New synonyms and combinations in Asiatic Ardisia (Myrsinaceae)

16 Pages·1999·0.99 MB·English

by  Ming, Hu Chi

BLUMEA 44 (1999)391-406
New synonyms and combinations in Asiatic Ardisia
(Myrsinaceae)
Hu+Chi Ming
South ChinaInstituteofBotany,Academia Sinica,Guangzhou510650,People'sRepublicofChina
Summary
StudiesonthegenusArdisiaSw. forFlora Malesiana haveledtothereduction of95taxa tosynon-
ymy,sevenspeciestotheinfraspecific level:A.demissa Miq.var.bambusetorum (King& Gamble)
C.M.Hu,A.polystictaMiq.subsp.punctipelata(Merr.) C.M.Hu,A.ternatensisScheff.varf.orstenii
(Scheff.)C.M.Hu,A.darlingiiMerr.subsp.podadenia(Stone)C.M. Hu,A.imperialisK.Schum.
var.novoguineensis(Mez)C.M.Hu,A.pubicalyxMiq.var.collinsae(Fletcher)C.M.Hu,A.tahanica
King & Gamble subsp. eucalyptifolia (Stone)C.M.Hu, and two new combinations are made:
A.forbesiiS.Moore var.cataractorum (Stone)C.M.Hu,A.fulvaKing& Gamblevar.multiflora
(Stone)C.M.Hu.
Key words:Ardisia,newsynonyms, newcombinations.
Introduction
Ardisiais thelargest genusin the Myrsinaceae, andrichly represented inthetropics
andsubtropics. Inthelastcomprehensive monographofthefamily, Mez(1902)recog-
nized 179 species ofArdisiain theOldWorld, ofwhich 105 were represented inthe
FloraMalesianaarea.Sincethennumerousnewspecies havebeendescribedby King
& Gamble(1906),Ridley(1923) andMerrill(1907-1923). Inthe 1980s,Stone(1982,
1989, 1990) madegreatcontributionstoour understanding of thegenus. Herevised
theMyrsinaceae ofPeninsularMalaysia, and described37 species and3 varietiesof
ArdisiafromBorneo, Sulawesi,andNew Guinea,thereby increasing the numberof
Ardisiaspecies toabout300 withintheFloraMalesianaregion. But thewholeofthe
genusofthisarea has notbeen reviewed.Thegenuswasdividedby Mezinto 14sub-
genera,of which5 are limitedto theAmericas; theother 9 are confinedto theOld
World. His treatment was followedby subsequent authors. Generally speaking, it
works well, butthereare manyexceptions; for example,Ardisia cymosa Blumeis a
distinctspecies, but itwas described six timesas anewspecies by differentauthors
(Table 1).
As can be seen from this table, these mistakes are due to theindividual authors
who placed theplant in a wrongsubgenus or based their study on limitedmaterial
from a limitedarea; thus it was inevitablethat specific names were multiplied and
muchconfusionresulted. Recently, Ihavehadtheprivilege ofstudying thecomplete
materialinherbariaA,AAU, BM,C, E, K,L, SING, UandUS through thekindness
ofthedirectors, andinthestudy itwas foundnecessaryto makethefollowingreduc-
tions. Alltherecognized species are placed undertheappropriate subgenus and ar-
ranged alphabetically.
392 BLUMEA Vol. 44, No. 2, 1999
Table 1.
SSppeecciieessnnaammee PPllaacceedd iinn ssuubbggeennuuss RReeccoorrdd ooffddiissttrriibbuuttiioonn
AArrddiissiiaa ccyymmoossaa BBlluummee ((11882266)) AAkkoossmmooss((aaccce..ttooMMeezz)) JJaavvaa,,BBoorrnneeoo,,TThhaaiillaanndd
ssyynn..AA..cchhiinneennssiiss BBeenntthh.. ((11886611)) BBllaaddhhiiaa ((aaccce..ttooMMeezz)) SSCChhiinnaa ttooNNVViieettnnaamm
ssyynn..AA..ppeerraakkeennssiiss
KKiinngg&& GGaammbbllee ((11990022)) CCrriissppaarrddiissiiaa EEnnddeemmiiccttooPPeenniinnssuullaarrMMaallaayyssiiaa
ssyynn..AAJ ..ssuuffffrruuttiiccoossaa RRiiddll.. ((11990088)) TTiinnuuss EEnnddeemmiiccttooPPeenniinnssuullaarrMMaallaayyssiiaa
ssyynn..AA..ccaaggaayyaanneennssiissMMeerrrr..((11991122)) AAccaarrddiissiiaa EEnnddeemmiicc ttootthheePPhhiilliippppiinneess
ssyynn..AA..ssiinnggaappoorreennssiissRRiiddll.. ((11991188)) CCrriissppaarrddiissiiaa ((RRiiddlleeyy)) EEnnddeemmiicc ttooSSiinnggaappoorree
AAkkoossmmooss((FFuurrttaaddoo,, SSttoonnee))
ssyynn..AA..uunndduullaattooddeennttaattaa
FFlleettcchheerr ((11993377)) BBllaaddhhiiaa EEnnddeemmiicc ttooTThhaaiillaanndd
Subgenus Acardisia Mez
1.Ardisia brachythyrsa Stapfvar. stellataStone
Ardisia brachythyrsa Stapfvar.stellata Stone,Proc.Acad. Nat. Sci. Philadelphia142(1990)28.
—Type: J.A.R.Anderson S30886 (holoE;iso L),Sarawak.
ArdisiakinabaluensisMerr.,Philipp. J.Sci.,Bot. 13(1918)114,syn.nov.—Type:Clemens10517
(n.v.),Borneo.
Distribution—Sabah, Sarawak(MtKinabalu).
Remarks —ArdisiakinabaluensisMerr.isidenticalwithA.brachythyrsa var. stel-
lata Stone.It differsfromA. brachythyrsa Stapf only by theslightly narrower sepals
andtheobscurely stellateindument.Thedistinctionsbetweenthe twoentities are too
slight forrecognition as separatespecies.
2.ArdisiacaudiferaMez
Ardisia caudiferaMez in Engl., Pflanzenr. 9(1902) 116.—Type:Beccari 1963 (holoP;iso K),
Borneo.
Ardisia subamplexicaulisMerr.,Philipp.J.Sci. 30 (1926)85,syn.nov.—Type:D.D. Wood 1283
(n.v.),Borneo.
Distribution—Sarawak, Sabah,KalimantanTimur.
Remarks —Theleafbases inA.caudifera vary fromroundedto cordulateormore
orless amplexicaul; thischaracterpreviously usedforthedistinctionofA.subamplexi-
caulisis unreliableforseparation.
3.ArdisialancifoliaMerr.
ArdisialancifoliaMerr.,Philipp. J.Sci., Bot. 13(1918)133.—Type:Clemens 10518(n.v.),Borneo.
Ardisia hewittiiFurtadoex Stone,Malay.Forest.45(1982)104,f. 1,syn.nov.—Type:Hewitts.n.
(holoSAR), Sarawak.
Ardisia miniscula Stone, Malay.Forest. 45 (1982) 106,f.2, syn.nov.—Type:J.A.R.Anderson
4524(holoSAR), Sarawak.
Distribution—Sarawak,Sabah, CentralKalimantan.
Hu Chi Ming: New synonyms and combinations in Asiatic Ardisia 393
Remarks — Ardisia hewittiiandA. miniscula are smaller plants in flower, with
smaller leaves andfewer-flowered inflorescences, but they merge intoand are not
clearlyseparable fromA. lancifolia Merr.
4.Ardisiaperissa Stone
Ardisiaperissa Stone,Proc.Acad. Nat.Sci.Philadelphia142(1990)37,f.11,12.—Type:J.A.R.
Anderson S 14871 (holoK; isoL), Sarawak.
Ardisia tryssa Stone, Proc.Acad. Nat. Sci.Philadelphia 142 (1990)42,f. 15,syn. nov.—Type:
Veldkamp8555 (holoL),Kalimantan.
Distribution Sarawak(Bau, Kuching, BukitManok); Kalimantan(Bukit Raya,
—
Gunung Sakoembang).
Remarks— When only a few specimens are compared, theremaybe seemingly
distinctivedifferences in the shape and margin ofleaves between A. perissa and
A. tryssa, but when more material is available for comparative study, the gradual
changes exhibitedinthesecharacterssuggestthatA. tryssa,characterizedbycoarsely
dentateleaves,ismerely ajuvenile stateofthesameplant.Aspecimen inBM(W.M.A.
Brook9898) hasboth deeply incisedandslightly dentateleaves onthe sameplant.
5.ArdisiavernicosaMez
Ardisia vernicosa Mez,Bull. Herb. Boiss. ser. 2,5 (1904)244. —Type:C. Hose 584 (isoE, K),
Sarawak.
Ardisia steiranthera Stone,Proc.Acad. Nat. Sci. Philadelphia 142 (1990) 32,f.8, syn.nov.—
Type:Ashton BRUN4(holoK; isoL),Borneo.
Distribution—Brunei, Sabah,KalimantanTimur.
Remarks—IndescribingA.steiranthera,Stonecompared itwitha groupofPhilip-
pine species, whichare quitedifferentandlater(1993) were describedas anewsub-
genus(Scherantha) by himself.Thetypeof/A.steirantheraisidenticaltoA. vernicosa
in every respect. The species is characterized by its coriaceous leaves with lateral
nerves meeting into a looped marginal vein, by puberulous, bipinnate-paniculate
inflorescence andtheovate-lanceolatesepals.
Subgenus Akosmos Mez
6.Ardisiacymosa Blume
Ardisia cymosa Blume,Bijdr. (1826)689. —Type:Blumes.n.(holoL,L930.255-123),Java.
Ardisia chinensis Benth.,Fl.Hongk.(1861)207,syn. nov.—Type:J.G.Champions.n.(holoK),
Hongkong.
ArdisiaperakensisKing& Gamble,J.Asiat.Soc.Bengal74 (1906) 147,syn. nov.—TVPe: Wray
3462 (holoK),Malaya.
Ardisia suffruticosa Ridl.,J. StraitsBranch Roy. Asiat. Soc. 50 (1908) 116,syn. nov.—Type:
Ridley 13010 (isoK), Malaya.
Ardisia cagayanensis Merr.,Philipp. J.Sci., Bot.7 (1912) 323,syn. nov.—Type: Catanduanes
FB 19620(isoK), Philippines.
Ardisia singaporensis Ridl.,J. Straits Branch Roy. Asiat. Soc. 79 (1918)94,syn. nov.—Type:
Ridley 2833 (lectoholohere designated,K;lectoiso SING),Singapore.
Ardisia undulatodentata Fletcher, Kew Bull. (1937) 39,syn. nov. —Type: Put 1631 (holoK),
Thailand.
394 BLUMEA Vol. 44, No. 2, 1999
Distribution—Thailand,Vietnam,China.InMalesia:Sumatra,PeninsularMalaysia,
Java,Borneo, Philippines.
Remarks — Although this species is widely distributed, its suffruticose habitis
consistentinall thespecimens. Someprevious authorsworkedonalimitedarea, and
othersputtheplantinwrongsubgenera duetolack ofauthenticmaterialforpurposes
ofcomparison, thus manysuperfluous nameshave beenpublished.
Subgenus Bladhia(Thunb.) Mez
7. ArdisiademissaMiq.
Ardisia demissaMiq.,Fl.Ind. Bat. 2(1858) 1018.—Type:Zollinger3509 (holoU; isoA,BM),
Java.
Ardisia metallica N.E.Br.,Illustr.Hort. 28 (1881)88,t.42,syn. nov.—Type:Linden s.n. (holo
K), Sumatra.
Ardisia odontophyllaWall, exDC. var.lobbiiC.B.Clarke,Fl. Brit.India 3(1882)526,syn.nov.
— Type:Lobb316(holoK), Malaya.
Ardisia odontophylla Wall, exDC. var.maingayi C.B.Clarke, Fl.Brit.India 3(1882)526, syn.
nov.—Type:Maingay 1016(holoK), Malacca.
Ardisia maingayi(Clarke) King& Gamble,J.Asiat.Soc. Bengal74 (1906)151,p.p.,syn.nov.
Ardisiacordulata Fletcher,Kew Bull.(1937)28,syn.nov.—Type:Kerr 7902 (holoK),Thailand.
ArdisiacordulataFletchervar.patulo-hirsutaFletcher,Kew Bull. (1937)29, syn.nov. —Type:
Put 3612 (holoK), Thailand.
Distribution— Thailand.InMalesia:Sumatra, PeninsularMalaysia, Java,Borneo.
Remarks — Mez(1902)considered.A. demissaataxonomicsynonymofA.fuligi-
nosa\ atthesametime,hecitedthespecimen Zollinger 3509underA. odontophylla,
unawarethat itis the typeofA. demissa. Thismistake was followedby subsequent
authorsandhas ledto manynewnamesbeen published.
8. Ardisia demissa Miq. var. bambusetorum(King & Gamble) C.M.Hu, stat. &
comb. nov.
Basionym:Ardisia bambusetorum King& Gamble,J.Asiat.Soc.Bengal74(1906) 153. —Type:
King'scollector 10095(holoCUL,n.v.),Malaya.
Distribution— PeninsularMalaysia (Perak).
Remark—Ardisiabambusetorumisglabrousthroughoutornearly so,butotherwise
scarcely differentfromvar. demissa.
9.Ardisia marginata Blume
ArdisiamarginataBlume,Bijdr. (1826)688. —Type:Blume s.n. (holoL,L900.211-144),Java.
ArdisiaapoensisElmer,Leafl.Philipp. Bot. 2(1910)663,syn.nov.—Type:Elmer 11890(isoL),
Philippines.
Distribution—Java,Borneo, Philippines.
Remark — Ardisia apoensis Elmer is known from only a few collections, and
scarcely differentfromA. marginata Blumeexceptforthepurplish blotchesonthe
leaves.
Hu Chi Ming:New synonyms and combinations in Asiatic Ardisia 395
Subgenus Crispardisia Mez
10.Ardisiapolysticta Miq.
Ardisiapolysticta Miq.,Fl.Ind.Bat.Suppl. 1(1861)576.—Type:Teijsmanns.n.(holoU), Sumatra.
Ardisia virens Kurz,For. Fl.Burma 2(1877)575. —Type:Griffith 3561 (iso K), EBengal.
ArdisiapardalinaMez inEngl.,Pflanzenr. 9(1902) 148,syn.nov. —Type: Warburg14361 (holo
Bf),Philippines.
Ardisiasinuato-crenata Mez in Engl.,Pflanzenr. 9(1902) 148,syn. nov.—Type:Haenke s.n.
(holoPRAG, n.v.),Philippines.
Ardisia jagoriiMez in Engl., Pflanzenr. 9(1902) 148,syn. nov.—Type: Jagor891 (holoBf),
Philippines.
Ardisia ramosii Merr.,Philipp.J.Sci.,Bot.7(1912)324,syn.nov.—Type:RamosBS13968 (iso
K),Philippines.
Distribution Widely distributedinSEAsiathroughPeninsularMalaysia andthe
—
Philippines toIndonesia.
Remarks — In herbaria,this species has long been identifiedas A. virens Kurz.
Afterexamining thetypeofA.polysticta Miq. inU,itturns out thatthetwo are iden-
tical,andtheearliernameA.polysticta mustbereinstated.Thespecies israthervariable,
butthevariationinsizeandmarginofleaves,andinthesizeof inflorescenceare con-
tinuousanditis difficultto findcharactersthatwill serve to dividethespecies.
11.Ardisiapolysticta Miq.subsp. punctipetala (Merr.)C.M. Hu,stat. & comb. nov.
Basionym:ArdisiapunctipetalaMerr.,Contrib. ArnoldArbor. 8(1934) 132.—Type:N.Walter&
C.M.Bangham796 (isoK), N Sumatra.
Distribution— Endemicto N Sumatra.
Remark—Differs fromvar. polysticta by itsmuch shortercalyx-lobes.
12. Ardisiarosea King & Gamble
Ardisia roseaKing& Gamble,J.Asiat.Soc.Bengal74 (1906)150.—Type;Wray402 (lectohere
designated,K), Malaya.
Ardisia longipedunculataKing& Gamble,J.Asiat.Soc. Bengal74(1906)150,syn. nov.—Type:
Wray296 (holoK; isoP),Malaya.
Distribution— Thailand.InMalesia: NSumatra,PeninsularMalaysia.
Remark— The rangeoflength ofthe peduncles found madeit impracticable to
maintainA. longipedunculata as aseparatespecies.
13.Ardisiasphenobasis Scheff.
ArdisiasphenobasisScheff.,Comm. Myrs.Archip. Ind.(1867)65.—Type:Korthalss.n. (holoL,
L900.211-185),Sumatra.
Ardisia wrayiKing& Gamble,J.Asiat.Soc.Bengal74 (1906)145,syn.nov.—Type:Wray3280
(lectohere designated,P),Malaya.
Distribution— NSumatra, PeninsularMalaysia (Perak, Johore,Selangor).
Remark— A distinct species characterizedby itslarge membranousleaves and
axillary racemose inflorescences.
396 BLUMEA—Vol. 44, No. 2, 1999
14. ArdisiavillosaRoxb.
Ardisia villosa Roxb.,Fl.Ind. ed.Carey 2(1824)274. —Type: Pulo-Pinang,India,notypema-
terialappears tohave beenpreserved.
Ardisia dolichosepalaMerr.,Philipp. J. Sci., Bot. 13 (1918) 112,syn. nov.—Type: Clemens
10819 (n.v.),Borneo.
Ardisia stenosepalaMerr.,Calif. Publ. Bot. 15(1929)236,syn.nov.—Type: Elmer21521 (n.v.),
Borneo.
Ardisiatawaensis Masam.,Enum.Phan. Bom.(1942)58,syn.nov.—Type:basedonA.stenosepala
Merr.
Distribution—Widely distributedinSEAsiathrough PeninsularMalaysiaandthe
Philippines to Indonesia.
Remarks — A very variable species in terms of density of indumentum.Both
A. dolichosepala andA. stenosepala are forms ofthis species, notableonly because
oftheslightly pubescent inflorescenceand nearly glabrous sepals.
15.Ardisia warburgiana Mez
Ardisia warburgianaMez in Engl.,Pflanzenr. 9(1902) 143.—Type:Warburg 14358 (holoBf),
Philippines.Neotype:Elmer 16471 (BM,K), Philippines.
ArdisiaproteifoliaMezinEngl.,Pflanzenr. 9(1902)143, syn.nov.—Type: Cuming 1867(holo
K),Philippines.
Distribution—Philippines.
Remarks — A transitionfromscallop-crenate leavesto coarsely serrate leaves is
evident, andthisvariationalsooccurs in otherspecies ofsubgenus Crispardisia, viz.
A. crenataSims, A.polysticta Miq.,etc.
Subgenus Pimelandra(A.DC.) Mez
16.Ardisiaapsidata Stone
Ardisia apsidata Stone,Proc.Acad. Nat. Sci. Philadelphia 142 (1990)22.—Type: Kostermans
10476 (holoL),Kalimantan.
Ardisiacosmetica Stone,Proc.Acad. Nat.Sci.Philadelphia142 (1990)25,f.3,syn.nov.—Type:
Kostermans 10477(holoL), Kalimantan.
Distribution—Kalimantan, Sarawak.
Remarks—Thetypespecimens ofA. apsidataandA. cosmeticawere collectedby
thesamecollectoratthesamelocality onthesameday withsuccessive fieldnumbers.
Thedifferencesintheleafmargins ofthetwotypesarenotconstantenough torecognize
two separatespecies.
17.Ardisiabrevipedata F. Muell.
ArdisiabrevipedataF.Muell.,Fragm.6(1868)163.—Type:Dallachys.n.,,AdSinumRockingham's
Bay,Australia(n.v.).
Ardisia apus Mez, Bot.Arch. 1(1922)256,syn. nov.—Type:Ledermann 12015 (holoBt),New
Guinea.
Distribution—Australia(Queensland); NewGuinea(Provinces; ESepik, Hollandia,
Morobe, MilneBay).
Hu Chi Ming:New synonyms and combinations in Asiatic Ardisia 397
Remark— Although thedistributionofA. brevipedata andA. apusare separated
bythe TorresStrait, no cleardistinctionsinthetwo entitiescanbemade.
18.Ardisiachrysophyllifolia King & Gamble
Ardisiachrysophyllifolia King& Gamble,J. Asiat. Soc.Bengal 74 (1906)123.—Type: Wray
1118(lectoheredesignated,K), Malaya.
ArdisiamysticaStone,Malay.Forest. 45 (1982) 114,f.5; Sida 16(1994)265, syn.nov.—Type:
Stone6546(holoKLU),Malaya.
Distribution PeninsularMalaysia.
—
Remarks—In describingA. mystica, Stonewas dealing with afruitingspecimen,
andhewasuncertainaboutitssubgeneric placement. Later(1994) henoticedthatitis
veryclosetoA.chrysophyllifolia, andemployed thelongerpedicel andtheminutely
tomentellousovary as principal criteria to separate it fromthat species. However,
manyspecies haveaccrescent pedicels whichmakesseparationby this characterun-
tenable.
19.Ardisiafuliginosa Blume
Ardisiafuliginosa Blume,Bijdr. (1825)692. —Type:Blumes.n. (holoL,L903.255-167),Java.
ArdisiafuliginosaBlume var.scopulata Stone,Proc.Acad. Nat. Sci. Philadelphia142(1990)26,
f. 4,syn.nov.—Type: H.O.Forbes 2142 (holoL), Sumatra.
Distribution—Sumatra, Java, Borneo,Lesser Sunda Islands.
Remark—Var.scopulata, withelongate racemoseinflorescences, isoneofnumer-
ous intermediatesandcannot reliably berecognized.
20.ArdisiaternatensisScheff.var. forstensii(Scheff.) C.M. Hu, comb. & stat.nov.
Basionym: Ardisiaforstensii Scheff.,Comm. Myrs.Archip.Ind.(1867)61. —Type: Forstens.n.
(holoL,L900.211-56),Celebes.
Distribution— EndemictoCelebes.
Remark— This variety differs from var. ternatensis by its shorter pedicels and
acute calyx-lobes.
21.ArdisiatomentosaC. Presl
Ardisia tomentosaC.Presl,Reliq. Haenk. 2(1835)66. —Type:Luzon,nospecimenwascited.
Ardisiaphilippinensis A.DC.inDC.,Prodr. 8(1844) 128,syn.nov.—Type:Cuming 1467(n.v.),
Philippines.
Ardisia cuprea Elmer,Leafl.Philipp. Bot. 8(1915)2777,syn.nov.—Type:Elmer 13287(iso C,
L),Philippines.
ArdisiapeninsulaElmer,Leafl.Philipp. Bot. 10(1939)3764,syn.nov.—Type: Elmer 16615(iso
C),Philippines.
Distribution—Sabah,Sarawak,Philippines.
Remarks—A variablespecies, showing variability insizeofleavesand density of
indumentum.Thespecies formerlyrecognized appeartobe merely formsintherange
ofgeneral variation,andtohavenotaxonomicsignificance.
398 BLUMEA Vol. 44, No. 2, 1999
Subgenus Pyrgus (Lour.) Mez
22.Ardisia darlingii Merr.
Ardisia darlingiiMerr.,Philipp. J. Sci., Bot. 5 (1910)215. —Type:DarlingFB 14611 (n.v.),
Philippines.
ArdisiagitingensisElmer,Leafl.Philipp.Bot. 4(1912)1498,syn.nov.—Type:Elmer 12430(iso
L,US),Philippines.
Distribution— Philippines (Luzon).
23.Ardisia darlingiiMerr. subsp. podadenia (Stone)C.M.Hu, comb.& stat. nov.
Basionym:ArdisiapodadeniaStone,Proc.Acad.Nat. Sci.Philadelphia 141(1989)300,f.23.—
Type: SAN25539(holoK; isoL), Borneo.
Distribution— Sabah(Sandakan District).
Remarks — Ardisiapodadenia Stone is similar in most respects toA. darlingii
Merr. from thePhilippines, butis widely separated fromit geographically. It might
well berecognized as asubspecies.
24. Ardisiapergamacea(Miq.) Mez
Ardisiapergamacea (Miq.) Mez in Engl.,Pflanzenr. 9(1902) 154.— Climacandra pergamacea
Miq.,Fl. Ind. Bat.Suppl. 1 (1861)576. — Type: Teijsmanns.n.(holoU), Sumatra.
ArdisiamerjimahStone,Proc.Acad.Nat.Sci.Philadelphia141(1989)277,f.4,syn.nov.—Type:
Sibu ForestOffice s.n. (holoSAR), Sarawak.
Distribution—Sumatra, Sarawak.
Remarks — Ardisiamerjimah Stone was originally placed insubgenus Akosmos,
butitseems tomethatitsaffinityis withthespecies ofsubgenus Pyrgus, especially
withA. darlingiisubsp. podadenia. Stoneregarded it as a distinctspecies mainly on
accountoftheleaveshavingshorthairson theunderside,thecalyx-lobes beingacute,
butthesecharactersareall identicalwithA.pergamacea(Miq.) Mez.
25.Ardisia serrata (Cav.) Pers.
Ardisia serrata (Cav.) Pers.,Syn. 1 (1805)233. —Anguillariaserrata Cav.,1c.6(1801)2,t. 503.
—Type: No specimenappears tohave been preserved; theplate must therefore serve asthe
type.
Ardisia angustifoliaA.DC. inDC., Prodr. 8(1844) 134,syn.nov.— Tinus candolleana Kuntze,
Revis.Gen. PI.2(1891)973.—Ardisiacandolleana (Kuntze)MezinEngl.,Pflanzenr.9(1902)
13,syn.nov.—Type:Cuming 1711 (iso BM,K), Philippines.
Ardisia whitfordiiMez, Philipp. J.Sci., Suppl. 1 (1906)271, syn. nov.— Type: Whitford 772
(n.v.),Philippines.
Ardisia mindorensis Merr.,Philipp. J. Sci.,Bot. 5 (1910)218, syn.nov. —Type:Merrill 6145
(n.v.),Philippines.
Ardisia curtipesMerr.,Philipp. J.Sci.,Bot. 5(1910)372,syn.nov.—Type: Curran, Merritt&
Zschokke 18137 (n.v.),Philippines.
Ardisia oliganthaElmer,Leafl.Philipp. Bot.4(1912)146,syn.nov.—Ardisia oligocarpaMerr.,
J.Arnold Arbor. 35(1954) 148,syn.nov.—Type:Elmer 12130 (iso L),Philippines.
Hu Chi Ming:New synonyms and combinations in Asiatic Ardisia 399
Ardisia dataensis Mez,Fedde Repert. 16(1920)414,syn. nov.—"type:MerrillBS 4496 (n.v.),
Philippines.
Ardisia abanii Stone,Proc.Acad. Nat. Sci.Philadelphia 142 (1990)53,f. 25,syn.nov.—Type:
Aban Gibot SAN99286 (isoL), Borneo.
Distribution Java,Borneo,Philippines.
—
Remarks— Apolymorphic ochlospecies. Fromplant toplant, or even withinthe
sameplantofdifferentage or vigor,thereis considerablevariationinthe size, shape
ofleaves,andin the densityofindumentum.Thesevariantsintergrade considerably
andshow nogeographical separation. They might wellbe regarded as a complex of
species, which havea distincttendency tomerge intoeachother, although extreme
formsare distinctenough.
Subgenus Stylardisia Mez
26.ArdisiaeximaMiq.
Ardisia exima Miq.,PI. Junghuhn.(1853) 196. —Type: Junghuhns.n. (holoL,L900.211-53),
Java orSumatra.
ArdisialampongaMiq., Fl.Ind.Bat.Suppl. 1 (1861)577,syn.nov.—Type: Teijsmanns.n.(holo
U),Sumatra.
Distribution—Sumatra,PeninsularMalaysia, Java, Borneo.
Remark—Ardisia lamponga Miq. resembles A. exima Miq. inall respects, and
there is no cleardistinctionbetweenthe two entities.
27.Ardisiaforbesii S. Moore
Ardisiaforbesii S. Moore,J.Bot. 52 (1914)291. —Type:H.O. Forbes 120 (lecto BM),New
Guinea.
Ardisia arborella Stone,Proc.Acad.Nat. Sci.Philadelphia 142(1990)44,f.16,syn.nov.—Type:
VanBalgooy3104 (holoPH;isoL), Sulawesi.
Distribution—Borneo, Celebes, NewGuinea.
Remarks—Althoughoriginallydescribedfrom NewGuinea,fairlyabundantmate-
rialshows thatitsdistributionextendsfromCelebestoBorneo.Inherbaria,thespeci-
mens collectedfromBorneoare commonly misidentifiedasA.synneura Scheff.
28.Ardisiaforbesiivar. cataractorum (Stone) C.M. Hu, comb. nov.
Basionym:Ardisia synneura Scheff. var.cataractorum Stone,Proc.Acad. Nat. Sci. Philadelphia
141(1989)290. —Type: P.Chai S18952 (holoSAR;isoA,K,L),Sarawak.
Distribution— Sarawak.
29.Ardisiamontana King & Gamble
Ardisia montanaKing & Gamble,J.Asiat.Soc.Bengal74 (1905) 128.—Type: Wray 327(lecto
here designated,K), Malaya.
Ardisia montana King& Gamble var.selangorensis Stone,TreeFl. Malaya4 (1989)274, syn.
nov.—Type:Stone 6877(holoSING),Malaya.
400 BLUMEA Vol. 44, No. 2, 1999
Distribution—PeninsularMalaysia.
Remark—A variablespecies intermsofleafshape and size.
30.Ardisia ophirensis (Clarke) Mez
Ardisia ophirensis (Clarke)Mez inEngl.,Pflanzenr. 9(1902)113.—Ardisia tuberculataA.DC.
var.ophirensis Clarke, Fl.Brit.India 3(1882)522. —Type:Hullet 754 (holoK; iso SING),
Malaya.
Ardisia labisiaefolia King& Gamble,J.Asiat.Soc.Bengal 74 (1906) 127,syn.nov. —Ardisia
ophirensis var.labisiaefolia(King& Gamble) Stone,Tree Fl.Malaya4(1989)273,syn.nov.
—Type: Wray1566(lectoholoheredesignated,K;lectoiso E),Malaya.
ArdisiapetricolaRidl.,J.Fed. Mai. St.Mus.6 (1915) 159,syn.nov. —Type:Ridley 16120 (holo
K), Malaya.
Ardisia andamanica Kurz var.latifolia Ridl.,Fl.Mai. Penins. 2(1923)245, syn. nov. —Type:
Ridleys.n. (iso SING),Malaya.
? ArdisiapraetermissaFurtado,Gard. Bull. Sing. 18(1958)283,syn.nov.—Type:Alvins679 =
342 (holoSING),Malaya.
Distribution— PeninsularMalaysia.
Remarks —Ardisiapraetermissa Furtadowas basedonapoorspecimenandorigi-
nally placed inthesubgenus Akosmos.Acareful studyontheholotype showsthat the
inflorescence,although rathershort,is clearlyterminal,andtheflowerwithlongexsert-
ed style. Thesecharacters indicate thatitis closely related with A. ophirensis, but
furthermaterialis necessary beforeitsstatus canbesettled.
31.Ardisia pterocaulis Miq.
Ardisia pterocaulis Miq.,PI. Junghuhn.(1853) 194.—Type: Junghuhns.n.(holoL,F900.211-
69),Sumatra.
Ardisia pterocaulis Miq. var.initida Scheff.,Comm. Myrs. Archip.Ind. (1867)80, syn. nov.—
Type:Korthalss.n.(holoL,L900.211-150),Sumatra.
Ardisiapterocaulis var.elongata Scheff.,Comm. Myrs.Archip.Ind.(1867)80,syn.nov.—Type:
Blumes.n. (holoF, L900.211-148),Java.
Ardisia synneuraScheff.,Comm. Myrs.Archip.Ind.(1867)80,syn.nov.—Tapeinospermasyn-
neura(Scheff.)Philipson, J. Bot. 77 (1939) 102, syn. nov.—Type: Korthals s.n. (holoF,
L900.211-191),Borneo.
Ardisiaplatyclada King& Gamble,J.Asiat.Soc.Bengal74 (1906)132,syn.nov.—Type: Scor-
techini 232 (holoK), Malaya.
Distribution—Sumatra,PeninsularMalaysia, Java,Borneo.
Remarks A distinctspecies characterized by itsbranchletsmarkedby flexuous
—
ridges, by the lateralnerves oftheleafforming alooped marginal vein, andby its
smallflowersandfruits.Ardisiasynneura Scheff.isknownonlyfromthetypematerial,
andis scarcely differentfromA.pterocaulis Miq.exceptfortheless scaly-pubescent
branchletsandleaves.
32.Ardisiasanguinolenta Blume
ArdisiasanguinolentaBlume,Bijdr. (1826) 685. —Type:Blume s.n.(holoL,L900.211-170),
Java.
Ardisiacolorata Roxb.,Fl.Ind. ed. Carey 2(1824)271 (nonFink, 1821),syn. nov.—Type: no
typewascited(Silhet).