:27)(cid:16)(cid:14)(cid:18)(cid:6)
Material and methods abiotic factors are more extreme than they are in the
intrazonal communities, and living conditions there-
This study is based mainly on the collections made fore are more adverse.
recently in Teriberka (ca 69°9’N, 35°8’E) by Leonid Intrazonal communities — communities distrib-
B. Rybalov and Victor B. Semenov in 3–11.VII.2008, uted in one or several zones, the typical representatives
as well as by Victor S. Piryugin in 3–7.VIII.2009. of this type are swamps and the river valley vegetation,
Some material was collected at Dal’niye Zelentsy (ca e.g., willow stands on floodplains, meadows, pebble
69°7’N, 36°3’E) by Anatoly B. Babenko in 4–26.VII. banks, rocky and/or grassy slopes, forest stands on
2009. Spiders were collected by the sifting of moss and river banks or river terraces, etc. Commonly, parts of
litter, soil sampling, pitfall trapping, and hand collect- intrazonal type of communities are situated on water-
ing. In addition, all available literature data on spiders sheds among zonal type of vegetation: there are willow
of the northern coast of the Kola Peninsula is com- stands in depressions, shrubby banks of lakes, flat-hill
piled. Linyphiids spiders recorded by Shokhin, et al. peatbogs with sedge fens or sphagnum bogs in hol-
[2004] from Teriberka were examined. All material is lows, grassy slopes of small brooks, etc. The intrazonal
stored in the personal collection of Andrei V. Ta- communities are smoothing gradient of the climatic
nasevitch, Moscow, Russia. factors (especially, temperature and humidity), making
For study of the zonal-landscape distribution of living conditions substantially better (less pessimal).
tundra spiders we used the scheme proposed by Cher- Some rivers penetrate several zones/subzones and riv-
nov [1978] for dividing plant communities in the Sub- er valleys are powerful channels for the penetration of
arctic. Based on this classification, there are three main southern forms to the North and vice versa. Some of
zonal types of communities in tundra (for more details, southern species can, using intrazonal biotopes, come
see Tanasevitch & Koponen [2007]): to watersheds and become part of the tundra fauna.
Zonal communities — plant communities distrib- Azonal communities — plant communities which
uted into one zone and which are situated at the water- not confined to a certain zone, but which are distribut-
shed. Zonal communities define the vegetation zonal ed in many zones; in the Kola tundra this type is sea-
face of the landscape. For the Kola tundra, this is the coast meadows and marshes.
polyvariant lichen-moss-undershrub-dwarf birch (or Anthropogenic communities — these kinds of com-
willow-dwarf birch) tundra. In zonal communities, the munities are very common in the North today. It is not
On spiders from the tundra zone of the Kola Peninsula 43
only anthropogenically changed natural vegetation Agyneta mossica (Schikora, 1993)
(meadow formation sites, roadsides of roads, etc.),
but also dump sites, dustbins, destroyed construc- MATERIAL. Intra: 1 (cid:1), Teriberka, Orlovka River valley,
tions, and so on. swampy meadow with Betula nana, Carex sp., Eriophorum sp.,
Vaccinium oxycoccus, Rubus chamaemorus, Menyanthes trifo-
Distributional pattern of species follows that sug-
liata, Andromeda polifolia, Sphagnum sp., 9.VII.2008, leg. L.R.
gested by Tanasevitch & Koponen [2007] with a few
& V.S.
minor changes. COMMENTS. Teriberka is the northernmost known
Abbreviation used in text: Zonal, Intra, Azonal & distribution of the species.
Anthro — zonal, intrazonal, azonal & anthropogenic RANGE. West Palaearctic boreo-nemoral.
communities, respectively, L.R. — L. Rybalov, V.S. —
V. Semenov, V.P. — V. Piryugin. Agyneta nigripes (Simon, 1884)
List of spiders MATERIAL. Zonal: 1 (cid:1), Dal’niye Zelentsy, lichen tundra on
top of hill, pitfall traps, 4–26.VII.2009, leg. A.B.
RANGE. Holarctic arcto-alpine.
Fam. THERIDIIDAE (2)
Allomengea vidua (L. Koch, 1879)
Robertus lividus (Blackwall, 1836)
2004 Allomengea vidua — Shokhin et al.: 143, examined.
2004 Robertus lividus — Shokhin et al.: 142. PREVIOUS RECORDS. Teriberka [Shokhin et al.,
MATERIAL. Intra: 3 (cid:1)(cid:1), Teriberka, Orlovka River valley,
2004].
elfin Betula sp. sparse forest on gentle slope with Vaccinium myr-
tillus, Cornus suecica, Geranium sanguineum, Epilobium angusti- COMMENTS. Teriberka is the northernmost known
folium, Calamagrostis sp., 9.VII.2008, leg. L.R. & V. S; 1 (cid:1), distribution of the species.
Orlovka River valley, elfin Betula sp. sparse forest on gentle slope RANGE. Holarctic.
with Vaccinium myrtillus, Gymnocarpium dryopteris, Cornus sueci-
ca, no moss or lichen, 9.VII.2008, leg. L.R. & V.S. Azonal: 2 (cid:1)(cid:1),
Bathyphantes gracilis (Blackwall, 1841)
Dal’niye Zelentsy, meadow-like community on slope to sea, pitfall
traps, 4–26.VII.2009, leg. A.B. Anthro: 1 (cid:2), ruderal plant commu-
nity with Poa sp., Deschampsia caespitosa, Epilobium angustifo- MATERIAL. Intra: 1 (cid:1), 1 (cid:2), Teriberka, Sphagnum-Carex
lium, etc., 3.VII.2008, leg. L.R. & V.S. bog in tundra, 3.VII.2008, leg. L.R. & V.S.; 1 (cid:1), 1 (cid:2), Carex-
PREVIOUS RECORDS. Teriberka [Shokhin et al., Sphagnum-Eriophorum bog with Salix sp. stands & Comarum
2004]. palustre, 6.VII.2008, leg. L.R. & V.S.; 1 (cid:1), Dal’niye Zelentsy,
swampy bank of lake, pitfall traps, 4–26.VII.2009, leg. A.B.
COMMENTS. Teriberka is the northernmost known
RANGE. Holarctic polyzonal.
distribution of the species.
RANGE. Palaearctic-Alaskan boreo-nemoral.
Bolephthyphantes index (Thorell, 1856)
Robertus scoticus Jackson, 1914
1912 Bolephthyphantes index — Fedotov: 461.
REMARKS. This species is absent from the materi-
MATERIAL. Zonal: 3 (cid:2)(cid:2), Teriberka, near Tretie Titovskoye
al examined.
Lake, dwarf birch-lichen mountain tundra with Betula nana, Cla-
donia sp., 4–6.VIII.2009, leg. V.P. Intra: 1 (cid:2), elfin Betula sp. PREVIOUS RECORDS. Murmansk Field Station
sparse forest on hill slope with Vaccinium myrtillus, Gymnocarpi- [Fedotov, 1912].
um dryopteris, Cornus suecica, Calamagrostis sp., no moss or COMMENTS. This is the northernmost known
lichen, 10.VII.2008, leg. L.R. & V.S.; 1 (cid:2), Betula tortuosa under- distribution of the species.
growth on airfield with Empetrum sp., Vaccinium myrtillus, Pleu-
RANGE. Palaearctic boreal-nemoral. Introduced to
rozium schreberi, Hylocomium sp., etc., 5.VII.2008, leg. L.R. &
V.S.; 1 (cid:2), same, 4–6.VIII.2009, leg. V.P. Anthro: 2 (cid:2)(cid:2), ruderal Greenland [Marusik et. al., 2006].
plant community with Poa sp., Deschampsia caespitosa, Epilobi-
um angustifolium, etc., 3.VII.2008, leg. L.R. & V.S. Bolyphantes luteolus (Blackwall, 1833)
COMMENTS. Teriberka is the northernmost known
distribution of the species.
1912 Bolyphantes luteolus – Fedotov: 461.
RANGE. Palaearctic boreo-nemoral. REMARKS. This species is absent from the materi-
al examined.
Fam. LINYPHIIDAE (65)
PREVIOUS RECORDS. Bolshoy Aynovskiy Island,
Rybachiy Peninsula, Lodeynaya Bay [Fedotov, 1912].
Agyneta conigera (O. Pickard-Cambridge, 1863)
COMMENTS. Bolshoy Aynovskiy Island is the
northernmost known distribution of the species.
MATERIAL. Intra: 2 (cid:2)(cid:2), Teriberka, Orlovka River valley,
RANGE. Palaearctic boreal-nemoral.
elfin Betula sp. sparse forest on gentle slope with Vaccinium myr-
tillus, Cornus suecica, Geranium sanguineum, Epilobium angusti-
folium, Calamagrostis sp., 9.VII.2008, leg. L.R. & V.S. Centromerus arcanus (O. Pickard-Cambridge,
COMMENTS. Teriberka is the northernmost known 1873)
distribution of the species.
RANGE. Palaearctic boreo-nemoral. 2004 Centromerus arcanus — Shokhin et al.: 143, examined.
44 A.V. Tanasevitch & L.B. Rybalov
MATERIAL. Zonal: 3 (cid:2)(cid:2), Teriberka, undershrub-lichen moun- PREVIOUS RECORDS. Teriberka [Shokhin et al.,
tain tundra with Betula nana, Vaccinium myrtillus, Empetrum sp., 2004].
Pleurozium schreberi, 4–6.VIII.2009, leg. V.P. Intra: 3 (cid:2)(cid:2), elfin
COMMENTS. Teriberka is the northernmost known
Betula sp. sparse forest on hill slope with Vaccinium myrtillus,
distribution of the species.
Gymnocarpium dryopteris, Cornus suecica, Calamagrostis sp., no
moss or lichen, 10.VII.2008, leg. L.R. & V.S.; 1 (cid:1), 3 (cid:2)(cid:2), Orlovka RANGE. Holarctic polyzonal.
River valley, elfin Betula sp. sparse forest on gentle slope with
Vaccinium myrtillus, Gymnocarpium dryopteris, Cornus suecica, Diplocephalus cristatus (Blackwall, 1833)
no moss or lichen, 9.VII.2008, leg. L.R. & V.S.; 3 (cid:2)(cid:2), Orlovka
River valley, swampy meadow with Betula nana, Carex sp., Erio-
MATERIAL. Anthro: 1 (cid:1), 1 (cid:2), Teriberka, meadow in village
phorum sp., Vaccinium oxycoccus, Rubus chamaemorus, Menyan-
with Calamagrostis sp., Epilobium angustifolium, Matricaria re-
thes trifoliata, Andromeda polifolia, Sphagnum sp., 9.VII.2008,
leg. L.R. & V.S.; 1 (cid:1), Dal’niye Zelentsy, Betula sp. sparse forest, cutita, Taraxacum sp., 11.VII.2008, leg. L.R. & V.S.
pitfall traps, 8–26.VII.2009, leg. A.B. Anthro: 6 (cid:2)(cid:2), Teriberka, COMMENTS. This is the northernmost known dis-
Betula tortuosa undergrowth on airfield with Empetrum sp., Vac- tribution of the species.
cinium myrtillus, Pleurozium schreberi, Hylocomium sp., etc., RANGE. Holarctic boreo-nemoral.
5.VII.2008, leg. L.R. & V.S.; 5 (cid:2)(cid:2), ruderal plant community with
Poa sp., Deschampsia caespitosa, Epilobium angustifolium, etc., Drepanotylus uncatus (O. Pickard-Cambridge,
3.VII.2008, leg. L.R. & V.S.
1872)
PREVIOUS RECORDS. Teriberka [Shokhin et al.,
2004].
2004 Erigoninae — Shokhin et al.: 143, examined.
COMMENTS. Teriberka is the northernmost known MATERIAL. Intra: 1 (cid:1), Teriberka, Sphagnum-Carex bog in
distribution of the species. tundra, 4–6.VIII.2009, leg. V.P.; 1 (cid:1), Carex-Sphagnum-Eriopho-
RANGE. Palaearctic polyzonal. rum bog with Salix sp. stands, Comarum palustre, 6.VII.2008, leg.
L.R. & V.S.; 2 (cid:2)(cid:2), Dal’niye Zelentsy, sedge-sphagnum peatbog,
Ceratinella brevipes (Westring, 1851) pitfall traps, 4–26.VII.2009, leg. A.B.; 1 (cid:2), Salix sp. with Sphag-
num sp. in depression on slope, pitfall traps, 4–26.VII.2009, leg.
A.B.
MATERIAL. Intra: 1 (cid:2), Dal’niye Zelentsy, Salix sp. with PREVIOUS RECORDS. Teriberka [Shokhin et al.,
Sphagnum sp. in depression on slope, pitfall traps, 4–26.VII.2009,
2004, as “Erigoninae”].
leg. A.B. Azonal: 1 (cid:2), meadow-like community on slope to sea,
pitfall traps, 4–26.VII.2009, leg. A.B. COMMENTS. Teriberka is the northernmost known
COMMENTS. This is the northernmost known dis- distribution of the species.
tribution of the species. RANGE. West Palaearctic.
RANGE. Palaearctic polyzonal.
Erigone arctica maritima (cid:1)(cid:2)(cid:3)(cid:4)(cid:5)(cid:6)(cid:7)(cid:8)(cid:9)(cid:10), 1902
Ceratinella brevis (Wider, 1834)
1912 Erigone arctica — Fedotov: 457.
MATERIAL. Intra: 1 (cid:2), Teriberka, Orlovka River valley, 2M0A04T EER. IaArcLti.c Ian —tra S: h2o k(cid:2)h(cid:2)i,n Deta la’ln.:i y1e4 3Z,e elexnatmsyin, esda.ndy bank of
swampy meadows with Betula nana, Carex sp., Eriophorum sp., lake, pitfall traps, 8–16.VII.2009, leg. A.B. Azonal: 1 (cid:2), Teriberka,
Vaccinium oxycoccus, Rubus chamaemorus, Menyanthes trifolia-
Orlovka River bay, seacoast marsh, under trash & seaweeds (Fucus
ta, Andromeda polifolia, Sphagnum sp., 10.VII.2008, leg. L.R. & sp., Laminaria sp.), 10.VII.2008, leg. L.R. & V.S. Anthro: 2 (cid:2)(cid:2),
V.S.
agricultural meadow with Deschampsia caespitosa, Phleum prat-
COMMENTS. Teriberka is the northernmost known
ense, Epilobium angustifolium, etc., 4.VII.2008, leg. L.R. & V.S.;
distribution of the species. A record of C. brevis from 1 (cid:1), agricultural meadow with Deschampsia caespitosa, Phleum
Taimyr [Osipov, 2003] is misidentification (material pratense, Epilobium angustifolium, etc., 4.–6.VIII.2009, leg. V.P.
examined). PREVIOUS RECORDS. Srednyaya Bay, Murman-
RANGE. Palaearctic polyzonal. sk Field Station [Fedotov, 1912, as Erigone a. (White,
1852)]; Teriberka [Shokhin et al., 2004, as E. a.].
Ceratinella scabrosa (O. Pickard-Cambridge, 1871) COMMENTS. Teriberka is the northernmost known
distribution of the species.
MATERIAL. Intra: 1 (cid:2), Teriberka, Orlovka River valley, RANGE. European.
swampy meadow with Betula nana, Carex sp., Eriophorum sp.,
Vaccinium oxycoccus, Rubus chamaemorus, Menyanthes trifolia- Erigone atra Blackwall, 1833
ta, Andromeda polifolia, Sphagnum sp., 9.VII.2008, leg. L.R. &
V.S.; 1 (cid:2), Sphagnum-Carex bog in tundra, 3.VII.2008, leg. L.R. &
1912 Erigone atra — Fedotov: 456.
V.S.
MATERIAL. Anthro: 2 (cid:2)(cid:2), Teriberka, agricultural meadow
COMMENTS. Teriberka is the northernmost known
with Deschampsia caespitosa, Phleum pratense, Epilobium angus-
distribution of the species. tifolium, etc., 4.VII.2008, leg. L.R. & V.S.; 1 (cid:2), ruderal plant com-
RANGE. West Palaearctic polyzonal. munity with Poa sp., Deschampsia caespitosa, Epilobium angusti-
folium, etc., 3.VII.2008, leg. L.R. & V.S.; 3 (cid:1)(cid:1), 4 (cid:2)(cid:2), meadow in
Cnephalocotes obscurus (Blackwall, 1834) village with Calamagrostis sp., Epilobium angustifolium, Matri-
caria recutita, Taraxacum sp., 11.VII.2008, leg. L.R. & V.S.; 1 (cid:2),
agricultural meadow with Deschampsia caespitosa, Phleum prat-
2004 Cnephalocotes obscurus — Shokhin et al.: 143, exam-
ense, Epilobium angustifolium, etc., 4–6.VIII.2009, leg. V.P.
ined. PREVIOUS RECORDS. Olen’ya Bay, Srednyaya
MATERIAL. Anthro: 2 (cid:1)(cid:1), Teriberka, Betula tortuosa under-
Bay [Fedotov, 1912].
growth on airfield with Empetrum sp., Vaccinium myrtillus, Pleuro-
zium schreberi, Hylocomium sp., etc., 4–6.VIII.2009, leg. V.P. RANGE. Holarctic polyzonal.
On spiders from the tundra zone of the Kola Peninsula 45
Erigone tirolensis L. Koch, 1872 MATERIAL. Intra: 1 (cid:1), 2 (cid:2)(cid:2), Teriberka, Orlovka River
valley, elfin Betula sp. sparse forest on gentle slope with Vaccini-
um myrtillus, Gymnocarpium dryopteris, Cornus suecica, no moss
1912 Erigone tirolensis — Fedotov: 454.
or lichen, 9.VII.2008, leg. L.R. & V.S.; 2 (cid:1)(cid:1), 3 (cid:2)(cid:2), Orlovka River
REMARKS. This species is absent from the materi-
valley, elfin Betula sp. sparse forest on gentle slope with Vaccini-
al examined.
um myrtillus, Gymnocarpium dryopteris, Cornus suecica, no moss
PREVIOUS RECORDS. Olen’ya Bay [Fedotov, or lichen, 9.VII.2008, leg. L.R. & V.S.; 1 (cid:1), 8 (cid:2)(cid:2), Sphagnum-
1912]. Carex bog in tundra, 3.VII.2008, leg. L.R. & V.S.; 2 (cid:2)(cid:2), Dal’niye
RANGE. Holarctic arcto-alpine. Zelentsy, sedge-sphagnum peatbog, pitfall traps, 4–26.VII.2009,
leg. A.B.; 1 (cid:1), sedge-sphagnum peatbog, pitfall traps, 4–
26.VII.2009, leg. A.B.; 3 (cid:1)(cid:1), 1 (cid:2), swampy bank of lake, pitfall
Gonatium rubens (Blackwall, 1833) traps, 4–26.VII.2009, leg. A.B.; 2 (cid:1)(cid:1), meadow with Geranium
sp. in Betula sp. & Salix sp. stands, pitfall traps, 13–26.VII.2009,
MATERIAL. Zonal: 1 (cid:2), Teriberka, near Tretie Titovskoye leg. A.B. Azonal: 2 (cid:1)(cid:1), 2 (cid:2)(cid:2), Dal’niye Zelentsy, meadow-like
Lake, dwarf birch-lichen mountain tundra with Betula nana, Cla- community on slope to sea, pitfall traps, 4–26.VII.2009, leg. A.B.
donia sp., 4–6.VIII.2009, leg. V.P.; 1 (cid:1), 3 (cid:2)(cid:2), Dal’niye Zelentsy, Anthro: 1 (cid:2), Teriberka, agricultural meadow with Deschampsia
lichen tundra on top of hill, pitfall traps, 4–26.VII.2009, leg. A.B.; caespitosa, Phleum pratense, Epilobium angustifolium, etc.,
1 (cid:1), 5 (cid:2)(cid:2), lichen tundra on slope of hill, pitfall traps, 4–26.VII.2009, 4.VII.2008, leg. L.R. & V.S.; 1 (cid:1), 13 (cid:2), ruderal plant community
leg. A.B.; 1 (cid:2), tundra with Betula nana, Vaccinium myrtillus, with Poa sp., Deschampsia caespitosa, Epilobium angustifolium,
Empetrum sp. on slope of hill, pitfall traps, 4–26.VII.2009, leg. etc., 3.VII.2008, leg. L.R. & V.S.
A.B. Intra: 1 (cid:1), Dal’niye Zelentsy, meadow with Geranium sp. in PREVIOUS RECORDS. Murmansk Field Station
Betula sp. & Salix sp. stands, pitfall traps, 13–26.VII.2009, leg. [Fedotov, 1912, as Dicyphus b.]; Teriberka [Shokhin
A.B. Azonal: 1 (cid:1), meadow-like community on slope to sea, pitfall et al., 2004].
traps, 4–26.VII.2009, leg. A.B. Anthro: 1 (cid:2), Teriberka, Betula
RANGE. Palaearctic polyzonal.
tortuosa undergrowth on airfield with Empetrum sp., Vaccinium
myrtillus, Pleurozium schreberi, Hylocomium sp., etc., 5.VII.2008,
leg. L.R. & V.S.; 1 (cid:2), meadow-like community on wasteland with Hypselistes jacksoni (O. Pickard-Cambridge, 1902)
Epilobium angustifolium, Calamagrostis sp., Polytrichum sp.,
7.VII.2008, leg. L.R. & V.S.; 1 (cid:2), stony wasteland with Deschamp- MATERIAL. Zonal: 1 (cid:2), Dal’niye Zelentsy, lichen tundra on
sia caespitosa, Phleum pratense, Empetrum sp. & Cladonia sp. top of hill, pitfall traps, 4–26.VII.2009, leg. A.B.; 2 (cid:2)(cid:2), lichen tundra
plots, 4–6.VIII.2009, leg. V.P. on slope of hill, pitfall traps, 4–26.VII.2009, leg. A.B.; 3 (cid:2)(cid:2), tundra
RANGE. Palaearctic polyzonal.
with Betula nana, Vaccinium myrtillus, Empetrum sp. on slope of
hill, pitfall traps, 4–26.VII.2009, leg. A.B. Intra: 2 (cid:1)(cid:1), Teriberka,
Gongylidiellum latebricola (O. Pickard-Cambridge, Sphagnum-Carex bog in tundra, 3.VII.2008, leg. L.R. & V.S.
1871) RANGE. Palaearctic-W-Nearctic boreo-nemoral.
MATERIAL. Intra: 1 (cid:1), 1 (cid:2), Teriberka, Orlovka River valley, Kaestneria pullata (O. Pickard-Cambridge, 1863)
swampy meadow with Betula nana, Carex sp., Eriophorum sp.,
Vaccinium oxycoccus, Rubus chamaemorus, Menyanthes trifoliata, MATERIAL. Intra: 1 (cid:2), Teriberka, Sphagnum-Carex bog in
Andromeda polifolia, Sphagnum sp., 9.VII.2008, leg. L.R. & V.S.
tundra, 3.VII.2008, leg. L.R. & V.S.
COMMENTS. Teriberka is the northernmost known
RANGE. Holarctic polyzonal.
distribution of the species.
RANGE. European polyzonal.
Leptorhoptrum robustum (Westring, 1851)
Hilaira herniosa (Thorell, 1875)
2004 Leptorhoptrum robustum — Shokhin et al.: 143, exam-
ined.
2004 Hilaira herniosa — Shokhin et al.: 143, examined. MATERIAL. Intra: 1 (cid:1), Dal’niye Zelentsy, meadow with
MATERIAL. Intra: 1 (cid:2), Dal’niye Zelentsy, meadow with Ge- Geranium sp. in Betula sp. & Salix sp. stands, pitfall traps, 13–
ranium sp. in Betula sp. & Salix sp. stands, pitfall traps, 13– 26.VII.2009, leg. A.B.; 2 (cid:2)(cid:2), Salix sp. with Sphagnum sp. in
26.VII.2009, leg. A.B. depression on slope, pitfall traps, 4–26.VII.2009, leg. A.B. An-
RECORDS. Teriberka [Shokhin et al., 2004]. thro: 1 (cid:2), Teriberka, agricultural meadow with Deschampsia cae-
RANGE. Holarctic boreal. spitosa, Phleum pratense, Epilobium angustifolium, etc., 4–
6.VIII.2009, leg. V.P.; 1 (cid:2), agricultural meadow on airfield with
Deschampsia caespitosa & Epilobium angustifolium, 5.VII.2008,
Hilaira nubigena Hull, 1911
leg. L.R. & V.S.
PREVIOUS RECORDS. Teriberka [Shokhin et al.,
2004 Hilaira sp. — Shokhin et al.: 143, examined. 2004].
MATERIAL. Intra: 1 (cid:2), Teriberka, Sphagnum-Carex bog in COMMENTS. Teriberka is the northernmost known
tundra, 3.VII.2008, leg. L.R. & V.S.; 3 (cid:2)(cid:2), Dal’niye Zelentsy,
distribution of the species.
sedge-sphagnum peatbog, pitfall traps, 4–26.VII.2009, leg. A.B.
PREVIOUS RECORDS. Teriberka [Shokhin et al., RANGE. Palaearctic boreo-nemoral.
2004, as Hilaira sp.].
RANGE. Palaearctic-Alaskan boreal. Macrargus multesimus (O. Pickard-Cambridge,
1875)
Hypomma bituberculatum (Wider, 1834)
MATERIAL. Zonal: 1 (cid:2), Teriberka, dwarf birch-lichen moun-
1912 Dicyphus bituberculatus — Fedotov: 454. tain tundra with Betula nana, Cladonia sp., 6.VII.2008, leg. L.R.
2004 Hypomma bituberculatum — Shokhin et al.: 143, exam- & V.S.
ined. RANGE. Holarctic boreal.
46 A.V. Tanasevitch & L.B. Rybalov
Maso sundevalli (Westring, 1851) Mecynargus paetulus (O. Pickard-Cambridge, 1875)
MATERIAL. Intra: 1 (cid:2), Teriberka, Orlovka River valley, elfin MATERIAL. Anthro: 1 (cid:1), 1 (cid:2), Teriberka, agricultural mead-
Betula sp. sparse forest on gentle slope with Vaccinium myrtillus, ow with Deschampsia caespitosa, Phleum pratense, Epilobium
Cornus suecica, Geranium sanguineum, Epilobium angustifolium, angustifolium, etc., 4–6.VIII.2009, leg. V.P.
Calamagrostis sp., 9.VII.2008, leg. L.R. & V.S.; 2 (cid:2)(cid:2), Sphagnum- RANGE. Holarctic boreo-nemoral.
Carex bog in tundra, 3.VII.2008, leg. L.R. & V.S. Anthro: 1 (cid:1),
Betula tortuosa undergrowth on airfield with Empetrum sp., Vac-
cinium myrtillus, Pleurozium schreberi, Hylocomium sp., etc., Mecynargus sphagnicola (Holm, 1939)
5.VII.2008, leg. L.R. & V.S.
COMMENTS. Teriberka is the northernmost known MATERIAL. Anthro: 1 (cid:2), Teriberka, ruderal plant community
distribution of the species. with Poa sp., Deschampsia caespitosa, Epilobium angustifolium,
RANGE. Holarctic polyzonal. etc., 3.VII.2008, leg. L.R. & V.S.
RANGE. Fennoscandian-Siberian-W-Nearctic bo-
real.
Mecynargus borealis (Jackson, 1930)
MATERIAL. Anthro: 1 (cid:1), 1 (cid:2), Teriberka, agricultural mead- Metopobactrus prominulus (O. Pickard-Cambridge,
ow on airfield with Deschampsia caespitosa & Epilobium angusti- 1872)
folium, 5.VII.2008, leg. L.R. & V.S.
RANGE. Holarctic boreal. MATERIAL. Zonal: 1 (cid:2), Teriberka, undershrub-moss tundra
with Empetrum sp., Rubus chamaemorus, Pleurozium schreberi,
Mecynargus monticola (Holm, 1943) 4–6.VIII.2009, leg. V.P.; 1 (cid:1), Dal’niye Zelentsy, lichen tundra on
top of hill, pitfall traps, 4–26.VII.2009, leg. A.B.
RANGE. Holarctic polyzonal.
MATERIAL. Zonal: 6 (cid:2)(cid:2), Teriberka, near Titovskoye Lake,
dwarf birch-lichen mountain tundra with Betula nana, Cladonia
sp., 4–6.VIII.2009, leg. V.P. Intra: 2 (cid:2)(cid:2), Carex-Sphagnum-Erio- Micrargus herbigradus (Blackwall, 1854)
phorum bog with Salix sp. stands, 4–6.VIII.2009, leg. V.P.
RANGE. Palaearctic-W-Nearctic boreal. MATERIAL. Anthro: 2 (cid:2)(cid:2), Teriberka, ruderal plant commu-
nity with Poa sp., Deschampsia caespitosa, Epilobium angustifo-
Mecynargus morulus (O. Pickard-Cambridge, 1873) lium, etc., 3.VII.2008, leg. L.R. & V.S.
COMMENTS. Teriberka is the northernmost known
distribution of the species.
2004 Semljicola faustus — Shokhin et al.: 143, examined.
MATERIAL. Zonal: 1 (cid:2), Teriberka, undershrub-moss tundra RANGE. Palaearctic boreo-nemoral.
with Empetrum sp., Rubus chamaemorus, Pleurozium schreberi,
4.VII.2008, leg. L.R. & V.S.; 1 (cid:1), 1 (cid:2), undershrub-lichen mountain Minicia marginella (Wider, 1834)
tundra with Betula nana, Vaccinium myrtillus, Empetrum sp., Pleu-
rozium schreberi, 7.VII.2008, leg. L.R. & V.S.; 1 (cid:1), 1 (cid:2), dwarf
MATERIAL. Anthro: 1 (cid:2), Teriberka, Betula tortuosa under-
birch-lichen mountain tundra with Betula nana, Cladonia sp.,
12.VII.2008, leg. L.R. & V.S.; 2 (cid:1)(cid:1), 2 (cid:2)(cid:2), undershrub-moss-lichen growth on airfield with Empetrum sp., Vaccinium myrtillus, Pleu-
rozium schreberi, Hylocomium sp., etc., 4–6.VIII.2009, leg. V.P.
mountain tundra with Empetrum sp., Vaccinium myrtillus, Pleuro-
COMMENTS. Teriberka is the northernmost known
zium schreberi, Dicranum sp., Cladonia sp., 10.VII.2008, leg. L.R.
& V.S.; 1 (cid:2), undershrub-lichen mountain tundra with Empetrum sp., distribution of the species.
Arctous alpina, Cladonia sp., no moss or lichen, 9.VII.2008, leg. RANGE. Palaearctic polyzonal.
L.R. & V.S.; 4 (cid:1)(cid:1), 3 (cid:2)(cid:2), near Titovskoye Lake, dwarf birch-lichen
mountain tundra with Betula nana, Cladonia sp., 4–6.VIII.2009,
leg. V.P.; 1 (cid:1), undershrub-moss-lichen mountain tundra with Betu- Minyrioloides trifrons (O. Pickard-Cambridge,
la nana, Empetrum sp., Pleurozium schreberi, Cladonia sp., Cetrar- 1863)
ia sp., 4–6.VIII.2009, leg. V.P.; 3 (cid:1)(cid:1), 4 (cid:2)(cid:2), near Tretie Titovskoye
Lake, dwarf birch-lichen mountain tundra with Betula nana, Cla- MATERIAL. Intra: 1 (cid:1), 2 (cid:2)(cid:2), Teriberka, Orlovka River val-
donia sp., 4–6.VIII.2009, leg. V.P.; 2 (cid:2)(cid:2), Sphagnum-Carex bog in
ley, elfin Betula sp. sparse forest on gentle slope with Vaccinium
tundra, 3.VII.2008, leg. L.R. & V.S. Intra: 4 (cid:2)(cid:2), Teriberka, Orlovka
myrtillus, Cornus suecica, Geranium sanguineum, Epilobium an-
River valley, swampy meadow with Betula nana, Carex sp., Erio-
gustifolium, Calamagrostis sp., 9.VII.2008, leg. L.R. & V.S.
phorum sp., Vaccinium oxycoccus, Rubus chamaemorus, Menyan- COMMENTS. Teriberka is the northernmost known
thes trifoliata, Andromeda polifolia, Sphagnum sp., 9.VII.2008, leg.
L.R. & V.S. Anthro: 1 (cid:1), 1 (cid:2), Teriberka, agricultural meadow on distribution of the species.
airfield with Deschampsia caespitosa & Epilobium angustifolium, RANGE. Holarctic boreo-nemoral.
5.VII.2008, leg. L.R. & V.S.; 2 (cid:1)(cid:1), 2 (cid:2)(cid:2), meadow-like community
on wasteland with Epilobium angustifolium, Calamagrostis sp., Poly- Minyriolus pusillus (Wider, 1834)
trichum sp., 7.VII.2008, leg. L.R. & V.S.; 1 (cid:2), stony wasteland with
Deschampsia caespitosa, Phleum pratense, Empetrum sp. & Cla-
MATERIAL. Anthro: 1 (cid:2), Teriberka, ruderal plant community
donia sp. plots, 4–6.VIII.2009, leg. V.P.
PREVIOUS RECORDS. Teriberka [Shokhin et al., with Poa sp., Deschampsia caespitosa, Epilobium angustifolium,
etc., 3.VII.2008, leg. L.R. & V.S.; 1 (cid:2), Betula tortuosa under-
2004, as Semljicola faustus (O. Pickard-Cambridge,
growth on airfield with Empetrum sp., Vaccinium myrtillus, Pleu-
1900)]. rozium schreberi, Hylocomium sp., etc., 4–6.VIII.2009, leg. V.P.
COMMENTS. Teriberka is the northernmost known COMMENTS. Teriberka is the northernmost known
distribution of the species. distribution of the species.
RANGE. European boreo-nemoral. RANGE. Palaearctic boreo-nemoral.
On spiders from the tundra zone of the Kola Peninsula 47
Oedothorax gibbosus (Blackwall, 1841) Oryphantes angulatus (O. Pickard-Cambridge, 1881)
MATERIAL. Intra: 1 (cid:1), Teriberka, Carex-Sphagnum-Erio- 1912 Lepthyphantes sp. — Fedotov: 459, pl. 8: 8, 9.
phorum bog with Salix sp. stands, Comarum palustre, 6.VII.2008, 1913 L. murmanicola Strand: 42, syn.n.
leg. L.R. & V.S. 1942 L. obscurus Fedotov. — Roewer: 551, incorrect citation.
COMMENTS. Teriberka is the northernmost known 2004 L. angulatus — Shokhin et al.: 143, examined.
distribution of the species. MATERIAL. Intra: 3 (cid:2)(cid:2), Teriberka, elfin Betula sp. sparse
forest on hill slope with Vaccinium myrtillus, Gymnocarpium dry-
RANGE. West Palaearctic polyzonal.
opteris, Cornus suecica, Calamagrostis sp., no moss or lichen,
10.VII.2008, leg. L.R. & V.S.; 1 (cid:2), Orlovka River valley, swampy
Oedothorax retusus (Westring, 1851) meadow with Betula nana, Carex sp., Eriophorum sp., Vaccinium
oxycoccus, Rubus chamaemorus, Menyanthes trifoliata, Androme-
da polifolia, Sphagnum sp., 9.VII.2008, leg. L.R. & V.S.; 2 (cid:2)(cid:2),
1912 Kulczynskiellum retusum — Fedotov: 454.
Sphagnum-Carex bog in tundra, 3.VII.2008, leg. L.R. & V.S.; 1 (cid:1),
2004 Oedothorax apicatus — Shokhin et al.: 143, examined.
MATERIAL. Zonal: 1 (cid:1), Teriberka, undershrub-moss tundra Dal’niye Zelentsy, swampy bank of lake, pitfall traps, 4–
26.VII.2009, leg. A.B.
with Empetrum sp., Rubus chamaemorus, Pleurozium schreberi,
4.VII.2008, leg. L.R. & V.S.; 2 (cid:2)(cid:2), Dal’niye Zelentsy, lichen tundra PREVIOUS RECORDS. Rybachiy Peninsula, Zem-
on slope of hill, pitfall traps, 4–26.VII.2009, leg. A.B. Intra: 1 (cid:1), lyanoye [Fedotov, 1912, as Lepthyphantes sp.]; Teri-
Teriberka, Orlovka River valley, swampy meadows with Betula nana, berka [Shokhin et al., 2004, as L. a.].
Carex sp., Eriophorum sp., Vaccinium oxycoccus, Rubus TAXONOMIC REMARKS. In his paper on the
chamaemorus, Menyanthes trifoliata, Andromeda polifolia, Sphag-
spiders of Murmansk Area, Fedotov [1912] reported
num sp. 10.VII.2008, leg. L.R. & V.S.; 2 (cid:1)(cid:1), 1 (cid:2), Dal’niye Ze-
on and illustrated a single female of Lepthyphantes sp.
lentsy, flood peddle bank of river, pitfall traps, 8–20.VII.2009, leg.
A.B.; 1 (cid:1), swampy bank of lake, pitfall traps, 4–26.VII.2009, leg. from Rybachiy Peninsula and noted that the species
A.B.; 4 (cid:2)(cid:2), Salix sp. with Sphagnum sp. in depression on slope, would probably be a new species similar to L. anguli-
pitfall traps, 4–26.VII.2009, leg. A.B. Azonal: 1 (cid:2), Dal’niye Ze- palpis (Westring, 1851). Later on, this species was
lentsy, meadow-like plant community on slope to sea, pitfall traps,
4–14.VII.2009, leg. A.B. Anthro: 1 (cid:1), 4 (cid:2)(cid:2), Teriberka, ruderal named by Strand [1913] as L. murmanicola Strand,
plant community with Poa sp., Deschampsia caespitosa, Epilobium 1913. According to the original figures by Fedotov
angustifolium, etc., 3.VII.2008, leg. L.R. & V.S.; 1 (cid:1), 1 (cid:2), agricul- [1912: pl. 8: 8, 9], the latter author unambiguously
tural meadow with Deschampsia caespitosa, Phleum pratense, Epi- dealt with a female of Oryphantes angulatus, and there-
lobium angustifolium, etc., 4.VII.2008, leg. L.R. & V.S.; 2 (cid:2)(cid:2),
fore it is safe to conclude that the name L. murmanico-
agricultural meadow on airfield with Deschampsia caespitosa &
Epilobium angustifolium, 5.VII.2008, leg. L.R. & V.S.; 1 (cid:2), meadow la is to be treated as its junior synonym (syn.n.). The
in village with Calamagrostis sp., Epilobium angustifolium, Matri- findings of O. angulatus in the regions close to the L.
caria recutita, Taraxacum sp., 11.VII.2008, leg. L.R. & V.S. murmanicola type locality [see Shokhin et al., 2004
PREVIOUS RECORDS. Pala Bay [Fedotov, 1912, and present paper] further support this conclusion.
as Kulczynskiellum r.]; Teriberka [Shokhin et al., 2004, COMMENTS. Rybachiy Peninsula is the northern-
as Oedothorax apicatus (Blackwall, 1850)]. most known distribution of the species.
RANGE. Palaearctic polyzonal. RANGE. European boreo-nemoral.
Oreoneta sinuosa (Tullgren, 1955) Pelecopsis mengei Simon, 1926
1912 Hilaira frigida — Fedotov: 457. 2004 Pelecopsis elongata — Shokhin et al.: 143, examined.
2004 Hilaira tatrica — Shokhin et al.: 143, examined. MATERIAL. Intra: 2 (cid:1)(cid:1), 8 (cid:2)(cid:2), Teriberka, Orlovka River
MATERIAL. Anthro: 1 (cid:2), Teriberka, stony wasteland with valley, elfin Betula sp. sparse forest on gentle slope with Vaccini-
Deschampsia caespitosa, Phleum pratense, Empetrum sp. & Cla- um myrtillus, Cornus suecica, Geranium sanguineum, Epilobium
donia sp. plots, 5.VII.2008, leg. L.R. & V.S.; 1 (cid:2), agricultural angustifolium, Calamagrostis sp., 9.VII.2008, leg. L.R. & V.S.; 1
meadow on airfield with Deschampsia caespitosa & Epilobium (cid:2), Orlovka River valley, elfin Betula sp. sparse forest on gentle
angustifolium, 5.VII.2008, leg. L.R. & V.S.; 1 (cid:1), stony wasteland slope with Vaccinium myrtillus, Gymnocarpium dryopteris, Cor-
with Deschampsia caespitosa, Phleum pratense, Empetrum sp. & nus suecica, no moss or lichen, 9.VII.2008, leg. L.R. & V.S.; 1 (cid:2),
Cladonia sp. plots, 4–6.VIII.2009, leg. V.P. Sphagnum-Carex bog in tundra, 3.VII.2008, leg. L.R. & V.S.; 1 (cid:2),
PREVIOUS RECORDS. Srednyaya Bay [Fedotov, Orlovka River valley, swampy meadows with Betula nana, Carex
1912, as Hilaira frigida (Thorell, 1872)]. Teriberka sp., Eriophorum sp., Vaccinium oxycoccus, Rubus chamaemorus,
[Shokhin et al., 2004, as H. tatrica (cid:1)(cid:2)(cid:3)(cid:4)(cid:5)(cid:6)(cid:7)(cid:8)(cid:9)(cid:10), 1915]. Menyanthes trifoliata, Andromeda polifolia, Sphagnum sp.
10.VII.2008, leg. L.R. & V.S.; 1 (cid:2), Dal’niye Zelentsy, sedge-sphag-
COMMENTS. Teriberka is the northernmost known
num peatbog, pitfall traps, 4–26.VII.2009, leg. A.B.; 2 (cid:2)(cid:2), meadow
distribution of the species.
with Geranium sp. in Betula sp. & Salix sp. stands, pitfall traps, 13–
RANGE. Fennoscandian. 26.VII.2009, leg. A.B.; 1 (cid:1), Betula sp. sparse forest, pitfall traps, 8–
26.VII.2009, leg. A.B. Azonal: 3 (cid:2), Dal’niye Zelentsy, meadow-like
Oreonetides vaginatus (Thorell, 1872) community on slope to sea, pitfall traps, 4–26.VII.2009, leg. A.B.
Anthro: 1 (cid:2), Teriberka, Betula tortuosa undergrowth on airfield
with Empetrum sp., Vaccinium myrtillus, Pleurozium schreberi,
1912 Oreonetides vaginatus — Fedotov: 458. Hylocomium sp., etc., 5.VII.2008, leg. L.R. & V.S.; 1 (cid:1), 1 (cid:2),
2004 Saaristoa abnormis — Shokhin et al.: 143, examined. ruderal plant community with Poa sp., Deschampsia caespitosa,
PREVIOUS RECORDS. Olen’ya Bay, Pala Bay Epilobium angustifolium, etc., 3.VII.2008, leg. L.R. & V.S.
[Fedotov, 1912]; Teriberka [Shokhin et al., 2004, as PREVIOUS RECORDS. Teriberka [Shokhin et al.,
Saaristoa abnormis (Blackwall, 1841)]. 2004, as Pelecopsis elongata (Wider, 1834)].
RANGE. Holarctic boreal. RANGE. Holarctic polyzonal.
48 A.V. Tanasevitch & L.B. Rybalov
Pocadicnemis pumila (Blackwall, 1841) Eriophorum bog with Salix sp. stands, 4–6.VIII.2009, leg. V.P.
Anthro: 1 (cid:2), Teriberka, agricultural meadow with Deschampsia
2004 Pocadicnemis pumila — Shokhin et al.: 143, examined. caespitosa, Phleum pratense, Epilobium angustifolium, etc., 4.–
MATERIAL. Intra: 5 (cid:1)(cid:1), 26 (cid:2)(cid:2), Teriberka, elfin Betula sp. 6.VIII.2009, leg. V.P.
sparse forest on hill slope with Vaccinium myrtillus, Gymnocarpi- RANGE. Fennoscandian-Siberian-W-Nearctic boreal.
um dryopteris, Cornus suecica, Calamagrostis sp., no moss or
lichen, 9–10.VII.2008, leg. L.R. & V.S.
Silometopus reussi (Thorell, 1871)
PREVIOUS RECORDS. Teriberka [Shokhin et al.,
2004].
MATERIAL. Anthro: 1 (cid:1), 7 (cid:2)(cid:2), Teriberka, agricultural mead-
COMMENTS. Teriberka is the northernmost known
ow with Deschampsia caespitosa, Phleum pratense, Epilobium
distribution of the species. angustifolium, etc., 4.VII.2008, leg. L.R. & V.S.
RANGE. Holarctic polyzonal. COMMENTS. This is the northernmost known dis-
tribution of the species.
Poeciloneta variegata (Blackwall, 1841) RANGE. Palaearctic boreo-nemoral.
1912 Poeciloneta variegata — Fedotov: 458.
Sisicus apertus (Holm, 1939)
REMARKS. This species is absent from the materi-
al examined.
MATERIAL. Zonal: 1 (cid:2), Teriberka, undershrub-lichen moun-
PREVIOUS RECORDS. Murmansk Field Station
tain tundra with Betula nana, Vaccinium myrtillus, Empetrum sp.,
[Fedotov, 1912]. Pleurozium schreberi, 4–6.VIII.2009, leg. V.P.
COMMENTS. This is the northernmost known dis- COMMENTS. This is the northernmost known dis-
tribution of the species. tribution of the species.
RANGE. Palaearctic-W-Nearctic polyzonal. RANGE. Holarctic boreo-nemoral.
Porrhomma convexum (Westring, 1851) Tapinocyba pallens (O. Pickard-Cambridge, 1872)
1912 Porrhomma norvegicum — Fedotov: 457. MATERIAL. Intra: 3 (cid:2)(cid:2), Teriberka, elfin Betula sp. sparse
REMARKS. This species is absent from the materi-
forest on hill slope with Vaccinium myrtillus, Gymnocarpium dry-
al examined. opteris, Cornus suecica, Calamagrostis sp., no moss or lichen,
PREVIOUS RECORDS. Pala Bay [Fedotov, 1912, 10.VII.2008, leg. L.R. & V.S.
as Porrhomma norvegicum Strand, 1901]. COMMENTS. This is the northernmost known dis-
COMMENTS. This is the northernmost known dis- tribution of the species.
tribution of the species. RANGE. European boreo-nemoral.
RANGE. European polyzonal. All records of P.
convexum in the Nearctic require confirmation. Tenuiphantes alacris (Blackwall, 1853)
Porrhomma egeria Simon, 1884 1912 Lepthyphantes alacris — Fedotov: 459.
2004 L. alacris — Shokhin et al.: 143, examined.
2004 Porrhomma convexum — Shokhin et al.: 143, examined. MATERIAL. Azonal: 1 (cid:1), 1 (cid:2), Dal’niye Zelentsy, meadow-like
PREVIOUS RECORDS. Teriberka [Shokhin et al., community on slope to sea, pitfall traps, 4–26.VII.2009, leg. A.B.
PREVIOUS RECORDS. Murmansk Field Station
2004, as Porrhomma convexum].
[Fedotov, 1912, as Lepthyphantes a.]; Teriberka
COMMENTS. This is the northernmost known dis-
[Shokhin et al., 2004, as L. a.].
tribution of the species.
COMMENTS. Teriberka is the northernmost known
RANGE. European boreo-nemoral.
distribution of the species.
RANGE. Palaearctic polyzonal.
Semljicola barbiger (L. Koch, 1879)
MATERIAL. Zonal: 1 (cid:2), Teriberka, undershrub-moss tundra Tenuiphantes nigriventris (L. Koch, 1879)
with Empetrum sp., Rubus chamaemorus, Pleurozium schreberi,
4–6.VIII.2009, leg. V.P. Intra: 1 (cid:1), Dal’niye Zelentsy, swampy MATERIAL. Intra: 1 (cid:1), Teriberka, Orlovka River valley,
bank of lake, pitfall traps, 4–26.VII.2009, leg. A.B.
elfin Betula sp. sparse forest on gentle slope with Vaccinium myr-
RANGE. Fennoscandian-Siberian arcto-boreal.
tillus, Cornus suecica, Geranium sanguineum, Epilobium angusti-
folium, Calamagrostis sp., 9.VII.2008, leg. L.R. & V.S.; 1 (cid:2),
Semljicola faustus (O. Pickard-Cambridge, 1900) Orlovka River valley, elfin Betula sp. sparse forest on gentle slope
with Vaccinium myrtillus, Gymnocarpium dryopteris, Cornus sueci-
MATERIAL. Azonal: 1 (cid:1), Dal’niye Zelentsy, meadow-like ca, no moss or lichen, 9.VII.2008, leg. L.R. & V.S.
community on slope to sea, pitfall traps, 4–26.VII.2009, leg. A.B. COMMENTS. Teriberka is the northernmost known
COMMENTS. This is the northernmost known dis- distribution of the species.
tribution of the species. RANGE. Holarctic boreo-nemoral.
RANGE. Fennoscandian-Siberian boreal.
Tenuiphantes tenebricola (Wider, 1834)
Semljicola lapponicus (Holm, 1939)
MATERIAL. Anthro: 1 (cid:1), 1 (cid:2), Teriberka, ruderal plant com-
MATERIAL. Intra: 2 (cid:2)(cid:2), Teriberka, Sphagnum-Carex bog in munity with Poa sp., Deschampsia caespitosa, Epilobium angusti-
tundra, 4–6.VIII.2009, leg. V.P.; 2 (cid:1)(cid:1), 2 (cid:2)(cid:2), Carex-Sphagnum- folium, etc., 3.VII.2008, leg. L.R. & V.S.
On spiders from the tundra zone of the Kola Peninsula 49
COMMENTS. This is the northernmost known dis- Walckenaeria kochi (O. Pickard-Cambridge, 1872)
tribution of the species.
RANGE. Palaearctic boreo-nemoral. MATERIAL. Intra: 1 (cid:2), Teriberka, Carex-Sphagnum-Erio-
phorum bog with Salix sp. stands, 4–6.VIII.2009, leg. V.P.
COMMENTS. This is the northernmost known dis-
Tiso aestivus (L. Koch, 1872)
tribution of the species.
MATERIAL. Zonal: 1 (cid:1), Dal’niye Zelentsy, lichen tundra on RANGE. Palaearctic boreo-nemoral.
top of hill, pitfall traps, 4–26.VII.2009, leg. A.B.; Anthro: 1 (cid:2),
Teriberka, meadow-like community on wasteland with Epilobium Walckenaeria mitrata (Menge, 1868)
angustifolium, Calamagrostis sp., Polytrichum sp., 7.VII.2008,
leg. L.R. & V.S.
2004 Walckenaeria mitrata — Shokhin et al.: 143, examined.
RANGE. Holarctic polyzonal.
PREVIOUS RECORDS. Teriberka [Shokhin et al.,
2004].
Wabasso replicatus (Holm, 1950)
COMMENTS. This is the northernmost known dis-
tribution of the species.
MATERIAL. Intra: 1 (cid:1), Teriberka, Carex-Sphagnum-Erio-
RANGE. European boreo-nemoral.
phorum bog with Salix sp. stands, Comarum palustre, 6.VII.2008,
leg. L.R. & V.S.; 2 (cid:2)(cid:2), Carex-Sphagnum-Eriophorum bog with
Salix sp. stands, 4–6.VIII.2009, leg. V.P. Walckenaeria nudipalpis (Westring, 1851)
RANGE. West Palaearctic boreal.
2004 Walckenaeria nudipalpis — Shokhin et al.: 143, exam-
Walckenaeria capito (Westring, 1861) ined.
MATERIAL. Anthro: 2 (cid:2)(cid:2), Teriberka, ruderal plant commu-
nity with Poa sp., Deschampsia caespitosa, Epilobium angustifo-
2004 Walckenaeria sp. — Shokhin et al.: 143, examined.
MATERIAL. Anthro: 1 (cid:2), Teriberka, Betula tortuosa un- lium, etc., 3.VII.2008, leg. L.R. & V.S.
PREVIOUS RECORDS. Teriberka [Shokhin et al.,
dergrowth on airfield with Empetrum sp., Vaccinium myrtillus,
Pleurozium schreberi, Hylocomium sp., etc., 4–6.VIII.2009, leg. 2004].
V.P. COMMENTS. This is the northernmost known dis-
PREVIOUS RECORDS. Teriberka [Shokhin et al., tribution of the species.
2004, as Walckenaeria sp.]. RANGE. Palaearctic polyzonal.
COMMENTS. This is the northernmost known dis-
tribution of the species. Zornella cultrigera (L. Koch, 1879)
RANGE. Holarctic polyzonal.
2004 Erigoninae — Shokhin et al.: 143, examined.
Walckenaeria cuspidata Blackwall, 1833 MATERIAL. Intra: 1 (cid:1), 1 (cid:2), Teriberka, Orlovka River valley,
elfin Betula sp. sparse forest on gentle slope with Vaccinium myr-
tillus, Gymnocarpium dryopteris, Cornus suecica, no moss or li-
2004 Walckenaeria sp. — Shokhin et al.: 143, examined. chen, 9.VII.2008, leg. L.R. & V.S.; 2 (cid:2)(cid:2), Dal’niye Zelentsy, Betula
MATERIAL. Intra: 3 (cid:2)(cid:2), Dal’niye Zelentsy, meadow with
sp. sparse forest, pitfall traps, 8–26.VII.2009, leg. A.B. Anthro: 1
Geranium sp. in Betula sp. & Salix sp. stands, pitfall traps, 13– (cid:2), Teriberka, ruderal plant community with Poa sp., Deschampsia
26.VII.2009, leg. A.B. Azonal: 1 (cid:2), meadow-like community on
caespitosa, Epilobium angustifolium, etc., 3.VII.2008, leg. L.R. &
slope to sea, pitfall traps, 4–26.VII.2009, leg. A.B.
V.S.
PREVIOUS RECORDS. Teriberka [Shokhin et al.,
PREVIOUS RECORDS. Teriberka [Shokhin et al.,
2004, as Walckenaeria. sp.].
2004, as “Erigoninae”].
COMMENTS. Teriberka is the northernmost known
RANGE. Holarctic boreal.
distribution of the species.
RANGE. Holarctic polyzonal.
Fam. TETRAGNATHIDAE (1)
Walckenaeria karpinskii (O. Pickard-Cambridge,
Tetragnatha extensa (Linnaeus, 1758)
1873)
1912 Tetragnatha extensa — Fedotov: 461.
2004 Walckenaeria karpinskii — Shokhin et al.: 143, exam- REMARKS. This species is absent from the materi-
ined. al examined.
MATERIAL. Zonal: 1 (cid:1), Teriberka, near Titovskoye Lake,
PREVIOUS RECORDS. Olen’ya Bay [Fedotov,
dwarf birch-lichen mountain tundra with Betula nana, Cladonia sp.,
4–6.VIII.2009, leg. V.P.; 3 (cid:1)(cid:1), 4 (cid:2)(cid:2), near Tretie Titovskoye Lake, 1912].
dwarf birch-lichen mountain tundra with Betula nana, Cladonia sp., RANGE. Holarctic polyzonal.
4–6.VIII.2009, leg. V.P. 1 (cid:2), Dal’niye Zelentsy, lichen tundra on
slope of hill, pitfall traps, 4–26.VII.2009, leg. A.B.; 1 (cid:2), tundra with
Fam. ARANEIDAE (3)
Betula nana, Vaccinium myrtillus, Empetrum sp. on slope of hill,
pitfall traps, 4–26.VII.2009, leg. A.B. Anthro: 1 (cid:2), Teriberka, ruder-
al plant community with Poa sp., Deschampsia caespitosa, Epilobi- Araneus diadematus Clerck, 1757
um angustifolium, etc., 3.VII.2008, leg. L.R. & V.S.
PREVIOUS RECORDS. Teriberka [Shokhin et al., 1912 Araneus diadematus — Fedotov: 462.
2004]. REMARKS. This species is absent from the materi-
RANGE. Fennoscandian-Siberian-Nearctic boreal. al examined.
50 A.V. Tanasevitch & L.B. Rybalov
PREVIOUS RECORDS. Murmansk Field Station PREVIOUS RECORDS. Teriberka [Shokhin et al.,
[Fedotov, 1912]. 2004].
RANGE. Holarctic polyzonal. COMMENTS. A record of P. monticola (Clerck,
1757) most probably refers to P. agrestis. For details,
Larinioides cornutus (Clerck, 1757) see Zyuzin [1979].
RANGE. Palaearctic polyzonal.
1912 Araneus cornutus — Fedotov: 462.
REMARKS. This species is absent from the materi- Pardosa agricola (Thorell, 1856)
al examined.
PREVIOUS RECORDS. Olen’ya Bay [Fedotov, MATERIAL. Zonal: 1 (cid:2), Teriberka, undershrub-moss tundra
1912, as Araneus c.]. with Empetrum sp., Rubus chamaemorus, Pleurozium schreberi,
RANGE. Holarctic polyzonal. 4.VII.2008, leg. L.R. & V.S.
COMMENTS. This is the northernmost known dis-
tribution of the species.
Larinioides patagiatus (Clerck, 1757)
RANGE. European boreo-nemoral.
1912 Araneus patagiatus — Fedotov: 462.
REMARKS. This species is absent from the materi- Pardosa amentata (Clerck, 1757)
al examined.
PREVIOUS RECORDS. Murmansk Field Station, 1912 Lycosa saccata — Fedotov: 467.
2004 Pardosa amentata — Shokhin et al.: 142.
Pala Bay [Fedotov, 1912, as Araneus p.].
MATERIAL. Intra: 8 (cid:1)(cid:1), 7 (cid:2)(cid:2), Dal’niye Zelentsy, Salix sp.
RANGE. Holarctic polyzonal.
with Sphagnum sp. in depression on slope, pitfall traps, 4–
26.VII.2009, leg. A.B.; 1 (cid:1), swampy bank of lake, pitfall traps, 4–
Fam. LYCOSIDAE (15) 26.VII.2009, leg. A.B.; 2 (cid:2)(cid:2), Betula sp. sparse forest, pitfall traps,
8–26.VII.2009, leg. A.B.; 12 (cid:1)(cid:1), 2 (cid:2)(cid:2), sandy bank of lake, pitfall
traps, 8–16.VII.2009, leg. A.B. Anthro: 1 (cid:2), Teriberka, agricultur-
Alopecosa aculeata (Clerck, 1757)
al meadow with Deschampsia caespitosa, Phleum pratense, Epilo-
bium angustifolium, etc., 4.VII.2008, leg. L.R. & V.S.; 1 (cid:2), ruderal
1912 Tarentula aculeata — Fedotov: 464. plant community with Poa sp., Deschampsia caespitosa, Epilobi-
2004 T. aculeata — Shokhin et al.: 142. um angustifolium, etc., 3.VII.2008, leg. L.R. & V.S.
MATERIAL. Intra: 3 (cid:1)(cid:1), Dal’niye Zelentsy, Salix sp. with PREVIOUS RECORDS. Shalim Island, Pala Bay,
Sphagnum sp. in depression on slope, pitfall traps, 4–26.VII.2009, Srednyaya Bay, Tyuva Bay [Fedotov, 1912, as Lycosa
leg. A.B.
saccata (Linnaeus, 1758)], Teriberka [Shokhin et al.,
PREVIOUS RECORDS. Murmansk Field Station,
2004].
Tyuva Bay [Fedotov, 1912, as Tarentula a.]; Teriberka
COMMENTS. Shalim Island is the northernmost
[Shokhin et al., 2004, as T. a.].
known distribution of the species.
RANGE. Holarctic polyzonal.
RANGE. European boreo-nemoral.
Arctosa alpigena (Doleschall, 1852)
Pardosa atrata (Thorell, 1873)
1912 Arctosa alpigena — Fedotov: 466.
2004 Pardosa atrata — Shokhin et al.: 142.
2004 Tricca alpigena — Shokhin et al.: 142.
MATERIAL. Zonal: 1 (cid:2), Teriberka, undershrub-moss-lichen MATERIAL. Intra: 15 (cid:1)(cid:1), 14 (cid:2)(cid:2), Dal’niye Zelentsy, sedge-
sphagnum peatbog, pitfall traps, 4–26.VII.2009, leg. A.B.; 2 (cid:2)(cid:2),
mountain tundra with Betula nana, Empetrum sp., Pleurozium
swampy bank of lake, pitfall traps, 4–26.VII.2009, leg. A.B.; 4
schreberi, Cladonia sp., Cetraria sp. 4.VII.2008, leg. L.R. & V.S.;
1 (cid:2), near Tretie Titovskoye Lake, dwarf birch-lichen mountain (cid:1)(cid:1), 4 (cid:2)(cid:2), sedge-sphagnum peatbog, pitfall traps, 4–26.VII.2009,
leg. A.B.
tundra with Betula nana, Cladonia sp., 4–6.VIII.2009, leg. V.P.; 1
(cid:2), Dal’niye Zelentsy, lichen tundra on slope of hill, pitfall traps, 4– PREVIOUS RECORDS. Teriberka [Shokhin et al.,
11.VII.2009, leg. A.B. Intra: 5 (cid:1)(cid:1), 3 (cid:2)(cid:2), Dal’niye Zelentsy, 2004].
sedge-sphagnum peatbog, pitfall traps, 4–26.VII.2009, leg. A.B.; 1 COMMENTS. Teriberka is the northernmost known
(cid:1), 1 (cid:2), meadow with Geranium sp. in Betula sp. & Salix sp. distribution of the species.
stands, pitfall traps, 13–26.VII.2009, leg. A.B.; 2 (cid:1)(cid:1), 1 (cid:2), sedge-
RANGE. Palaearctic boreo-nemoral.
sphagnum peatbog, pitfall traps, 4–26.VII.2009, leg. A.B. Azonal:
2 (cid:1)(cid:1), 3 (cid:2)(cid:2), Dal’niye Zelentsy, meadow-like community on slope
to sea, pitfall traps, 4–26.VII.2009, leg. A.B. Anthro: 1 (cid:2), Teriber- Pardosa eiseni (Thorell, 1875)
ka, ruderal plant community with Poa sp., Deschampsia caespito-
sa, Epilobium angustifolium, etc., 3.VII.2008, leg. L.R. & V.S. 2004 Pardosa eiseni — Shokhin et al.: 142.
PREVIOUS RECORDS. Lodeynaya Bay [Fedotov, MATERIAL. Intra: 1 (cid:2), Dal’niye Zelentsy, meadow with Ge-
1912]; Teriberka [Shokhin et al., 2004, as Tricca a.]. ranium sp. in Betula sp. & Salix sp. stands, pitfall traps, 13–
RANGE. Holarctic polyzonal. 26.VII.2009, leg. A.B. Azonal: 1 (cid:2), meadow-like community on
slope to sea, pitfall traps, 4–14.VII.2009, leg. A.B.
PREVIOUS RECORDS. Teriberka [Shokhin et al.,
Pardosa agrestis (Westring, 1861)
2004].
COMMENTS. Teriberka is the northernmost known
2004 Pardosa monticola — Shokhin et al.: 142, not seen.
distribution of the species.
REMARKS. This species is absent from the materi-
RANGE. Palaearctic boreal.
al examined.