New species and new records of Eastern Palaearctic scythridids from the Irano-turanian Region (Lepidoptera: Gelechioidea: Scythrididae)

2008·0.78 MB·

by  Pietro Passerin D’Entrèves

©EntomologicaFennica.11December2008
New species and new records of Eastern Palaearctic
scythridids from the Irano-turanian Region
(Lepidoptera: Gelechioidea: Scythrididae)
PietroPasserind’Entrèves&AngelaRoggero*
Passerind’Entrèves,P.&Roggero,A.2008:Newspeciesandnewrecordsof
Eastern Palaearctic scythridids from the Irano-turanian Region (Lepidoptera:
Gelechioidea:Scythrididae).—Entomol.Fennica19:159–167.
FournewScythrididaespeciesaredescribedfromtheEasternPalaearcticregion.
Scythris pamirica sp.n., S. balkhi sp.n., and S. brandti sp.n. were collected in
North-EasternAfghanistan,whileS.elburzisp.n.wascollectedinNorthernIran.
The lectotype of S. paelopyga (Staudinger, 1880) is established here, and the
malegenitaliaaredescribed.Morphologicalaffinitiesanddifferencesofthenew
speciesarereported, andrelatedtothescythrididtaxathatarecloser tothem.
NewrecordsoflittleknownspeciesfromtheIrano-turanianregionarelisted.
P.Passerind’Entrèves,UniversityofTorino,DepartmentofBiologiaAnimalee
dell’uomo,ViaAccademiaAlbertina13,I-10123Torino,Italy
A.Roggero,*correspondingauthor,Dept.BiologiaAnimaleedell’Uomo,Via
Accademia Albertina 13, I-10123 Torino, Italy; E-mail: angela.roggero@
unito.it
Received17May2007,accepted28September2007
1.Introduction Huemer 2003). Moreover, being very few new
speciesidentifiedandassignedtothisfamilytill
DuringtheXIXthandXXthcenturiesexpeditions now,thescythrididfaunaoffareasternareasre-
weresuccessivelyorganizedintheIrano-turanian mains poorly studied (Passerin d’Entrèves &
Region (Lederer 1869, Brandt 1938, Wiltshire Roggero2007).
1957).AslistedinAmsel(1949a),lepidopteran Twentynine species are recorded from vari-
material was collected by Bienert (1858–59), ouspartsofIranandAfghanistan(Table1),but
Haberhauer Jr. (1877–97), Wiltshire(1935–38), till now Scythris limbella alone was reported
Brandt(1936–37)andRichter(1954–56)among fromboththecountries(Landry1991).Basedon
others.Manynewlepidopteranspecieswerede- currentknowledgeofScythrididaeworlddistri-
scribed fromthe area, but the results were rela- bution,weexpectthefaunaofIranandAfghani-
tivelyscarcefortheScythrididae(i.e.,Staudinger stantobefarmorediversethanpresentlyknown,
1880, Christoph 1885, Meyrick 1918, 1924, andlikely mostofthespeciesarenotidentified
Amsel 1949b). More recently, further material yet. We also assume that there is more or less
wasobtainedaftertheHackerExpeditioninPaki- close relationships to the fauna of surrounding
stanin1988(Bengtsson1997a),andtheAustrian centralAsia(Turkmenistan,Tadjikistan,andPa-
ExpeditioninnorthernIranin2001(Bengtsson& kistan) and more remote areas (W Palaearctic,
160 Passerind'Entrèves&Roggero (cid:127) ENTOMOL.FENNICAVol.19
Table1.Listofthespeciesfromthestudyarea.
Afghanistan Iran
ApostibesafghanaPasserind’Entrèves&Roggero,2003 X –
Apostibesinota(Meyrick,1924) X –
CatascythriskebirellaAmsel1935 – X
Eretmoceramedinella(Staudinger1859) – X
Scythrisapicalis(Zeller1847) – X
ScythrisbiacutellaBengtsson2002 – X
ScythriscamelellaWalsingham1907 – X
ScythriscanceroidesBengtsson1997 – X
Scythriscanescens(Staudinger1880) X –
ScythriscurlettiiBengtsson1997 – X
ScythrisdecrepidellaBengtsson1997 – X
ScythrisethmiellaAmsel1974 – X
ScythrisfissurellaBengtsson1997 – X
Scythrisflabella(Mann1861) – X
ScythrisfriedeliBengtsson1997 – X
ScythrisghaemiiBengtsson&Huemer2003 – X
ScythrisheratiPasserind’Entrèves&Roggero2004 X –
Scythrislimbella(Fabricius1775) X X
Scythrismonochreella(Ragonot1896) – X
ScythrismusWalsingham1898 – X
ScythrisnielseniPasserind’Entrèves&Roggero2004 X –
ScythrispangalactisMeyrick1933 – X
Scythrissatyrella(Staudinger1880) – X
ScythrissenecaiBengtsson1997 – X
ScythrissinuosellaBengtsson2002 – X
Scythrissubclavella(Rebel1901) – X
ScythrissubparachalcaBengtsson2002 – X
ScythristessulatellaRebel1903 – X
ScythrisvalgellaBengtsson2002 – X
Arabian Peninsula, Afrotropical Region, India) usuallyproposedinliterature(Bengtsson1997b).
as is known for many other taxa (Larsen 1984, The preparation of genitalia followed the stan-
Akhanietal.1997,Maceyetal.1998,Ribera& dardprocedure(Landry1991,Bengtsson1997b),
Blasco-Zumeta1998,Sruoga&Puplesiené1998, butmountingfollowedthemethoddescribedby
(cid:1)iplak2003,Frisch2006,Mey2006,Nabozhen- Passerind’Entrèves(1976)forScythrididaemale
ko2006). genitaliainordertoavoidflatteninganddistort-
Theaimofthepresentworkistodescribefour ingofparts.
newspeciesofIrano-turanianScythrididaeandto FournewspeciesfromIrano-turanianRegion
updatetheknowledgeofthefamilybypresenting wereidentified(Fig.1),andaredescribedhere.
newdistributionaldataonanumberofotherspe- Thetypematerialsofthefournewspeciesarede-
cies. positedinStaatlicheMuseumfürNaturkundeof
Karlsruhe(SMNK,Germany),MuseumfürNa-
turkunde der Humboldt-Universität of Berlin
2.Materialandmethods (MNHB, Germany) and Naturhistoriska Riks-
museetofStockholm(NHRM,Sweden).
We examined typical and not-typical material Inthispaperwefollowedthetraditionalclas-
fromIranandAfghanistanpreservedinmanyin- sification,inwhichscythrididsarerankedatfam-
stitutional and private collections. To identify ilylevel(Nye&Fletcher1991,Kaila2004),re-
specimens, genitalia features were employed as jecting at present the taxonomy proposed by
ENTOMOL.FENNICAVol.19 (cid:127) Irano-turanianscythridids 161
Fig.1.Typelocalitiesof
thefournewscythridid
species:S.elburzisp.n.
(1),S.balkhisp.n.(2),
S.pamiricasp.n.(3)
andS.brandtisp.n.(4).
Hodges(1998)wholoweredthescythrididstatus third. Vinculum one third long as valvae, distal
tosubfamilylevel(inXylorictidaefamily). margin rounded. Aedeagus almost as long as
valvae,tubularanddown-turned,withdistalhalf
sigmoid and abruptly narrowing toward apex;
3.Newspecies juxta short and pyriform. Sternum VIII
subtriangular,aslongasvalvae,bifidatapex,api-
3.1.Scythrispamiricasp.n. ces diverging, carrying two small processes at
distal third, the proximal margin with deep,
Typematerial.Holotype:#(Fig.2a):NO[st]Af- arched notch and two thickened articular plicae
ghanistan, Khinch-e-Andarab, westl. Pamir, onsides.
3,550–4,000 m a.s.l., G. Ebert leg., 17.–27.VII.
1957.GenitalSlide288ZMB(MNHB).
Diagnosis.Thespeciesisexternallysimilarto
manyotherscythrididsandcanbeidentifiedonly
by examinationofgenitalia.Onthebasisofthe
male genitalia the similarity with S. satyrella
(Staudinger, 1880) is evident. Main diagnostic
characters for the species are the laminar uncus
andthewell-developed,spatulatevalvae.
Description. Wingspan 20 mm. Forewing
brown,withapexverydarkbrownandanarrow,
whitish line subapically at costal margin, one
thirdlongaswing;twowhitishpatchesinmiddle
ofwingsurface:onelongandY-shaped,extended
totwothirdsofwing,thesecondroundedatcell
end. Fringe brownish, darker near apex. Hind-
wingsgreyish,glossy.Fringelong,lightbrown.
Head, thorax and abdomen brown. Legs pale
brown. Antenna brown, three-fourths length of
forewing.
Malegenitalia(Fig.3a–c).UncusY-shaped,
distal arms very short and rounded; gnathos as
longasvinculum,laminar,asymmetrical,withta-
pered and apically pointed distal arm. Valvae Fig.2.–a.S.pamiricasp.n.male(Scalebar=5mm).
symmetrical, spatulate, with narrow proximal –b.S.elburzisp.n.male(Scalebar=3mm).
162 Passerind'Entrèves&Roggero (cid:127) ENTOMOL.FENNICAVol.19
Fig.3.S.pamiricasp.n.,malegenitalia.–a.Uncus-
gnathos-tegumen-juxta-rightvalva-vinculumcomplex.
–b.VIIISternite.–c.Aedeagus.Scalebar=1mm. Fig.4.S.elburzisp.n.,malegenitalia.–a.Aedeagus.
–b.Uncus-tegumencomplex.–c.VIIISternite.–d.
Valvaeandvinculum.Scalebar=1mm.S.paelopyga
Femalegenitalia.Unknown. (Staudinger,1880),malegenitalia.–e.Uncus-tegu-
mencomplex.–f.VIIISternite.–g.Valvaeand
Bionomy.ThespecieswascollectedinJulyat
vinculum.–h.Aedeagus.Scalebar=1mm.
highelevation,inanareacharacterizedbyalpine
vegetation.Hostplantsareunknown.
Distribution. Only known from the Pamir in the description of S. satyrella genitalia,
RangeinnorthernAfghanistan. Bengtsson(1997b:62)perhapsmistookthegna-
Etymology. The species is named after the thosfortheuncus.
Pamirarea,inCentralAsia. We hypothesized that S. pamirica and S.
Remarks.Thespecieshasacloseresemblance satyrellaarebothmorelikelyrelatedtothepalae-
toS.satyrelladescribedfromIran(Shahrud)and arcticthantoafrotropicalandorientalscythridid
Turkmenistan (Ashkrabad and Nukhur, Achal- fauna.Inordertofindotherspecimensbelonging
Tekkeregion),althoughBengtssonquoteditalso tothesamespecies-group,wearenowexamining
from Turkey (1997b:62) and Russia (from scythrididcollectionsfromthesameareatoana-
Sarepta;1997b:223).Likely,thetwospeciesbe- lyzetheirphylogeneticrelationships.
long to the same species-group. They are sepa-
ratedonthebasisofthemarkeddifferenceinfea-
tures of valvae and sternum VIII and different 3.2.Scythriselburzisp.n.
lengthoftheaedeagus,theuncus-gnathos-tegu-
mencomplexbeingsimilar.Itmustbenotedthat Type material. Holotype: # (Fig. 2b): Iran,
ENTOMOL.FENNICAVol.19 (cid:127) Irano-turanianscythridids 163
Table2.AcomparisonofScythrispaelopyga(Staudinger)andS.elburzisp.n.
Structure S.paelopyga S.elburzi
Uncus Short,halfaslongasthetegumen,sharp, Verylong,needle-like,withaglobular
narrowandthorn-like expansionnearthebase
Gnathos Absent Absent
Tegumen Symmetrical,globose,short,withtwo Symmetrical,bilobed,shortandlarge,with
triangularlaminaeatsuperiormargin twoquadrangularplatesatsuperiormargin
Juxta Shell-shaped,concave,withafinger-like Shell-shaped,smallerthaninS.paelopyga
processposteriorly
Aedeagus Bisinuate,narrowingtoapex Rectilinear,downturnedattwothirdsoflength
Valvae Asymmetrical,therightvalvarectilinear Asymmetrical,therightvalvawitha
withatootlikeprocessinwardturnedat fingerlikeprocessrectilinearatapex,the
apex,theleftvalvainwardturned leftvalvatriangularshaped,withtwo
sharpvertices
Vinculum Veryshort,marginrounded Veryshort,marginbisinuate
SternumVIII Rectangular,short,onlyslightlynotchedin Quadrangular,hindmarginwithalargeV-
themiddleathindedge shapednotch,andforemargincarrying
anotchtwicedeeperthanthehindone
TergumVIII Largelysubtrapezoidal,hindmarginwith Quadrangular,theforeanglesrounded
adeep,triangularnotch,withweakly
sclerotizedlateralbulges
Elbursgebirge Kerodj (= Keredji, now Karaj), bilobed. Aedeagus cylindrical, down-arched,
Brandtleg.,20.VI.1936.GenitalSlide1105PdE blunt, narrowing at apex, longer than valvae.
(NHRM). Tergum VIII (not drawn) quadrangular, distal
Diagnosis.Thespeciescanbeidentifiedonly margin rounded, slightly notched; sternum VIII
byexaminationofthegenitalia,diagnosticchar- quadrangular,bilobed,withlargeV-shapednotch
acters being the long and sharp uncus, and the atdistalmargin,andtwicedeepernotchatproxi-
asymmetricalvalvae. malmargin.
Description. Wingspan 16 mm. Forewings Femalegenitalia.Unknown.
evenly dark brown, fringe brown. Hindwings Bionomy. The species was collected in a
verylightbrown,glossy,fringebrownish.Head, mountainous area at an unspecified elevation.
thorax and abdomen dark brown. Legs dark Hostplantsareunknown.
brown, with dense, thick and dark brown vesti- Distribution. Known only from Northern
ture. Antennae dark brown, two thirds long as Iran.
forewings. Etymology. The species is named after the
Malegenitalia(Fig.4a–d).Uncuslongerthan ElburzRange,wherethetypelocalityissituated.
tegumen, narrow, sharp, pointed, with bulge at Remarks. The species is similar to S. paelo-
base.Gnathosabsent.Tegumenveryshort,sub- pyga (Staudinger, 1880) from Amasya, Turkey
triangular,withtworectangularlaminaeatdistal (Fig.4).Thetwospeciescanbeeasilyseparated
edge, extending upward. Valvae asymmetrical, on genital differences (Table 2). For further de-
fusedatproximalmargin;middleofinnermargin tails,seethegenitaliadescriptionofS.paelopyga
withdown-turned,thorn-shapedprocess,similar below.
in both valvae; proximal margin of both valvae ThedistributionofbothS.elburzisp.n.andS.
with inwardly arched, short and thick process; paelopyga shows that the species are related to
leftvalvatwo-thirdsaslongasrightone,triangu- palaearctic fauna more than to afrotropical and
larly shaped; right valva twice as long as wide, oriental ones. Palaearctic species are relatively
narrowing to apex, finger-like and inwardly common in Iran, which is part of a transitional
turned, with tuft of long setae and triangular, zonebetweenPalaearctic,AfrotropicalandOri-
smalllaminaonouterside.Vinculumveryshort, entalRegions(Frisch2006).
164 Passerind'Entrèves&Roggero (cid:127) ENTOMOL.FENNICAVol.19
Fig.5.–a.S.balkhisp.n.male(Scalebar=3mm).
–b.S.brandtisp.n.male(Scalebar=3mm).
3.3.Scythrisbalkhisp.n. Fig.6.S.balkhisp.n.,malegenitalia.–a.VIIITergite
andsternite.–b.Uncus-tegumen-gnathos-aedeagus-
Typematerial.Holotype:#(Fig.5a):NAfghani- valvacomplex,sideview.Scalebar=1mm.S.
stan,Balkh,400ma.s.l.,H.G.Amselleg.,24.V. brandtisp.n.,malegenitalia.–c.Uncus-tegumen-
1956.GenitalSlide836PdE(SMNK). gnathos-valvacomplex,sideview.–d.VIIISternite;
Diagnosis.Easilyconfusedwithotherspecies e.Aedeagus.Scalebar=1mm.
fromtheIrano-turanianRegiononthebasisofex-
ternalfeaturesalone,thisspecieshasdistinctgen- TergumVIIIsubtriangular,distallyrounded,with
italia,withvalvaeveryreducedandtheaedeagus well-developed extrusion at outer side, with
sickle-shaped. deeplyarched,thickenedbase,joinedtosternum
Description. Wingspan 9 mm. Forewings VIII.SternumVIIIsubtriangular,withtwoshort,
whitish with two light brown dots near apex. parallel prongs at apex, and narrow, triangular
Fringe brownish. Hindwings and fringe very notchinmiddleofthickenedproximalmargin.
light brown. Head, thorax and abdomen light Femalegenitalia.Unknown.
brown.Legslightbrown.Antennabrown,almost Bionomy.ThespecieswascollectedinMayat
aslongasforewing,withantennalscapewhitish. lowelevation.Hostplantsunknown.
Male genitalia (Fig. 6a, b). Uncus elongate, Distribution.KnownonlyfromnorthernAf-
bifid, with deep incisure. Gnathos well-devel- ghanistan.
oped,withjoiningarmsnarrowandelongate,dis- Etymology. The species is named after the
tal arm cylindrical, as long as uncus, slightly typelocality,Balkh,innorthernAfghanistan.Itis
down-turned.Tegumenglobose.Valvaesymmet- theoldesttowninAfghanistan,andwasknownin
rical,reduced,roundedatapex,fusedtotegumen. thepastasBactra.
Vinculum very small. Aedeagus shorter than Remarks.Thespeciesverylikelydoesnotbe-
uncus,superiormarginbisinuate,upward-turned longtoanyknownspecies-group,buttheshape
with sharp apex. Juxta short and upturned. of sternum VIII indicates that the new species
ENTOMOL.FENNICAVol.19 (cid:127) Irano-turanianscythridids 165
might be related to the S. subfasciata-group of Remarks.Thespeciesdoesnotseemtobelong
species(K.Nupponen,pers.comm.),orS.mus- toanyknowspecies-group.
groupofspecies.Thetypelocalityliesinaplain
atthefoothillsoftheEasternHinduKushRange,
wherethesemountainsleadinthePamir. 4.Descriptionofmalegenitaliaof
Scythrispaelopyga(Staudinger,1880)
3.4.Scythrisbrandtisp.n. Type material. Lectotype here designated: #,
Turkey,Amasya.“25/7”,“AmasiaM”,“Origin”,
Typematerial.Holotype:#(Fig.5b):NAfghani- “Paelopyga”, “Prep. Gen. 1365 PdE”, “Lecto-
stan, Hindukusch Doab, 1,400 m a.s.l., H. G. typus Butalis paelopyga (Stgr.) Passerin &
Amsel leg., 4.VI.1956. Genital Slide 829 PdE Roggero 2007”. Genital Slide 1365 PdE
(SMNK). (MNHB).
Diagnosis.Thespeciescannotberecognized Diagnosis. The species is similar to many
onthebasisofexternalappearance,butiseasily other scythridids, but the asymmetrical valvae,
identified from its genitalia: sternum VIII with thetegumenwithtwotriangularexpansionsand
twoprolongationsnearthehindmargin,andthe theaedeagusareusefuldiagnosticcharactersfor
fishhook-like aedeagus are unique. Also the thespecies.
valvaeandthecomplexuncus-gnathos-tegumen Description. The external features of S.
arewell-characterized. paelopyga were exaustively described by Stau-
Description. Wingspan 10 mm. Forewings dinger(1880),buttheauthordidnotelucidatethe
lightbrown,withtwowhiteirregularpatchesand characteristic shape of the genitalia. Here, we
transversewhitelineatdistalthird,fringebrown. presentthedrawinganddescriptionofthemale
Hindwingbrownwithbrownfringe.Head,tho- genitalia.
rax and abdomen brownish. Legs very light Male genitalia (Fig. 4e–h). Uncus narrow,
brown. Antenna light brown, almost as long as thorn-shaped, sharp at apex, half long as tegu-
forewing. men. Gnathos absent. Tegumen globular, carry-
Male genitalia (Fig. 6c–e). Uncus laminar, ing at distal edge two asymmetrical, triangular
dome-like.Gnathoswithtriangularmedianpro- laminae with sharp apices upward and outward
cess,verylonganddownwardcurved.Tegumen extending.Valvaeasymmetrical:leftvalvarecti-
narrow,aslongasthevalvae.Valvaesymmetri- linear,withblunttoothinwardlyturnedatexter-
cal, small, triangular, proximal margin rounded nalapex,andtriangularlaminaextendingatdistal
and apex blunt; inner margin inwardly curved half of inner margin; right valva narrower than
withspiniformprocessnearapexatinnermargin. left one, inwardly turned, apex rounded and
Aedeagus three times longer as valvae, narrow, spatular,ovallaminaatmiddleofinnermargin;
strongly curved at basal half, tip hooked. Juxta vinculumshortandrounded.Aedeagusbisinuate,
long. Sternum VIII subtriangular, distally trun- largeratbaseandtaperedtowardapex.Juxtahalf
catedandslightlyemarginate,basethickenedand aslongasaedeagus,triangular,concave.Sternum
deeply notched, two triangular lateral processes VIII rectangular, hind margin rounded and
atdistalthird. slightlynotchedinthemiddle.TergumVIIIsub-
Femalegenitalia.Unknown. trapezoidal, posterior margin deeply and nar-
Bionomy.Thespecieswascollectedinearly rowly notched, with weakly sclerotized lateral
JuneintheHinduKushRange,atrelativelylow bulges.
elevation.Hostplantsandlifecycleareunknown. Bionomy. The species was collected from
Distribution.KnownonlyfromnorthernAf- ApriltoAugust(Staudinger1880).
ghanistan. Distribution. Known in the past only from
Etymology. The species is named after Dr. Amasya (Turkey), the species was collected re-
Brandt, who made expeditions in Iran and col- cently in Aksehir, Sultan Daglari (Central Tur-
lectedmanyinterestingLepidopterafromthatre- key)byK.NupponenatthebeginningofAugust;
gion. thehabitatisaxerothermic,chalkslope;themoth
166 Passerind'Entrèves&Roggero (cid:127) ENTOMOL.FENNICAVol.19
isnocturnalandcomereadilytoartificiallightat Remarks. The Baluchestan constitutes the
night(K.Nupponen,pers.comm.). southernpartoftheSistanandBaluchestanPro-
Remarks. Likely the species belongs to the vince,andislocatedinthesoutheastofIran,bor-
samespecies-groupasS.elburzisp.n.Staudinger deringPakistanandAfghanistan.Itischaracter-
collectedfourmalespecimensinhisTurkishEx- izedbyadesertclimateandisoneofthedriestre-
pedition during 1875 (the lectotype and three gionsofthecountry.
othersinthesamelocalityasthelectotype,10.V.,
3.VI. and 1.VIII.1875); two other male speci- ScythrisfissurellaBengtsson,1997
menswerecollectedfromAmasyaon26.IV.1875 O[st] Afghanistan, Sarobi, 1,100 m a.s.l.,
byJ.Mann,whogavethemtotheauthor,asre- 3.VII.1956,H.G.Amselleg.,1#[Genitalslide
ported by Staudinger (1880: 391), who also 806 PdE]. O[st] Afghanistan, Sarobi, 1,100 m
pointedoutthatoneofthemwasdamaged. a.s.l.,28.VI.1956,H.G.Amselleg.,1$[Genital
slide854PdE].
Distribution.Sudan,Yemen,Oman,Iran,Pa-
5.Newrecords kistanandsouthernRussia(K.Nupponen,pers.
comm.).
ApostibesgriseolineataWalsingham,l907 Remarks. New to Afghanistan. Sarowbi
Afghanistan,Herat,970ma.s.l.,5.V.1956,H. (Surobay)isplacedintheKabulProvince,atlow
G.Amselleg.,1#[GenitalSlide816PdE]. elevationneartheHinduKushRange,inNEAf-
Distribution.Algeria,Tunisia,Libya,andIs- ghanistan.
rael.
Remarks.NewtoAfghanistan.Herat(WAf- Scythrislimbella(Fabricius,1775)
ghanistan)islocatedinthefertilevalleyofHari NIran,Masandaran,Golestan-Wald,60kmE
RudRiver. Ganbad Qabus [=Gonbad-e Kavus, Golestan],
510ma.s.l.,Ebert&Falknerleg.,8.VII.1972,1
Scythriscanescens(Staudinger,1880) # [Genital Slide 374 PdE]. Afghanistan, Herat,
N. Afghanistan, Polichomri, 700 m a.s.l., 970ma.s.l.,25.IV.1956,H.G.Amselleg.,1#
5.VI.1956, H. G. Amsel leg., 5 ## [Genital [GenitalSlide835PdE].
slides810,813,838,839and844PdE]. Distribution.OvermostofEuropetoCentral
Distribution. Morocco, Algeria, Tunisia, Asia.Thespeciesisalsointroducedinthenorth-
Libya,Turkey,Syria,Afghanistan,andPakistan. easternNearcticRegion.
Remarks. Pol-e-Khomri (Baghlan Province) Remarks. The species was already known
islocatedintheNEAfghanistan,neartheHindu from Iran and Afghanistan. Golestan Province
KushRange. (NEIran,southoftheCaspianSea)wasseparated
fromtheprovinceofMazandaranonlyin1997.
Scythriscapitalis(Erschoff,1874)
NO[st] Afghanistan, Khinch-e-Andarab, Scythrismonochreella(Ragonot,1896)
westl. Pamir, 3,550–4,000 m a.s.l., 17.– SO[st] Iran, (Djiroft), Anbar-Abad, 1.–
27.VII.1957,G.Ebertleg.,3##[Genitalslides 18.V.1956,W.Richterleg.,1#[Genitalslide842
289 ZMB, 808 and 3275 PdE], 4 ## and 1 $ PdE].
[Genitalslide3276PdE]. Distribution.Turkey,Syria,Lebanon,Israel,
Distribution. Turkey, Turkmenistan, Tajiki- Egypt,UnitedArabEmirates,Iran,Iraq,andIn-
stan,Uzbekistan,andKyrgyzstan. dia.
Remarks.NewtoAfghanistan. Remarks. Anbar-Abad is located near Jiroft,
inthesouthofKermanProvince,mostofwhichis
ScythrisdecrepidellaBengtsson,1997 largely steppe or sandy desert, with scattered
Iran, Belutschistan, Jranshar, 800 m a.s.l., oases.
11.–21.IV.1954, Richter u. Schäuffele leg., 1 #
[Genitalslide819PdE]. Scythristributella(Zeller,1847)
Distribution.EgyptandsouthernIran. NIran,Masandaran,Golestan-Wald,60kmE
ENTOMOL.FENNICAVol.19 (cid:127) Irano-turanianscythridids 167
GanbadQabus,510ma.s.l.,Ebert&Falknerleg., Hodges,R.W.1998:Gelechioidea.—In:Kristensen,N.P.
8.VII.1972,3##[GenitalSlides386,505,506 (ed.), Lepidoptera, Moths and Butterflies. Vol 1.
HandbuchderZoologie/HandbookofZoology[Wal-
PdE].
terdeGruyter,Berlin&NewYork],IV/35:131–158.
Distribution. Europe, fromSpain to Greece,
Kaila,L.2004:PhylogenyofthesuperfamilyGelechioi-
Russia(SUrals),andLibya dea(Lepidoptera:Ditrysia):anexemplarapproach.—
Remarks.NewtoIran. Cladistics20:303–340.
Landry,J.-F.1991:SystematicsofNearcticScythrididae
(Lepidoptera:Gelechioidea):phylogenyandclassifi-
Acknowledgements. The research was supported by the
cationofsupraspecifictaxa,withareviewofdescribed
grantsfromtheitalianMinisterodellaRicercaScientificae
species.—Mem.Entomol.Soc.Canada160:1–341.
Tecnologica (MURST 60% “Biosistematica dei Lepi-
Larsen, T. B. 1984: The Zoogeographical Composition
dotteriScythrididae”).WeareverygratefultoDr.H.G.
anddistributionoftheArabianButterflies(Lepidop-
Amsel(†)(StaatlicheMuseumfürNaturkundeofKarls-
tera;Rhopalocera).—J.Biogeogr.11:119–158.
ruhe,Germany),andtotheCuratorsofEntomologyCol-
Lederer,J.1869:VerzeichnisdervonHerrnJos.Haber-
lections of the Museum für Naturkunde der Humboldt-
hauerbeiAstrabadinPersiangesammeltenSchmet-
UniversitätofBerlin(Germany)andtheNaturhistoriska
terlinge.—HoraeSoc.Entomol.Ross.6:73–93.
RiksmuseetofStockholm(Sweden)fortheloanofthema-
Macey,J.R.,SchulteII,J.A.,AnanjevaN.B.,LarsonA.,
terial. We would also thank Dr. W. Mey (Museum für
Rastegar-PouyaniN.,ShammakovS.M.&Papenfuss
Naturkunde der Humboldt-Universität of Berlin, Ger-
T.J.1998:PhylogeneticRelationshipsamongAgamid
many)fortheloanofS.paelopygatypematerial,andDr.B.
LizardsoftheLaudakiacaucasiaSpeciesGroup:Tes-
Å. Bengtsson for his useful comments. We are greatly
tingHypothesesofBiogeographicFragmentationand
obliged to Dr. K. Nupponen for his information about
anAreaCladogramfortheIranianPlateau.—Mol.
Irano-turanianscythridids.
Phylogenet.Evol.10:118–131.
Mey,W.2006:RevisionofthegenusPhyllobrostisStau-
References dinger,1859(Lepidoptera,Lyonetiidae).—Dt.Ento-
mol.Zeit.53:114–147.
Amsel,H.G.1949a:DieMicrolepidopterenderBrandt’- Meyrick, E. 1918: Exotic Microlepidoptera II (1916–
schenIran-Ausbeute.1.—Bull.Soc.Fouad.I.Ento- 1923).—Classey,Hampton.640pp.
mol.33:227–269. Meyrick, E. 1924: Exotic Microlepidoptera III (1923–
Amsel,H.G.1949b:OntheMicrolepidopteracollectedby 1930).—Classey,Hampton.640pp.
E.P.WiltshireinIrak,andIranintheyears1935to Nabozhenko,M.V.2006:ArevisionofthegenusCatomus
1938.—Bull.Soc.Fouad.I.Entomol.33:271–351. Allard,1876andthealliedgenera(Coleoptera,Teneb-
Akhani,H.,Trimborn,P.&Ziegler,H.1997:Photosynthe- rionidae)fromtheCaucasus,MiddleAsia,andChina.
ticpathwaysinChenopodiaceaefromAfrica,Asiaand –Entomol.Rev.86:1024–1072.
Europewiththeirecological,phytogeographicaland Nye,I.W.B.&Fletcher,D.S.1991:ThegenericNamesof
taxonomicalimportance.—Pl.Syst.Evol.206:187– MothsoftheWorld.Volume6.Microlepidoptera.—
221. NaturalHistoryMuseumPublication,London.368pp.
Bengtsson,B.Å.1997a:ScythrididsfoundsbyH.Hacker Passerin d’Entrèves, P. 1976: Revisione degli Scitrididi
inPakistan1988,andIndia1992.—Esperiana4:467– (Lepidoptera, Scythrididae) paleartici. II. I tipi di
481. ScythrisdelMuséumNationald’HistoireNaturelledi
Bengtsson,B.Å.1997b:Scythrididae.Microlepidoptera Parigi.—Boll.Mus.Zool.Univ.Torino1976:27–70.
ofEurope.Vol.2.—ApolloBooks,Stenstrup.301pp. Passerind’Entrèves,P.&Roggero,A.2007:Scythrididae.
Bengtsson,B.Å.&Huemer,P.2003:EineneueArtderFa- Fascicle44.—In:Heppner,J.B.(ed.),Lepidoptero-
milie Scythrididae aus dem Iran (Lepidoptera, rum Catalogus (New Series) Vol 3. Association for
Scythrididae).—CarinthiaII193/113:573–578. TropicalLepidoptera,Gainesville.86pp.
Brandt, W. 1938: Beitrag zur Lepidopteren-Fauna von Ribera, I. & Blasco-Zumeta, J. 1998: Biogeographical
Iran.—Entomol.Rdsch.55:698–699. linksbetweensteppeinsectsintheMoneagrosregion
Christoph,H.1885:LepidopteraausdemAchal-Tekke- (Aragón,NESpain);theEasternMediterranean,and
Gebiet.2.Teil.—In:N.M.Romanoff.Mémoiressur CentralAsia.—J.Biogeogr.25:969–986.
lesLépidoptères.St.Petersburg.2:119–171. Sruoga, V. & Puplesiené, J. 1998: Contribution to the
(cid:1)iplak,B.2003:DistributionofTettigoniinae(Orthoptera, knowledge of Central asiatic Microlepidoptera with
Tettigoniidae)bush-cricketsinTurkey:theimportan- descriptionofnewspecies.—ActaZoologicaLitua-
ceoftheAnatolianTaurusMountainsinbiodiversity nicaEntomologia8:38–48.
andimplicationsforconservation.—Biodiversityand Staudinger,O.1880:Lepidopteren-FaunaKleinasien’s.—
Conservation12:47–64. HoraeSoc.Entomol.Ross.15:159–435.
Frisch,J.2006:ThegenusScopaeus(Coleoptera,Staphy- Wiltshire,E.P.1957:TheLepidopteraofIraq.—Ministry
linidae,Paederinae)inIran,withdescriptionofnew of Agricolture, Government of Iraq. Kaye Limited,
speciesfromtheZagrosMountains.—Dt.Entomol. London.162pp.
Zeit.53:5–22.