New species of Paleogene cypraeoideans (Gastropoda) from the Pacific slope of North America

2011·5.6 MB·English

by  Lindsey T Groves

THE NAUTILUS 125(2):45-52, 2011 Page 45
New
species of Paleogene cypraeoideans (Gastropoda)
from the Pacific slope of North America
LindseyT. Groves
Natural HistoryMuseumofLosAngeles County
MalacologySection,900 Exposition Boulevard
Los Angeles, CA90007 USA
[email protected]
ABSTRACT ous authors, only a brief overview of the stratigraphic
nomenclature and agewillbe mentioned. Readers willbe
A new species of Bemaya sensu stricto (Cypraeidae) from referred toadditional sources fordetaileddescriptions.
EocenerocksofWashingtonand fournewspeciesofEocypraea
sensu stricto (Eocypraeidae), one from Paleocene strata of
northern California, two from Eocene strata of Washington,
and one from Baja California Sur, Mexico, are described. The Martinez Formation
new species ofBemaija sensu stricto and the Washington spe-
cies of Eocypraea sensu stricto represent the northernmost Whitney in Gabb (1869: xiii) provisionally proposed tbe
Cenozoie records for their respective genera in western North Martinez Group to “include a series of beds, of small
A(mLearmiacrac.k,A18t0e2n)t,atpirveeviroeucsolrydknoofwEnocoynplryaferaom(EtohceyLpurtaeetaia)ninSftlaagtea ogenogtrhaephniocratlheerxntefnlta,nkfooufnMdoanttMearDtiiabnleoz.”[CHaleifaolrsnoiai]ncaonrd-
(middle Eocene) of France, Belgium, and England, is noted rectly noted that the group may eventually prove to be
from the middle Eocene Domengine Formation of Kings worthyofrankingonlyas asubdivisionofthe Cretaceous
County,California. Chico Group. In the same volume, Gabb (1869: 129)
Additionalkeywords: Bemaya, Eocypraea, fossils,paleontology, referred to the Martinez Group as “the upperportion of
Paleocene, Eocene ‘Division A’ ofthe California reports.” These beds were
provisionally demonstrated to be “Paleocene” age by
Clark and Vokes (1936). Brice (1953) referred to Paleo-
cene strata in Lake County, California, as Martinez For-
mation. He noted that “fossils characteristic of the
INTRODUCTION Paleocene Martinez are found in scattered localities
through the sandstone, and the lithologic assemblage is
Five new species of Paleogene cypraeoideans are similar to that of the type Martinez formation.” There-
described from localities on the Pacific slope of North fore, for the lack ofa more appropriate name, the usage
America (Table 1). The new species of Bemaija sensu of Brice will be followed and these uppermost lower or
stricto represents the first record of this genus from lowermost upper Paleocene (Danian/Thanetian stages)
Washington and is the northernmost record of the beds in Lake County will be referred to as “Martinez”
genus in western North America. Four new species of Formation.
Eocypraea sensu stricto are described from strata in Lake
County, California, Thurston and Lewis counties, Wash-
ington, and Baja California Sur, Mexico. All four species Crescent Formation
represent first records from their respective regions
and the species from Washington are the northernmost The Crescent Formation of Arnold (1906: 460-461)
representatives of the genus in western North America. was described for a “series black basalt and greenish
These taxa are indicators of shallow, warm water depos- basalt tuffs and tuffaceous sands found in the vicinity of
itional environments. Port Crescent,” Clallam County, Washington. Weaver
(1937) referred to the Crescent Formation as middle
Eocene (“Capay” California provincial molluscan stage
STRATIGRAPHYAND GEOLOGIC AGES [CPMS]) tuffaceous shales and sandstones and basaltic
agglomerate of marine origin exposed on the northern
The formations listed below, from oldest to youngest, flank of the Olympic Peninsula. Recently, much confu-
are those from which the new species are described. sion has been ascribed to Eocene deposits near
Because these formations have been mentionedbyprevi- Maynard on Discoveiy Bay. Durham (1944) referred
B
)
Page 46 THE NAUTILUS, Vol. 125, No. 2
Table 1. List of the species of Bemaya (Protocypraea), Most recently Spencer (1984) mapped the area as
Bemaya sensu stricto, and Eocypraea sensu stricto from a “sedimentary member” ofthe Crescent Formation.
western North America and their generalized localities
(formation is included for new species).
Domengine Formation
Bemaya (Protocypraea) andBemaya sensustricto
Anderson (1905) described the “Domijean Sands” for
Late Cretaceous
Bemaya Protocypraea argonautica (Anderson, 1958) widespread outcrops on the north flank of Mt. Diablo,
[Jackso(nCo., Oregon)] Fresno County, California and the type locality is in the
B. (P.)berryessae (Anderson, 1958) [Yolo Co., California] NE14 of section 17, north of Coalinga. The molluscan
B. (P.)gualalaensis (Anderson, 1958) [MendocinoCo, faunas were described by Clark and Vokes (1936) and
California] Vokes (1939). These faunas became the basis of the
B. (P.)popenoei Groves, 2004 [Orange Co., California] “Domengine” [CPMS] ofClarkandVokes (1936), which
B. (P.) rineyi Groves, 1990 [San Diego Co., California] theybelievedtobe middle Eocene (Squires (1988). Most
B. (Bemaya)beardi Groves,2004 [VancouverId., British recently, based on calcareous nannoplankton and mag-
Columbia]
netic stratigraphy, Prothero (2001) considered the for-
B. (B.)crawfordcatei Groves, 1990 [San Diego Co., California] mation to be earlyto middle Eocene age.
B. ( .)jeanae Groves, 2004 [Butte Co., California]
Eocene
Bemaya (Protocypraea)grovesi Squires and Demetrion, 1992 Bateque Formation
[BajaCaliforniaSur, Mexico]
Bemaya (Bemaya)squiresi newspecies [Crescent Formation, The Bateque FormationofMina (1956, 1957) was named
Jefferson Co., Washington] for outcrops along the Pacific side of Baja California,
Eocypraea sensustricto Mexico, from the north end and east side of Laguna San
Late Cretaceous Ignacio to the San Juanico area about 105 km to the
Eocypraea (Eocypraea)louellae Groves, 1990 [YoloCo., south (Squires and Demetrion, 1992). The formation
California] ranges in age from middle early Eocene to late middle
Eocene (“Capay” [CPMS] through “Tejon” [CPMS])
Paleocene
Eocypraea Eocypraea novasumma (Nelson, 1925) [Ventura based on calcareous nannoplankton, planktonic foramini-
Co., Calif(ornia] ) fers, andmollusks (Squires and Demetrion, 1992).
Eocypraea (Eocypraea)takeosusukii newspecies [“Martinez”
Formation, Lake Co., California]
ABBREVIATIONS
Eocene
Eocypraea (Eocypraea)batequensis newspecies [Bateque Abbreviations used for institutional catalog and/or local-
Formation, BajaCaliforniaSur, Mexico] itynumbers, affiliations, and condensed terminology are
Eocypraea (Eocypraea)bayerquei (Gabb, 1864) [ContraCosta
as follows (unless indicated otherwise, collections are in
Co., California]
California): CIT, California Institute ofTechnology, Pas-
Eoc[yKperraneCao.(,EoCcaylipfroarenai)a]castacensis (Stewart, 1926 [1927]) adena (collections now at LACMIP); CPMS, California
Eocypraea (Eocypraea)crescentensis newspecies [Crescent provincial molluscan stage; CSUN, California State
Formation, ThurstonCo.,Washington] University, Nortlnidge (collections now at LACMIP);
Eocypraea (Eocypraea)jimgoederti newspecies [Crescent IGM, Instituto de Geologia Universidad Nacional
Formation, Lewis Co.,Washington] Autonoma de Mexico, Mexico Citv; LACMIP, Natural
Eocypraea (Eocypraea ef. E. (E.)inflate (Lamarck, 1802) Histoiy Museum of Los Angeles County, Invertebrate
[Kings Co., California] Paleontology Section; LACMVP, Natural History Museum
Eocypraea (Eocypraea)maniobraensis Squires andAdvocate, of Los Angeles County, Vertebrate Paleontology Sec-
1986 [RiversideCo., California] tion; UCLA, University of California, Los Angeles (col-
lections now at LACMIP); UCMP, University of
to these beds as Eocene basalts, sandstones, shales, California, Museum of Paleontology, Berkeley; USGS,
and small limestone lenses. Armentrout and Berta United States Geological Survey, Menlo Park.
(1977) mentioned Narizian/Refngian benthic foraminif- Measurement parameters are defined as follows:
eral stages (late middle to late Eocene) aged foraminif- length = greatest distance between anterior and poste-
era from a locality within the Townsend Shale Member riorends;width = greatestdistancebetween lateral mar-
of the Lyre Formation on the east side of Discovery gins; and height = greatest distance between base and
Bay. Tabor and Cady (1978) mapped outcrops on the dorsum.
west side of Discovery Bay as a “sandstone and minor
siltstone member” of the Lyre Formation, a younger SYSTEMATIC PALEONTOLOGY
unit than the Crescent Formation. Armentout and
others (1983) referred to these beds as “sandstone of The classification used here for Cypraeidae follows that
Maynard” and proposed that it interfingers with the of Schilder and Schilder (1971) and Fehse (2001) for
lower to middle Eocene upper Crescent Formation. Eocypraeidae.
1
L.T. Groves, 201 Page 47
SuperfamilyCypraeoidea Rafinesque, 1815 fied as B. (£>.) obesa, figured by Perrilliat and others
FamilyCypraeidae Rafinesque, 1815 (2003: 43, figs. 5-6) from the middle Eocene San Juan
Subfamily Bernayinae Schilder, 1927 Formation, central Chiapas, southern Mexico, lacks
Tribe Bernayini Schilder, 1927 enough shell material for adequate comparison to the
new species.
Genus Bemaya Jousseaume, 1884
Type Species: Cypraea media Deshayes, 1835, by Discussion: Post-burial crushing has damaged the an-
original designation. Upper middle Eocene (Bartonian terior portion of the aperture and dorso-ventrally dis-
torted the specimen Generic and subgeneric assignment
Stage), Auvers-sur-Oise, Val-d’Oise (northwest ofParis), .
are based on the wide aperture, deep anterior and poste-
France.
rior canals, and medium-height spire. Bemaya squiresi
Diagnosis: Shell medium to large size; anterior end represents the northernmost occurrence ofthe genus in
somewhat carinate; dorsum smooth; spire of medium Cenozoic strata and the only representative of the genus
height and partially covered; aperture wide, sides described from Washington.
rounded; anterior and posterior canals deep; fossula
smooth, concave, wide. Material: The new species is represented by a single
fairlywell preservedspecimen that exhibits original shell
Remarks: Schilder and Schilder (1971) recognized 22 material on the base and minor amounts oforiginal shell
species and 11 subspecies ofworldwide Bemaya sensu material on the dorsum.
stricto. Four of their subspecies have been raised to
specific status, one species and one subspecies have Type Material: Holotype LACMIP 13644, measures
mm mm mm
beenreassignedtoothergenera, three species havebeen 52.7 in length, 40.0 in width, and 26.3 in
described subsequent to 1971 (see Groves, 1990; 2004), height.
and another new species is described here. This brings
the present total to 29 species and six subspecies. Only Type Locality: LACMIP loc. 22341 [ex UCLA
four of these species are from western North America loc. 2341], southwest end of Discovery Bay, Jefferson
(Table 1). County, Washington. Middle lower Eocene (“Capay”
[CPMS] = Ypresian Stage), Crescent Formation.
Subgenus Bemaya Jousseaume, 1884
Bemaya (Bemaya)squiresi newspecies Etymology: This species is named after friend and
colleague Richard L. Squires (CSUN Geological Sci-
(Figures 1-2)
ences) for his extraordinary contributions to molluscan
B. (B.) n. sp. Groves, 1997: 7. paleontology.
Diagnosis: Bernaya of large size, anterior and poste- Family Eocypraeidae Schilder, 1924
rior canals deep, spire of medium height, fossula con- Subfamily Eocypraeinae Schilder, 1924
cave and smooth (although columellar dentition extends Tribe Eocypraeini Schilder, 1924
Genus Eocypraea Cossmann, 1903
slightly onto fossula), posterior terminal ridges extend to
margins.
Type Species: Cypraea inflata Lamarck, 1802, by ori-
Description: Shell of medium to large size, con- ginal designation, middle Eocene (Lutetian/Bartonian
stricted anteriorly; maximum height of shell nearly stages), Paris Basin, France.
centered; maximum width of shell slightly posterior of
Diagnosis: Inflated-pyriform shell ofsmall to medium
center; aperture wide, straight; dentition coarse to
medium; columellar lip with 25 teeth, labral lip with 24 size; spire involute; narrow elongate aperture; fossula
broad, smooth, concave.
teeth; prominent anterior terminal ridges that form a
slight marginal callus.
Remarks: Schilder and Schilder (1971) recognized 23
Comparison: The new species is unlike any other species and nine subspecies ofEocypraea sensu stricto.
Bemaya from western North America. Flowever, it does Six of their subspecies have been elevated to specific
resemble Bemaya obesa (Deshayes, 1865) from the status, five species have been described subsequent
upper middle Eocene (Bartonian) of the Paris Basin, to 1971, and lour new species are described here which
France, particularly the specimen figured by Cossmann brings the total to 38 species and three subspecies.
and Pissarro (1911: pi. 32, fig. 162-1). The new species Nine of these species are from western North America
is less inflated, more constricted anteriorly, has a (Table 1). Schilder (1924) established the subfamily
widerless sinuous aperture, and has finer dentition than Eocypraeinae for various genera of extinct and living
B. obesa. Because the posterior terminal ridges are eypraeids and ovulids including Eocypraea ofCossmann
missing in B. squiresi they cannot be compared; how- (1903). Schilder (1932) includedEocypraea in the family
,
ever, the anterior terminal ridges of B. obesa are more Amphiperatidae (= Ovulidae), subfamily Cypraediinae,
prominent than those ofB. squiresi. A specimen identi- but included fossil and living genera of ovulids
Page 48 THE NAUTILUS, Vol. 125, No. 2
lFl79Eoe0io—cn4gc.1g5uyt0,2rph3.re13Eas847oe.1-c8a1,8y-.mp(15Er2Em2oa..oce7clyayepmnpP(rgraEmataloeheec.aloaye)5gpn(-esrgE6pnato.„eehc.aEcyc)fopy3.cipr-Eiry4a.map.egera(ooEaEe)i.oedcd)caereyriean(ptsnEficrsloea.ancntetyee1aepw-n(r2(sLsaE.iapeosemBaccaneyi)reepmbcsrwaak,at,yseehapqoa1eul8)c(eo0iBtn2teaes)ysk,i,pmeseahohsynoyLulpeaAsoowu)tCtkysysMiqpppiuIeeeicPnrLiUeeeAs1sCwi,3CM6nMsh4pPoeIel1wcP,oi1ftse5yr1ps8pe3o,1ce6m5ih4eIo6Lfsl,GA,roMofCthmryoM5oplUIe1mo7tPCLLy4MplAAfoePcCCr.LoMMl4mAIoP1cC5LP.7MlAAo311c-C,.31PM621142I861.52P6/3,56f5m4l2r,o47cmo,..1m10fl1.rLme69no9Amgm5mtC1hmL,lM.eAI9n1l.gCPe41tnM—hgm.l1Ito2hmcP...
only. In 1971 Schilder and Schilder included the sub- Eocypraea (Eocypraea) takeosusukii new species
family Eocypraeinae in the family Ovulidae and again (Figures 3-4)
included living and fossil genera. Fehse (2001) elevated
E. (E.) n.sp. Groves, 1997: 7
Eocypraeinae to lull family status (Eocypraeidae) but
included onlyextinct genera. Diagnosis: An Eocypraea with inflated shell and
Subgenus Eocypraea Cossmann, 1903 slightlys-shaped aperture.
P )
L.T. Groves, 2011 Page 49
Description: Shell moderately inflated, of medium fossula smooth, wide; all surfaces smooth; anterior canal
size, constricted anteriorly; spire completely covered; shallow, posterior canal deep; anterior and posterior
dorsum highly arched; maximum height nearly cen- terminal ridges short; slight basal marginal callus on
ter; maximum width nearly center; aperture slightly outer lip; base rounded.
s-shaped and widens anteriorly, curves sharply toward
columella posteriorly; denticulation semi-coarse with Comparison: The new species is unlike any known
smooth interstices; outer lip with 16 teeth, inner lip species in the eastern Pacific. However it somewhat
with 11 teeth; fossula smooth, wide; all surfaces smooth; resembles Eocypraea inflate (Lamarck, 1802) from the
posterior columella highly inflated; posterior canal middle Eocene (Lutetian Stage) ofParnes, Oise Depart-
deep, anterior canal missing; anterior and posterior ment, Paris Basin, France as illustrated by Cossmann
basal ridges slight and do not form basal calluses; base and Pissaro (1911: pi. 32, fig. 162-7) and E. cotteri Cox
rounded. (1930: pi. 19, figs. 8a-8c) from the late Paleoeene
(Thanetian Stage) of the Samana Range of northwest
Comparison: Eocypraea (E.) takeosusukii is unlike India. Eocypraea (E.) batequensis differs from both
other eocypraeid in the eastern Pacific and is most simi- mainly by its smaller size but also by it's narrower aper-
lar to Eocypraea bartlettiana (Maury, 1912: 86-87, ture, finerdentition, and less calloused outer lip.
pi. 1 1, figs. 11-13) from the Paleoeene (Thanetian Stage)
ol Soldado Rock, Trinidad, Trinidad and Tobago. The Discussion: Excellent preservation ol the holotype
new species differs from E. bartlettiana by the lack ofa permits unequivocal generic assignment. Eocypraea (E.)
prominent basal callus, coarser dentition, and more sin- batequensis differs from all other eoeypraeids in the
nous aperture. Western Hemisphere andis the firstdescribedfrom Baja
California Sur, Mexico.
Discussion: Good preservation of the holotype per-
mits unequivocal generic assignment. Eocypraea (£. Material: Represented by a single well preserved
takeosusuki differs from all other eoeypraeids in the internal mold.
Western Hemisphere and is the first eocypraeid de-
scribed from the “Martinez” Formation. Type Material: Holotype IGM 5174, measures 9.4 nun
mm mm
in length, 6.5 inwidth, and5.1 in height.
Material: The new species is represented by a single,
moderately well preserved specimen exhibiting original Type Locality: LACMIP loc. 16951 (= CSUN loc.
shell material on its base and dorsum but with minor 1220b), Mesa La Salma, Baja California Sur, Mexico,
amounts ofshell missing from the dorsum. The anterior Bateque Formation.
end ofthe shell is missing.
Type Material: Holotype LACMIP 13645, measures Etymology: This species is named for the Bateque
mm mm Formation.
18.8 inlength, 14.4 in width, 11.2 nun in height.
Eocypraea Eocypraea crescentensis newspecies
Type Locality: LACMIP loe. 7045 {ex CIT loc. 1589), (Figures 7-8() )
EastofLowerLake, Lake County, California, “Martinez”
Formation. E. (E.) n. sp. 2. Groves, 1997: 7 (in part).
Etymology: This species is named in honorofthe late Diagnosis: An eocypraeid with inflated shell and
Takeo Susuki {ex UCLA) for his numerous important slightly s-shaped aperture.
contributions to the study of invertebrate paleontology
ofsouthern California. Description: Shell moderately inflated, of small size;
constricted anteriorly and slightly produced; spire cov-
Eocypraea (Eocypraea) batequensis new species ered; maximum height slightly posterior ofcenter; max-
(Figures 5-6) imum width nearly center; aperture wide and veiy
Eocypraea sp. Squires and Demetrion, 1992: 31, figs. slightly s-shaped; denticulation coarse with smooth
77-79. interstices, outerlipwith 14 teeth, inner lip with 6 teeth;
fossula smooth, wide; dorsal surface exhibits linear pat-
E. (E.) n. sp.l. Groves, 1997: 8.
tern which could represent growth lines; anterior and
Diagnosis: An Eocypraea withinflatedshell and slightly posterior canals shallow; anterior and posterior basal
s-shaped aperture. ridges reduced forming a slight collumellar basal callus;
slight posterior spiral sulcus present; base rounded.
Description: Shell moderately inflated, of small size,
constricted anteriorly and posteriorly; spire covered; Comparison: Eocypraea (E.) crescentensis n. sp. most
maximum height of dorsum slightly posterior of cen- closely resembles E. (E jimgoeclerti n. sp. (this paper)
ter; maximum width nearly center; aperture slightly but has a straighter ap.e)rture, a less prominent basal
s-shaped; denticulation semi-coarse with smooth inter- collumellar callus, more produced extremities, and a
stices; outer lip with 24 teeth, inner lip with 18 teeth; posteriorspiral sulcus.
)
Page 50 THE NAUTILUS, Vol. 125, No. 2
Discussion: Excellent preservation of the holotype Western Hemisphere and is the second eoeypraeid
permits positive generic assignment. Eocypraea (E.) described from the Crescent Formation.
crescentensis differs from all other eocypraeids in the
Western Hemisphere and is the first eoeypraeid de- Material: The new species is represented by a single
scribed from the Crescent Formation. The new species fairlywell preserved specimen thatexhibits original shell
represents the northernmostrecordofthegenus inwest- material.
ern North America.
Type Material: Holotype LACMIP 13647, measures
Material: The new species is represented by the 12.7 mm in length, 8.2 mm in width, and 6.9 mm in
fairly well preserved holotype specimen that exhibits
height.
original shell material and a poorly preserved topo-
mm
typie internal mold that measures 7.8 in length,
5.6 mm in width, and 4.3 mm in height, and three Type Locality: LACMIP loc. 41573 (ex CSUN loc.
small fragments 1573), Doty Hills, Lewis County, Washington, Crescent
Formation.
l\|je Material: Holotype LACMIP 13646, measures
10.9 mm in length, 7.6 mm in width, and 6.1 mm in Etymology: This species is named for colleague }im
heigOht. Goedert, Gig Harbor, Washington, who collected the
holotype and donated it to LACMIP and for his numer-
Type Locality: LACMIP loc. 16655 (ex CSUN loc. ous important contributions to Tertiary molluscan pale-
1563), Larch Mountain, Thurston County, Washington, ontologyofWashington.
Crescent Formation.
Eocypraea (Eocypraea) sp., cf. £. (E.) inflata (Lamarck,
Etymology: This species is named for the Crescent 1802)
Formation, Washington. (Figures 1 1-12)
Eocypraea (Eocypraea)jimgoederti new species Eocypraea castacensis (Stewart, 1926 [1927]): Vokes,
(Figures 9-10) 1939: 26, 154, pi. 20, fig. 14.
Cypraea castacensis Stewart, 1926 [1927]: Ingram,
E. (£.) u.sp. 2. Groves, 1997: 7 (in part). 1942: 103, pi. 8, fig. 6.
Eocypraea (Eocypraea) moumieti Dolin and Dolin,
Diagnosis: Aneoeypraeidwithinflatedshellandslightly
1983: 36; Groves', 1997: 8.
s-shapedaperture.
Description: Shell slightly inflated, of small size; Remarks: The hypotype of Vokes (1939, pi. 20,
constricted anteriorly and posteriorly and slightly pro- fig. 14) [UCMP 15815] most closely resembles £. (£.)
duced; spire covered; dorsum highly arched; maximum inflata (Lamarck, 1802) from middle Eocene strata
height slightly posterior of center; maximum width (Lutetian/Bartonian stages) of France, Belgium, and
nearlycentered; aperture wide for size, slightlys-shaped; England. This is particularly evident from the illustra-
dentieulation coarse with smooth interstices; outer lip tions of Cossmann, 1903 (pi. 9, figs. 18-19) and
with 14 teeth, inner lip with 12 teeth; fossula smooth Cossmann and Pissarro, 1911 (pi. 32, fig. 162-7). It also
and wide; all surfaces roughened by preservation; poste- superficially resembles E. (£.) maniobraensis Squires
and Advocate, 1986 from the lower Eocene (“Capay”
rior columella moderately inflated; anterior canal shal-
[CPMS]) Maniobra Formation of Riverside County,
low, posterior canal deep; anterior terminal ridges
prominent, posterior terminal ridges slightly reduced; California. Eocypraea (£.) maniobraensis is more elon-
basal marginal callus moderate on outer lip; base gate, has coarser dentition, has a prominent basal callus,
rounded. and is significantly larger than £. (£.) sp. cf. £. (E.)
inflata.
Comparison: Eocypraea (E.) jimgoederti new spe- Vokes (1939) and Ingram (1942) both misidentified
cies is different from all other eocypraeids from the this poorly preserved specimen as £. (£.) castacensis
eastern Pacific but is somewhat similar to a specimen (Stewart, 1926 [1927]). Most specimens of £. (£.)
oEfocEe.ne(£.()Ludtoeltlifuasni S(tLaagueb)rioefreP,ar1n8es8,1)OifsreomDetphaertmmiedndtl,e claessstagcleonsbiossehatvheana£f.air(l£y.)prsop.m,incfe.nt£b.as(a£.l)cainlflluastaa.ndDoiltiins
Paris Basin, France as figured by Cossmann and Pissaro and Dolin (1983) described £. (£.) moumieti from the
(1911: pi. 33, fig. 162-8). The new species has coarser Gan Basin, Pyrenees Atlantique Deptartment, France.
dentition, straighter aperture, and has a less inflated They included the hypotype of Vokes (1939) as this
columellar region than E. (E. dollfusi. new species but did not refigure it. Dolin and Ledon
) (2002) tentatively reassigned £. (£.) moumeti to the
Discussion: Good preservation of the holotype per- eoeypraeid genus Sulcocypraea. Eocypraea (£.) sp. cf.
mits positive generic assignment. Eocypraea (£. £. (£.) inflata differs from E. (£.) moumeti by its finer
jimgoederti differs from all other eocypraeids in the dentition and a slightly more inflated columellar region.
L.T. Groves, 2011 Page 51
ACKNOWLEDGMENTS
Deshayes, G.-P. 1835. Description des eoquilles fossils des
environs de Paris. 2: 1-783 + Atlas, pis. 1-105. F.-G.
Manythanks to Richard L. Squires (CSUN) and LouElla Levrault, Paris.
R. Saul (LACMIP) who reviewed an early draft of the Deshayes, G.-P. 1865. Description des animauxsans vertebres
manuscript and offered valuable comments and sugges- deeouverts dans le basin de Paris. 3: 1-667 + Atlas, pis.
tions. Richard L. Squires (CSUN) collected the eoey- 1-107. J.-B. Bailliere et Fils, Paris.
praeid specimen from the Bateque Formation and Dolin, C. and L. Dolin. 1983. Revision des Triviaeea et
made it available for study. Maria del Carmen Perrilliat Cypraeacea (Mollusea, Prosobranchiata) Eocenes recoltes
(IGM) kindly loaned the Bateque Formation specimen. dans les localities de Gan (Tuilerie et Acot) et Bosdarros
(Pyrenees Atlantiques, France). Mededelingen van de
James L. Goedert (Gig Harbor, Washington and
LACMVP Research Associate) collected the eocypraeid Werkgroep voor Tertiaire en Kwartaire Geologie 20(1):
5-48, figs. 1-31.
specimens from the Crescent Formation, made them Dolin, L. and D. Ledon. 2002. Nouveaux taxons et discussion
available for study, and provided pertinent information de la systematique des genres correspondants d’Ovulidae
regarding their stratigraphic occurence. Special thanks (Mollusea, Caenogastropoda) de l’Eocene inferieur de
to David Lindberg (UCMP) who loaned compara- Gan (France). Geodiversitas 24: 329-347, figs. 1-5.
tive material. The late Don McNamee (LACM Research Durham, J.W. 1944. Megafaunal zones of the Oligocene of
Library) processed numerous interlibrary loans and northwestern Washington. University ol California Publi-
assisted in the acquisition ofseveral obscure references. cations, Bulletin of the Department of Geological Sci-
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