New species of Anthonomus germar in the A. dogma and A. furcatus species groups (Coleoptera: Curculionidae)

9 Pages·1994·2.8 MB·

by  W E Clark

PROC. ENTOMOL. SOC. WASH.
96(2). 1994. pp. 199-207
NEW SPECIES OF ANTHONOMUS GERMAR IN THE A. DOGMA AND
A. FURCATUS SPECIES GROUPS (COLEOPTERA: CURCULIONIDAE)
Wayne E. Clark
Department of Entomology and Alabama Agricultural Experiment Station, Auburn
University, Alabama 36849-5413.
Abstract.—Four previously undescribed species of Neotropical Anthonomini are as-
signed to the Antlwnomus dogma group. These are A. dogma (Colombia, Panama), A.
idea (Brazil), A. opinionis (Panama, Venezuela) and A. ratio (Brazil). Characters ofthe A.
dogma group and ofeach ofthe species in the groupare described and a key to the species
is presented. The habitus of each species, the pygidium, abdomen, femur and tibia of
some of the species, and the male genitalia of one of the species are illustrated with
photographs. A new species oftheAnthonomusfurcalusgroup, A. blik(Venezuela), is also
described and illustrated.
Key Words: Coleoptera, Curculionoidea, Curculionidae, Anthonomus dogma. Anthon-
omus idea, Anthonomus opinionis. Anthonomus ratio. Anthonomus hlik
A recently recognized new species ofAn- (CWOB); Collection of H. P. Stockwell,
thonomus in the A. furcatus group differs Smithsonian Tropical Research Institute,
somewhat in external appearance from the Panama (HPSC); Universidad Central de
eleven species previously assigned to that Venezuela, Maracay, Venezuela (IZAV);
group(Clark 1988a).Thetypeofthisspecies Museu Paraense Emilio Goeldi, Belem,
bears impressive resemblance to specimens Brazil (MPEG): Museu de Zoologia, Uni-
representing four previously undescribed versidade de Sao Paulo, Sao Paulo, Brazil
A&M
species that belongto a group related to the (MZSP): Texas University, College
.4. furcatus group. This group is described Station, Texas, USA (TAMU); National
herein as the A. dogma species group. This Museum of Natural History, Washington,
paper includes descriptions and illustra- D.C., USA (USNM). Measurements were
tions and a key to the species in the A. dog- made according to criteria described by
ma group, as well as a description and il- Clark (1994). Exact label data are cited for
lustrations of the new species in the A. types. Separate labels are indicated by
furcatus group. brackets ([]), each separate line by a slash
Materials and Methods
The Anthonomus dogma Group
Specimens of 1 1 adult weevils were ex-
amined. These are from the collections of Recognition characters.—The members
the following individuals and institutions oftheA. dogmagroupare moderately large,
(letter codens identify the collections in the shining. hXackAnthonomus {\QX\g\h = 2.88-
text): Collection of C. Bordon, Maracay, 3.60 mm; width = 1.32-1.76 mm) with
Venezuela (CCBM); Collection of C. W. sparse lacteous scales above, and broader,
USA
O'Brien, Tallahassee, Florida, dense lacteous scales beneath (Figs. 1-8).
200 PROCEEDINGS OFTHE ENTOMOLOGICALSOCIETY OF WASHINGTON
^
*^9fv*"y8|5^: . St-
^jk-
Jg^^— 'Ss
Figs. 1-8. Antlioiiomus dogma group members, habitus, lateral and dorsal views. 1, 2) .-1. dogma, female.
Fort Clayton, Panama, Panama; 3, 4) .1. idea, female, hololype; 5, 6) .A. opinwius. female, holotype; 7. 8) .4.
ratio, female, holotype.
.
VOLUME NUMBER
96, 2 201
The pronotum and elytra are mostly gla- groups. Therostrum of.4. subulatus(Cham-
brous, with lacteous scales in a narrow dor- pion) is "abruptly and strongly narrowed at
sal midline on the pronotum, alongthe pos- the antennal insertions, the distal por-
. . .
terior margin of the pronotum (Figs. 2, 4, tion . . . extraordinarily narrow, smooth,
6, 8), in a conspicuous postocular vitta on glabrous, and subparallel sided in dorsal
the pleuron, and in an apicolateral elytral view" (Clark 1993). The rostrum is similar
vitta (Figs. 1, 3, 5, 7) (the latter feature is in two ofthe species in theA. dogma group,
inconspicuous in A. dogma itself (Fig. 1)). A. dogma and A. idea (Figs. 1, 3). The en-
The thoracic and abdominal sterna have dophallus ofsome of the species in the A.
dense lacteous scales. Similar scales are es- furcatus group is like that in A. subulatus
pecially dense on the metepimeron and which has"twoproximal spinesandone
. . .
metepisternum and form aconspicuousline more distal sclerite" (Clark 1993). The
. . .
that connects the pleural and apicolateral cndophallus ofA. dogma (Fig. 18) (males of
elytral vittae (Figs. 1, 3, 5, 7). Each ofthe the other species are unknown) is similar,
small to moderately large punctures on the but more like that ofother members ofthe
remainder ofthe pronotum and on the up- A.furcatusgroup, including the new species
per portion of the pleuron has a minute, described herein (Fig. 19), in having an ex-
inconspicuous, aeneous seta. The profem- tra pair of spines between the proximal
ora are moderately to strongly inflated, 1.4- spines and the distal sclerite.
1.9X as wide as the metafemur. Plant associations.—Unknown.
Relationships.—The most striking attri- Distribution.—The species ofthe A. dog-
butes ofthe species in the A. dogma group ma group are known from Brazil, Colom-
are the smooth, shining, black integument bia, Panama and Venezuela.
and the contrasting sparse lacteous scales of
the convex pronotum and elytra (Figs. 1- Key to Species in the
8). Some unrelated species ofAnthonoinus Anthonomus dogma Group
with similarcharacteristicswereassigned to 1 Rostrum stout basally, strongly, evenly ta-
the A. alholineatus group by Clark (1987). pered to narrow apex in male, abruptly nar-
These had been described by Champion rowed slightly beyond antennal msertions \n
(o1f9O0t3i)doacsehpahvailunsg'"'.a.n.dgebnyerSallefeapceires(1li9k5e8t)haats offemdaelnese(Fligasc.teIo.u3s);scelayllersa(wFiigtsh.o2u,t4)h;uemyeersalstpraotncgh-
ly, evenly convex, not prominent (Figs. 1^);
"superficially" resembling otidocephaline
abdominal sternum 5 of female with poste-
Curculionidae. They were inelegantly des- romedianclaspthatapparentlyreceivesapical
ignated "otidocephaline-like" by Clark portion ofpygidium (Figs. 13. 14); pygidium
(1987), who asserted that the peculiar con- offemale with slight apicomedian projection
dition ". . . is the result ofconvergent evo- (tFoiogtsh. d9i,st1a0l);tomleatragefevmeuntrrawlitthooatnh o(Fbisgo.le1s5c)ent 2
lution in Otidocephalinae and Anthonom- r. Rostrum slender from base to apex (Figs. .5,. .
inae [and] ... in different groups of 7);elytrawithhumeralpatchofdenselacteous
Anthonominae." Convergence was also in- scales (Figs. 6, 8); eyes most strongly convex
vokedwhenanthonomineswithsimilarfea- posteriorly,slightlyprominent(Figs. 5-8);ab-
tures turned up in the genus Loncophorus dominal sternum 5 offemale without poste-
romedian clasp; pygidium offemale without
Germar (Clark 1988b). apicomedian projection (Figs. II, 12); meta-
The somewhat "otidocephaline-like" femurwithoutobsolescenttoothdistaltolarge
species in the A. dogma group appear not ventral tooth (Figs. 16, 17) 3
to be closely related to the anthonomines 2. Elytrawithdenselacteousscalesonbasalpor-
in these othergroups. They seem instead to tionsofsuturalinterstriae(Fig.4);apicolateral
elytralscalesdistinct, lacteous(Fig. 3);protib-
be allied to "non-otidocephaline-like" spe- iaoffemalewithoutpreapicaltooth;metafem-
cies in the A. subulatus and A. furcatus oral tooth conical A. idea
202 PROCEEDINGS OFTHE ENTOMOLOGICAL SOCIETY OF WASHINGTON
2'. Elytra without lacteous scales on basal por- 9); eyes strongly, evenly convex, not prom-
tionsofsuturalinterstriae(Fig. 2);apicolateral inent (Figs. 1, 2); protibia offemale with a
elytral scales indistinct, aeneous and lacteous small, acute, preapical ventral tooth; meta-
(Fig. 1);protibiaoffemalewithpreapicaltooth; femur with a slightly uncinate, acute, ven-
metafemoral tooth slightly uncinate (Fig. 15)
4. dogma tral tooth and an obsolescent more distal
3. Rostrum tapered distally (Fig. 7); metafemur tooth (Fig. 15); profemur ca. 1.4x as wide
with a short, conical ventral tooth (Fig. 17) . as metafemur.
1. ratio Male.-Length: 3.28-3.40 mm (n = 2).
y. Rfeomsutrruwmitnhoatltoanpge,resdlendidsetralvleynt(rFailg.to5o);thme(Ftiag-. Width: 1.72-1.76 mm (n = 2). Head: eyes
16) I. opmionis separatedbydistanceca. 0.8x width ofros-
trum at base. Rostruni: length 1.26-1.59 x
Anthonomus dogma. New Species (n = 2) pronotal length; proximal portion
Figs. 1, 2, 9, 13, 15, 18 carinate, sparsely setose, with narrow lac-
Holotype.—PANAMA. Panama: $ [Pan- teous scales at extreme base and on frons;
ama: CanalZone/Albrook ForestSite/ Fort distal portion glabrous, sparsely punctate,
Clayton] [Lot No. 174/ Feb. 23/24, 1968/ length 46-48%(n = 2)oftotalrostrallength.
R. Hutton/ Black light trap] [University/ Prothorax: pronotum punctate, interspaces
Arkansas/ Collection] (CWOB). broad, smooth, shining; pleuron more
Paratypes (4).-COLOMBIA. Norte de coarsely punctate, interspaces narrower, ru-
Santander: 1KfmemNalWe [Colombia/ Norte de gulose; pronotum with narrow midline of
Santander/4 Malpaso/8July 1982/ small, dense, lacteous scales, and with lon-
Clark & Cave] (TAMU). PANAMA. Pan- ger, sparser, lacteous scales along posterior
ania: female [Panama: Canal Zone/ Al- margin; each punctureon remainderofdor-
1
brook Forest Site/ Fort Clayton] [Lot No. sum and upper portion of pleuron with a
174/ Feb. 23/24, 1968/ R. Hutton/ Black minute, inconspicuous seta; pleuron with
light trap] [University/ Arkansas/ Collec- conspicuous postocular vitta ofbroad lac-
tion](CWOB); 1 male[Panama:CanalZone/ teous scales. Elytra: striae not impressed,
Albrook ForestSite/FortClayton] [LotNo. with elongate punctures; interstriae slightly
176/ Feb. 29/Mar. 12, 1968/ R. Hutton/ convex, smooth, shining, glabrous, except
Black light trap] [University/ Arkansas/ forapicolateral scales. Abdomen:sterna with
Collection] (CWOB); 1 female [Panama: lacteous scales that are broader laterally.
CanalZone/AlbrookForestSite/FortClay- Legs: coxae with dense, broad, lacteous
ton] [Lot No. 130/ Sept. 7/8, 1967/ Hutton scales; protibia slightly curved in basal '/:,
& Llaurado/ Black light trap] [University/ with slight inner marginal prominence in
Arkansas/ Collection] (CWOB). distal Vv. protibial uncus short, slender,
Recognitioncharacters(Figs. 1, 2).—Ros- curved, acute; metatibial mucro stout, par-
trum stout basally, strongly, evenly tapered allel tolongaxisoftibia. Genitalia(Fig. 18):
to narrow apex in male, abruptly narrowed aedeagus narrow, strongly sclerotized be-
slightly beyond antennal insertions in fe- low, subtruncate apically; endophallus with
male (Fig. 1); elytra without lacteous scales two small basal spines, twosimilar, median
on sutural interstriae (Fig. 2); apicolateral spines, and a small distal spine.
elytral scales minute, inconspicuous, aene- YemaXe.—Length: 2.88-3.28 mm (mean
ous and lacteous (Fig. 1); abdominal ster- = 3.09, n = 3). Width: 1.42-1.72 mm (mean
num 5 offemale with posteromedian clasp = 1.58, n = 3). Rostrum:length 1.49-1.60x
that apparently receives apical portion of (mean = 1.54, n = 3)pronotal length; length
pygidium (Fig. 13); median emargination of ofdistal portion 46-49% (mean = 47, n =
clasp broad (Fig. 13); pygidium of female 3) oftotal rostral length.
with rounded apicomedian projection (Fig. Distribution.—,-t/;//;6»/;o/H»5 dogma is
VOLUME 96. NUMBER 2 203
ia
ia
13 14
16
Figs. 9-17. Anlhononius dognui group members. 9-12. Pygidium. female, dorsal views: 9) A. dogma. Fort
Clayton, Panama, Panama; 10) ,-1. idea, holotype; 11).-1. opinionis, hololype; 12) .-1. ratio, holotype. 13, 14.
.Abdomen, female, ventral views: 13)A. dogma. Fort Clayton. Panama. Panama; 14)A. idea, holotype. 15-17.
Metathoracic legs: 15) 1. dogma. Fort Clayton, Panama, Panama; 16) .1. opinionis. holotype; 17) .-1. ratio.
holotype.
204 PROCEEDINGSOFTHE ENTOMOLOGICAL SOCIETY OF WASHINGTON
known only from the type series from Co- scaleson sutural interstriaeandapicolateral
lombia and Panama. scales.Abdomen:sterna with lacteousscales
that are broader laterally. Legs: coxae with
Anthonomus idea, NF^^ Species dense, broad, lacteous scales; protibia
Figs. 3, 4, 10, 14 slightly curved in basal '/:, with slight inner
Holotype.-BRAZIL. Para: 9 [Sta. Isabel marginal prominence in distal '/r, protibial
do Para/ PA, Brasil/ 30.III.1962/ J. Be- uncus short, slender, curved, acute; meta-
chyne col.] (MZSP). tibial mucro stout, parallel to long axis of
Paratype (1).-BRAZIL. Para: 1 female tibia.
[BRAZIL Para/ Belem/ 1-26-1969 L &/ C. 'Dx^XrihuXxon.—Anthonomus idea is
W. O'Brien (CWOB). known only from the type series from Bra-
Recognition characters (Figs. 3, 4).—.4/?- zil.
thonomus idea is similar to A. dogma from
which it differs in the following characters: Anthonomus opinionis. New Species
Elytra with dense lacteous scales on basal Figs. 5, 6, 11, 16
portions ofsutural interstriae (Fig. 4); api- Holotype.—PANAMA. Panama: 9
NW
colateral elytral scales distinct, lacteous (Fig. [PANAMA: C.Z./ 5 mi. Gamboa/ 12
3); prolibia of female without preapical July '76 Lubin] [Malathion/ canopy fog]
tooth: metafemoral tooth conical, acute: [Antho./ #17] (from HPSC, deposited in
median emargination of posteromedian USNM).
clasp of abdominal sternum 5 of female Paratype (1).-VENEZUELA. Tdchira: I
deep, narrow (Fig. 14); pygidium offemale female [Sn. Joaquin/ de Navay/ Venezue-
with smaller apicomedian projection (Fig. la - Tachi-/ ra. 400 m./ 21-22-1979] [col./
10). A. Chacon] (IZAV).
Male.—Unknown. Recognition characters (Figs. 5, 6).—An-
¥Qma\e.—Length: 3.12-3.24 mm (n = 2). thonomus opinionis is distinguished by the
Width: 1.50-1.58 mm (n = 2). Head: eyes following combination of characters: ros-
round, strongly, evenly convex, not prom- trum slender, evenly, broadly curved, not
inent, separated by distance ca. 0.8 x width tapereddistally(Fig. 5); elytra with humeral
of rostrum at base. Rostrum: length 1.43- patch ofdense lacteous scales (Fig. 6); api-
1.70x (n = 2) pronotal length; proximal colateral elytral scalesdistinct, lacteous (Fig.
portion carinate, sparsely setose, with nar- 5); eyes most strongly convex posteriorly,
row lacteous scales at extreme base and on slightly prominent (Figs. 5, 6); metafemur
frons;distalportionglabrous, length 56-58% with a long, slender ventral tooth, without
(n = 2) of total rostral length. Prothorax: more distal tooth (Fig. 16); abdominal ster-
pronotum punctate, interspaces broad, num 5 of female with posterior margin
smooth, shining; pleuron more coarsely slightlyproduced; profemurca. 1.9x aswide
punctate, interspaces narrower, rugulose; as metafemur.
pronotum with narrow midline of small, Male.—Unknown.
dense, lacteous scales, and with longer, Female.—Length: 3.44-3.60 mm (n ^ 2).
sparser, lacteous scalesalongposteriormar- IVidth: 1.52-1.58 mm (n = 2). Head: eyes
gin; each puncture on remainder ofdorsum separated by distanceca. 0.4x width ofros-
andupperportion ofpleuron witha minute, trum at base. Rostrum: length 1.70-1.93 x
inconspicuous seta; pleuron with conspic- (n = 2) pronotal length; proximal portion
uous postocular vitta of broad lacteous feebly carinate, apparently glabrous; distal
scales. Elytra: striae not impressed, with portion smooth,glabrous, length 39^6%(n
elongate punctures; interstriae slightly con- = 2) oftotal rostral length. Prothorax: pro-
vex, smooth, shining, glabrous, except for notum sparsely, minutely punctate, inter-
VOLUME 96. NITMBER 2 205
spacesbroad, smooth, shining; pleuron more coarselypunctate, interspacesnarrower, ru-
coarsely punctate, interspaces narrower, ru- gulose; pronotum with narrow midline of
gulose; pronotum with narrow midhne of small, dense, lacteous scales, and with lon-
small, dense, lacteous scales, and with lon- ger, sparser, lacteous scales along posterior
ger, sparser, lacteous scales along posterior margin; each puncture on remainderofdor-
margin; each punctureon remainderofdor- sum and upper portion of pleuron with a
sum and upper portion of pleuron with a minute, inconspicuous seta; pleuron with
minute, inconspicuous seta; pleuron with conspicuous postocular vitta of broad lac-
conspicuous postocular vitta of broad lac- teous scales. Elytra: striae not impressed,
teous scales. Elytra: striae not impressed, with minute punctures; interstriae slightly
with minute punctures; interstriae slightly convex, smooth, shining, glabrous, except
convex, smooth, shining, glabrous, except for scales on humeri and in apicolateral vit-
forscaleson humeri andapicolateral scales. ta. Pygidium: broad, without apicomedian
Pygidium: broad, without apicomcdian projection (Fig. 12). Abdomen: sterna with
projection (Fig. 1 1). Abdomen: sterna with lacteous scales that are broader laterally.
lacteous scales that are broader laterally. Legs: coxae with dense, broad, lacteous
Legs: coxae with dense, broad, lacteous scales; protibiacurved inbasal '/:, with inner
scales;protibiacurved inbasal '/:. with inner marginal prominence indistal 'A, with small,
marginal prominence in distal '/3, without acute, preapical tooth; protibial uncus slen-
preapical tooth; protibial uncus slender, der, slightlycurved,acute; metatibialmucro
slightly curved, acute; metatibial mucroob- obsolete.
solete. T>\sXribuUor\.—Anthononius ratio is
XyhXnhuUon.—Anthonomus opinionis is known only from the type series from Bra-
known only from the type series from Pan- zil.
ama and Venezuela.
The Anthonomus furcatus Group
Anthonomus ratio. New Species Recognition characters.—The species of
Figs. 7, 8, 12, 17 Anthonomus in the A. furcatus group were
Holotype.-BRAZIL. Para: 2 [Marituba/ characterized by Clark (1988a) as "relative-
23.6.1961] [Brasil, PA/ J. & B. Bechyne] ly large anthonomincs . . . with the rostrum
(MPEG). relatively slender and of moderate length,
Recognition characters (Figs. 7, ^).—An- the antennal funiculus 7-articulate and rel-
thononius ratio is similar to A. opinionis atively large, and the eyes strongly and
from which it differs in the following char- evenly convex." Cited as unique characters
acters; Rostrum tapered distally (Fig. 7); of the group were the "aedeagus with dis-
metafemur with a short, conical ventral tinct apicolateral prominences," and "en-
tooth (Fig. 17). dophallus with two proximal endophallic
Male.—Unknown. spines and one straight or uncinate, more
YemzXa.—Length: 2.96 mm (n = 1). distal sclerite," most species also with "two
Width: 1.32 mm (n = 1). Head: eyes sepa- spines between the proximal spines and the
rated by distance ca. 0.4x width ofrostrum distal endophallic sclerite." Additional
at base. Rostrum: length 1.65 x (n = 1) pro- characters cited, these shared with species
notal length; proximal portion feebly cari- in other groups, included "abdominal ster-
nate, apparently glabrous; distal portion num 5 ofmaleapicomediallydepressedand
smooth, glabrous, length 45% (n = 1) of with a posteromedian prominence which is
total rostral length. Prothorax: pronotum part of an isolated posteromedian sclerite
sparsely, minutely punctate, interspaces andprofemoraswollen,withalargeven-
. . .
broad, smooth, shining; pleuron more tral tooth and a smaller distal tooth."
206 PROCEEDINGS OFTHE ENTOMOLOGICAL SOCIETY OF WASHINGTON
1S 13
Figs. 18-19. Anihonomiisspecies. 18..-1. c/f\?/?;a. holotype. malegenitalia,dorsal view. 19..1. />///:. holotype,
malegenitalia, dorsal view.
The new species described below has all Ua\Q.-Length: 3.28 mm (n = 1). Width:
ofthese A. furcatus group characters. Nev- 1.48 mm (n = 1). Head:eyes round, strong-
ertheless, it is quite distinct in appearance ly, evenly convex, prominent, separated by
from the species previously assigned to the distance ca. 0.5x width ofrostrum at base.
groupand was overlooked until now. main- Rostrum: slender, length 1.52 x (n = 1)
ly because the vestiture is so different. In pronotal length; proximal portion carinate,
the previously described species, the scales sparsely setose, with narrow lacteous scales
on thepronotum andelytra are fairly dense, at extreme base and on frons; distal portion
relatively broad, those on the elytral inter- glabrous, sparsely punctate, length 33% (n
striae "short, narrow, the apices directed = 1) oftotal rostral length. Prothorax: pro-
toward the midline ofthe interstria" (Clark notum punctate, interspaces smooth, shin-
1988a:360). By contrast, the scales on the mg; pleuron more coarsely punctate, inter-
pronotum and elytra ofthe new species are spaces narrower, rugulose; pronotum with
small and sparse and the shining, black in- narrow midline of small, dense, lacteous
tegument is broadlyexposed between them. scales, and with intermixed sparse, long,
narrow, lacteous scales and narrower, in-
Anthonomus blik. New Species conspicuous aeneous scales: pleuron with
Fig. 19 conspicuous postocular vitta of broad lac-
Holotype.-VENEZUELA. Lara: 6 teousscales. Elytra:striaenottoonly slight-
[VENEZ. Bordon/ leg. 13 XII 1978] [aired, ly impressed, with elongate punctures: in-
de Bobare./ m. 900. Edo. Lara] (CCBM). terstriae nearly flat, smooth, shining, with
Recognition chaTaciers.—Anthonomus sparse, narrow, lacteous scales; lacteous
blik is distinguished from the other mem- scalesbroaderanddenseinlongapicolateral
bers of the A. furcatus group by the black, \itta; interstriae 2-5 slightly prominent near
smooth, shining integument and the small, base. Abdomen: sternum 5 with isolated
sparse, lacteous scales on the prothorax and posteromedian sclerite. Legs: procoxa with
elytra. The scalesaredenseonly in a narrow large, blunt, anterior tooth: profemur 1.8x
medianvittaonthepronotumandinabroad as wide as metafemur, with a small slightly
vitta that extendsacross the pleuron, across curved tooth anterobasal to large, broad-
the metepimeron and metepistemum, and based, ventral tooth; protibia curved in bas-
around the apicolateral portions of the el- al -A, with slight inner marginal prominence
ytra. This pattern gives the species the gen- in distal '6; protibial uncus slender, curved,
eral "otidocephaline-like" appearance not- acute; metafemur with one stout, conical
ed in thespeciesin the.-I. dogmagroup. The tooth; metatibial mucro stout, parallel to
large procoxal tooth is also unique to the long axis of tibia. Genitalia (Fig. 19): ae-
species. deagus with acute, apicolateral promi-
VOLUME 96. NUMBER 207
nences; endophalluswith twobasal sclerites culioninae (part), vol. 4. pt. 4. pp. 145-312. //;
amid a dense field ofdenticles, two median Godman, F. D.. and O. Salvm, eds.. 1879-1911.
spines anterior to a median row of small. BiologiaCentrali-Americana.Insecta.Coleoptera.
7 volumes in 17 parts. London. Dulau.
sparse denticles, and one stout distal spine. Clark.W. E. 1987. Thespeciesof.l/j/Zio/xi/f/i/.vmthe
Female.—Unknown. alholmcatus group (Coleoplera: Curculionidae).
DislnhuUon.—Amhononiusblikisknown Transactions ofthe American Entomological So-
only from the holotype from Venezuela. ciety 113(4): 309-359.
Etymology.—The name ofthis species is of.th1e98w8eae.vilRegveinsuisonAonflthhoen/oim»rutsinG«esrpemcaires(gCrooluep-
a term applied by Rolston (1987) for any optera:Curculionidae).TheColeopteristsBulletin
presupposition that might insulate a theory 42(4): 359-377.
from refutation. . 1988b. Revision of the weevil genus Lon-
cophonis Chevrolat (Coleoplera: Curculionidae,
Acknowledgments Anlhonominae).QuaeslionesEntomologicae24(3):
465-518.
Thanks are extended to the individuals . 1993. L«?/()rr/n7K-/;iHChampion,anewsyn-
and institutions listed in the Materials and onym oiAnlhonomus Germar (Coleoptera: Cur-
Methodssection forloansofspecimens used culionidae).TheColeopteristsBulletin47(2): 192-
in this study. H. R. Burke, John D. Free- 194.
man, and Michael Williams provided crit- . 1994. The Anlhonomus gibMcrus species
group(Coleoptera:Curculionidae). Proceedingsof
ical review ofthe manuscript published as the Entomological Society ofWashington 96(2):
Alabama Agricultural Experiment Station 19.V198.
Journal Series No. 17-933549. Rolston. H., III. 1987. Scienceand Religion: ACrit-
ical Survey. Temple University Press. Philadel-
Literature Cited phia. 357 pp.
Sleeper,E.L. 1958. NotesontheAnlhonominae(Co-
Champion, G. C. 1903. Insecla. Coleoplera. Rhyn- leoptera,Curculionidae).TheOhioJournalofSci-
chophora. Curculionidae. Curculionoidea. Cur- ence 58(6): 366-370.