New species and new records of Calliostomatidae (Gastropoda: Trochoidea) from New Caledonia and Solomon Islands

2009·17.8 MB·English

by  Claude Vilvens

C. VlLVENS Novapex 10(4): 125-163, 10 décembre 2009
New species and new records of Calliostomatidae
(Gastropoda: Trochoidea)
from New Caledonia and Solomon Islands
Claude VlLVENS
Rue de Hermalle, 1 13 - B-4680 Oupeye, Belgium
Scientific Collaborator, Muséum national d'Histoire naturelle, Paris.
vilvens.claude(âskynet.be
KEYWORDS. Gastropoda, Calliostomatidae, New Caledonia, Solomon Islands, Calliostoma,
Bathyfautor, Thysanodonta, new species.
ABSTRACT. New records of 16 known Calliostomatidae species from New Caledonia and
Solomon Islands area are listed, extending the distribution area of some of them. Seven new
species are described and compared with similar species: Calliostoma (Calliostoma) cochlias n.
sp., C. (Fautor) aprosceptum n. sp., C. (F.) diaphoros n. sp., C. (Benthastelena) hexalyssion n. sp.,
C. (B.) malaita n. sp., C. (Ampullotrochus) tropis n. sp., C. (A.) aporia n. sp. A list of the
Calliostomatidae ofthe Indo-Pacific area is provided with theirdistribution.
RESUME. De nouveaux relevés de 16 espèces connues de Calliostomatidae provenant de
Nouvelle-Calédonie et des îles Salomon sont listés, étendant ainsi l'aire de distribution d'un certain
nombre d'entre elles. Sept nouvelles espèces sont décrites et comparées avec des espèces
similaires : Calliostoma (Calliostoma) cochlias n. sp.. C. (Fautor) aprosceptum n. sp., C. (F.)
diaphoros n. sp., C. (Benthastelena) hexalyssion n. sp., C. (B.) malaita n. sp., C. (Ampullotrochus)
tropis n. sp., C. (A.) aporia n. sp. Une liste des Calliostomatidae d'Indo-Pacifique est fournie avec
leurdistribution.
INTRODUCTION The présent paper gives a synthetic report on the
Calliostomatidae species collected during thèse most
New Caledonia. and the adjacent Vanuatu and Fiji récent (cited above) expéditions involving MNHN
Islands, has reveal numerous new calliostomatid scientists in New Caledonia and Solomon Islands
species since thèse 15 last years: Marshall (1995b) areas. Besides new records for 24 known species,
described not less than 27 new species (Calliostoma, which sometimes extends their distribution and/or
Thysanodonta, Laetifautor, ...); Vilvens (2005) added introduces some variability ofshell characters, 7 new
5 Calliostoma species to this large list (extending to species are described hère.
Tonga Islands); finally Vilvens & Maestrati (2006) What also makes fundamental the papers of Marshall
described 3 new Thysanodonta from this area. (1995a. 1995b) is his systematic work, after having
Ail thèse descriptions were based on a large material created the subfamily Thysanodontinae (Marshall.
coming from the French expéditions (Héros et al., 1988) : clarification ofthe numerous subgenera ofthe
2007; Vilvens. 2007; Bouchet et al., 2008; Bouchet et genus Calliostoma (while placing some of them in
al., 2009) conducted ail around this area for the last synonymy), description of new gênera {Bathyfautor.
thirty years by IRD (Institut de Recherche pour le Dactylastele, Selastele, Fautrix) and interprétation of
Développement, Paris - ex-ORSTOM) and MNHN Maurea as a subgenus ofCalliostoma. This important
(Muséum national d'Histoire naturelle, Paris) works were used, among others, to build a provisional
:
MUSORSTOM and BATHUS, BERYX, BIOCAL, list of the Calliostomatidae of the Indo-Pacific area,
BIOGEOCAL, BORDAU, CALSUB, CHALCAL, with their respective distribution, that is provided as
CORAIL, HALICAL, HALIPRO, LAGON, LIFOU, appendix at the end ofthis paper.
LITHIST, MONTROUZIER, NORFOLK, PALEO-
SURPRISE, SMIB. VOLSMAR. However, the Material and methods
calliostomatids material brought by the most récent
campaigns (NORFOLK 2, CONCALIS) was not yet The material studied in the présent paper was brought
studied. by 5 IRD-MNHN expéditions: NORFOLK 2,
Unlike New Caledonia, no régional study about CONCALIS, SALOMON 1, SALOMON 2 and
calliostomatids (and even Trochoidea as far as I know) SALOMONBOA 3.
from the Solomon Islands has been published in the As the continuation of the NORFOLK 1 campaign
last years. The spécifie campaigns leaded again by (6/2001), the NORFOLK 2 campaign (10-11/2003)
MNHN and IRD (especially SALOMON 1 & 2, took place around Norfolk Ridge and was mainly
SALOMONBOA 3) were therefore welcome to fill dedicated to the study of speciation and endemism
this gap. phenomena on the seamounts. The CONCALIS cruise
125
C Vil \l \s Calliostomatidae from New Caledonia and Solomon [slands
(4-5/2008) took place in the Grand-Passage and 112004) thaï explored the western part of the
Surprises [slands north off New Caledonia's main Solomon [slands, with spécial care for the hard
m SALOMON
Island and lu focused on deep-water cônes for their bottoms (a bit neglected by I ) and for the
venom glands. But, among other interesting biological l'aima assoeiated to the "sunken woods". This new
material, il also provided an additional valuable interesting topie of the récent years has justified the
malacological material, among which some BOA ("Bois coulés et Organismes Associés [Sunken
rrochoidea (and especialk Calliostomatidae) were Wood and Associated Organisms]) cruises séries:
found. BOA (1 1/2004), BOA 1 (9/2005) and SANTOBOA
Ihe SALOMON 1 campaign (9-10/2001) surveyed (10/2006) off the larger islands in northern Vanuatu
the deep waters off the central part of the Solomon and southern New Caledonia, and SALOMONBOA 3
[slands (from Guadalcanal to Malaita and Makira). (9-10/2007) lhat operaled off the larger Ibrested
lins survej was continued by SALOMON 2 (10- islands ofGuadalcanal and Malaita (Samadi, 2009).
155° 60e 165 70e 175(
5
'
-»••»»
SALOMON 2 » i 4
Atn. *
SALOMON
Tf
1
SOLOMON ISLANDS
'*** *"k TU
\ -,
I
-i
15* i «V»!^
*AHHUATU
(ON( AL1S
< ' \ »*>
20e o
S. îiflfc
if N o t !
B a s < n
mmC**ào**Vi ***** "t\ mtâ
%mm
— — — c FUI
.
25 i X
l
'PACIFORIFOLKC2
n0
*=r*S«£«r !•«•
t h
Map 1 : Prospecting areas surveyed in this paper- approximate locations ofNORFOLK, CONCALIS and
SALOMON
cruises.
126
C. VlLVENS Novapex 10(4): 125-163, 10 décembre 2009
The range ofthe known and new species is provided gênerai shape ofthe shell (depressed, high spired
forail the available spécimens and also forthe only - conical, cyrtoconoidal, coeloconoidal);
living spécimens ifthey hâve been found; when thèse shape ofthe vvhorls (convex, concave, straight -
ranges are the same, the common range is cited once with orwithout shoulderorkeel);
with the "(living)" annotation; ifail the spécimens are spiral cords ofthe whorls (onthogeny, number,
dead collected. the range is cited with the "(dead)" beads, strength);
annotation. Regarding the description methodology, spiral cords on the base (number. beads, distance
the main conchological features used are (see Text between);
Figure 1 below): shape ofthe aperture. the outerand the inner lip.
Text Figure 1 : Features ofCalliostomatidae shells: H : height; W : width; HA : height ofthe aperture; PI, P2,
P3, ... : primary cords; SI, S2, S3, ... : secondary cords; Tl, T2. ... : tertiary cords (shell : Calliostoma (Fautor)
chlorum Vilvens, 2005, Fiji, BORDAU 1, stn DW1454, 13.6 x 10.4 mm).
Abbreviations BMNH Natural History Muséum. London. England.
:
NSMT National Muséum ofScience, Tokyo, Japan.
:
Repositories RMBR Raffles Muséum of Biodiversity Research,
:
IMT Institute of Malacology of Tokyo, Tokyo, Singapore.
:
Japan. ZMA Zoologisch Muséum, Amsterdam. The
MNHN Muséum national d'Histoire naturelle, Paris, Netherla:nds.
:
France.
127
C \ II \l \s Calliostomatidae from New Caledonia and Solomon [slands
Other ahbrt-xiations SALOMONBOA 3. stn CP2816, 09°49'S, 161 32'H,
464-819m.
Il : heighl
w width Material examined. Solomon Islands.
SALOMONBOA
11 \ heighl ofthe aperture 3: stn CP2816, 09°49'S, 161°32'E,
I \\ numberofteleoconch whorls 464-819 m. I l\ (holotype).
PI. P2. P3,...: primai} cords (PI is the most adapical)
Distribution. Only known from the type locality.
SI. S2, S3,...: secondai-) cords (SI is the mosl
adapical) Diagnosis. A typical rather large Calliostoma species
II. 12. T3,...: tertiary cords (numbered following
with a rather elevated, conical spire, six strong,
appearing order)
prominent spiral cords on the last whorl, adapical
stn station
: granular cords and abapical smooth cords, an almost
l\ : live-taken spécimens présent in sample liât base with a large médian smooth area bordered by
dd no live-taken spécimens présent in sample
: an external and three internai spiral cords, without
sub subadult spécimen
: umbilicus.
juv juvénile spécimen
:
Description. Shell ofmédium large size for the genus
SYSTEMATICS (height 17.0 mm, width 15.8 mm), higher than wide,
conical in shape; spire elevated, height l.lx width,
We tbllow hère the classification of Marshall (1995), 3.3x aperture height; angulate periphery;
used by Bouchet & Rocroi (2005), where anomphalous.
Calliostomatidae, earlier treated as a subfamily of Protoconch 400 pm wide, of whorl, rounded,
1
Trochidae (Hickman & McLean, 1990), are ranked as covered by a network of ridges producing polygonal
a family of superfamily Trochoidea (besides true areas; apical fold nearly straight with a rounded,
Trochidae and Solariellidae), vvith the two subfamilies clearly visible terminal varix.
C'alliostomatinae and Thysanodontinae. Teleoconch of 7.0 whorls; whorls straight except
convex last whorl, without keel. Suture visible, not
TROCHOIDEA
Superfamily : Rafinesque, 1815 canaliculated. First whorl convex, sculptured by axial,
Family : CALLIOSTOMATIDAE Thiele, 1924 weakly prosocline, strong threads and 3 spiral cords;
Subtamily : CALLIOSTOMATINAE Thiele, 1924 PI, P2 and P3 appearing immediately, ail granular by
Tribe Calliostomatini Thiele, 1924 [= Ziziphininae intersection with axial threads; PI weaker than other
:
Gray, 1847] cords; P2 closer to PI than to P3; interspace between
Genus : Call'wstoma Swainson, 1840 threads 2.5x larger than threads; P4 almost completely
Subgenus : Call'wstoma (s. s.) Swainson, 1840 hidden by succeeding whorl. On second whorl, PI, P2
Type species Trochus conulus Linnaeus, 1758 (by and P3 similar, granular; axial threads slightly
:
s.d. Herrmannsen, 1846)- Récent, Mediterranean Sea. broader, distance between 3x larger than threads. On
third whorl, PI strongest at mid whorl, with large,
Call'wstoma (Call'wstoma) codifias n. sp. round beads; P2 weakest; SI and S2 appearing at end
Colour Figs B1-B2, Figs 1-2 ofwhorl, thin, granular; S3 absent.
On fourth whorl, beads ofP3 and S2 obsolète, the two
Type material. Holotype (17.0 x 15.8 mm) MNHN cords smooth after mid whorl, P3 making keel; PI
(MNHN 22073). strongest, P3 weaker than PI but stronger than other
cords; SI and P2 similar in strength, granular; axial
Type locality. Solomon Islands, South ofMalaita, sculpture weakening and disappearing on abapical
part.
Colour plate 1. Figures A1-G2. Holotype MNHN ofthe new species described. Scale bars: 5 mm.
Al-2. C. (Benthastelena) hexalyssion n. sp., Solomon Islands, 460-487 m, 22.2 x 18.1 mm; Bl-2. Calliostoma
(Calliostoma) cochlias n. sp, Solomon Islands, 464-819 m, 17.0 x 15.8 mm; Cl-2. Calliostoma (Benthastelena)
malaitan. sp., Solomon Islands, 487-541 m, 20.7 x 227 mm; Dl-2. Calliostoma (Ampidlotrochus) tropis n. sp.,
Solomon Islands, 309-328 m, 15.9 x 12.9 mm; El-2. Calliostoma (Fautor) diaphoros n. sp., Solomon Islands,
307-310 m, 1 1.2 x 9.1 mm; Fl-2. Calliostoma (Ampidlotrochus) aporia n. sp., Solomon Islands, holotype
MNHN (MNHN 20082), 105-128 m [SALOMON 2, stn CP2294], 1 1.2 x 8.8 mm; Gl-2. C. (Fautor)
aprosceptum n. sp., NorthernNew Caledonia, holotype MNHN (MNHN 20074), 380-430 m [CONCALIS, stn
DW2943], 12.5x9.9 mm.
128
C. VlLVENS Novapex 10(4): 125-163, 10 décembre 2009
129
C \ Il \ I \s Calliostomatidae from New Caledonia and Solomon Islands
On fifth whorl, beads of V2 obsolète; axial sculpture similar in size to other cords except S3, a higher ll/W
also obsolète on adapical part. On sixth whorl, P2 ratio ( 1.2 to 1.3) with a slightly concave shape.
nearl) smooth; beads ofSl weakening. On last whorl. The new species may be compared to C.
P4 l'ullv visible, peripheral, smooth; SI nearl) (Benthastelena) margaritissimum (Habe & Okutani,
smooth.; ail spiral cords similar in size, except PI 1968) (Figs 11-12) from Midvvay (described as
stronger; PI >.ti11 granular, with beads weaker and belonging to the subgenus Trîstîchotrochus, synonym
louer than on preceding whorls; interspace between ofBenthastelena), but this similar in size species hâve
cords similar in size to cords in abapical part, about primary spiral cords ail granular, the primary cords
1.5x to 2\ size ofcords in adapical part. much stronger than secondary ones, a rounded
Aperture subelliptic; outer lip thin. slightly curved; periphery and 8-9 evenly spaced on spiral cords the
basai part rounded. producing a round angle with base.
outer lip and an obtuse angle at meeting point with The gênerai shape of C. cochlias n. sp. reminds
inner lip. Calliostoma (B.) uranipponensc Okutani, 1969 (Figs
Columella weakly arcuated, almost vertical, without 3-4) from Japan (originally described in the subgenus
tooth; eallus completely covering umbilicus. Tristichotrochus), but although this species has a
Base weakly convex. almost flat, with an extemal comparable ontogeny ofprimary and secondary cords
subgranular spiral cord, 1 to 3 internai, nearly smooth, (but SI and S2 appear later), the spiral cords are ail
closely packed cords around the umbilical area and a granular, tertiary cords are présent and the base has
large intermediate smooth zone. mainly about 12 subgranular, evenly spaced spiral
Colourofteleoconch and protoconch pinkish white. cords.
Operculum corneous, circular, multispiral with a short The new species also reminds C. (Fautor) takujii
growing edge, brown. Kosuge, 1986 (Figs 13-14) from Japan, but if this
species has also a PI that is the most prominent, ail
Discussion. The placement of the new species in the cords are granular, the ontogeny of cords is
Calliostoma s. s. is justified by the strong, mainly différent (PI appearing later, S3 présent) and the base
smooth or subgranular spiral cords, the weak axial has about 10 evenly spaced granularspiral cords.
sculpture and the absence ofumbilicus.
The combination ofconical shape, prominent, smooth Etymology. Screw (Ancient Greek : x°X^-iaÇX used
spiral cords except PI and almost flat, smooth base as a noun in apposition - with référence to the regular
(except 1 external and 3 internai smooth spiral cords) spiral sculpture on the whorls.
makes the new species rather hard to confuse with
other existing species from the same area, except Calliostoma (Calliostoma) vilvensi Poppe, 2004
maybe Calliostoma trotini Poppe, Tagaro & Dekker, Colour Figs 11-12, Figs 15-18
2006 (Figs 7-8) from the Philippines, but this smaller
species has a deep umbilicus and 8 subgranular to Calliostoma vilvensi Poppe, 2004: 6-8, pi.2, figs 2a-
nearly smooth, evenly spaced spiral cords on the base. 2b-2c. Type locality: Philippines, off Aliguay Island,
Calliostoma cochlias n. sp. is resembles C. (C.) 80-200 m.
occidentale (Mighel & A.Adams, 1842) (Figs 9-10) Calliostoma vilvensi -- Poppe, Tagaro & Dekker,
from northern Atlantic, but this slightly smaller 2006: 127 pi. 66 figs 1-2.
species has more convex whorls, usually only 4 strong
primary cords, SI appearing occasionally but much Material examined. Solomon Islands. SALOMON
later (5' whorl) and absence ofS2, a narrower smooth 2: stn CP2234, 06°51'S, 156°24'E, 182-277 m, 1 dd. -
area on the base with two external cords. Stn CP2237, 06°53'S, 156°22'E, 400 m, 4 dd, 1 juv dd.
Calliostoma cochlias n. sp. may be compared to
Bathyfautor rapuhia Marshall, 1995 (Figs 5-6) from Distribution. Philippines, Aliguay Island, 80-150 m
New Zealand and Norfolk Ridge, but this taller (range computed from Poppe et al., 2006); Salomon
species has a prominent S3 making keel, a smooth PI Islands, 82-83 m.
Colour plate 2. Figures Hl-02. Species with an extension ofthe distribution. Scale bars: 5 mm.
Hl-2. Calliostoma (Ampullotrochus)peregrinum Marshall, 1995, Norfolk Ridge, 370-371 m, 26.0 x 20.6 mm;
11-2. Calliostoma (Calliostoma) vilvensi Poppe, 2004, Solomon Islands, 400 m, 12.0 x 1 1.4 mm;
Jl-2. Calliostoma (Ampullotrochus)xanthos Marshall, 1995, Norfolk Ridge, 405-455 m, 14.3 x 10.1 mm;
Kl-2. Calliostoma (Ampullotrochus)philippei Poppe, 2004, Solomon Islands, about 258-283 m, 14.7 x 10.5
mm; Ll-2. Thysanodontacassis Vilvens & Maetrati, 2006, Norfolk Ridge, 680-980 m, 13.0 x 8.9 mm;
M1-M2. Calliostoma (Fautor) scobinatum (A.Adams in Reeve, 1863), Solomon Islands, 82-83 m, 7.5 x 5.9 mm;
Nl-2. Calliostoma (Fautor) boucheti Marshall, 1995, northern New Caledonia, 350 m, 9.9 x 8.4 mm;
01-2. Calliostoma (Fautor)periglyptum Marshall, 1995, Solomon Islands, 180-187 m, 7.1 x 5.9 mm.
130
C. VlLVENS Novapex 10(4): 125-163, 10 décembre 2009
131
C. Vil \ I \s Calliostomatidae from New Caledonia and Solomon Islands
Remarks, lhe type material ofthis species was oot Remarks. The dead spécimen from northern New
available bul us author lias provided accurate pictures Caledonia is eroded, but shows the correct ontogeny.
ofthe type material and recentl) collected spécimens, Il implies an extension of the distribution, allhough
rhe two shells from the Solomon Is. arc smaller additional material is elearly needed to draw a firm
(height up to 12.2 mm) than the types from the conclusion.
Philippines (height up to 22.0 mm), but they lune
about 2 whorls less. They share the same cords Calliostoma (Fautor) metivieri Marshall, 1995
ontogenj (PI. P2 and P3 starting immediately, SI Figs 33-34
appearing al 2"J whorl, S2 absent and S3 starting at 3r
whorl, the 5 cords quickly o\' the same si/e on next Calliostoma (Fautor) metivieri Marshall, 1995: 397-
whorls). Poppe did not plaee his new species into a 398. figs 16-18, 120, 153. Type locality: southern
New Caledonia, 400-420 m.
subgenus. Taking the absence of axial sculpture and
the blunt shape of the beads ofcords into account, it Material examined. Norfolk Ridge. NORFOLK 2:
sCaelelmisosttohamta s.thsitsr..species belong to the subgenus sSttnn DDWW22031039,, 2233°°4379''SS,, 116687°°1473''EE,,42423-04-94550m,m,3 d1d.lv. -
Subgenus : Fautor Iredale, 1924 Distribution. South of Ile des Pins, southern New
Type species : Ziziphinus comptus A.Adams, 1855 (by Caledonia and northern Norfolk Ridge, 390-535 m,
o.d.) - Récent, southern Australia. living at 415-535 m (range computed from Marshall,
1995).
Calliostoma (Fautor) boucheti Marshall, 1995
Colour Figs N1-N2. Figs 21-24 Calliostoma (Fautor)paradigmatum Marshall, 1995
Figs 25-26
Calliostoma (Fautor) boucheti Marshall, 1995: 389-
392, figs 3, 7-9. 117, 150, 155. Type locality: off Calliostoma (Fautor) paradigmatum Marshall, 1995:
southern New Caledonia, 600 m. 395-397, figs 13-15, 1 19, 155. Type locality: southern
Calliostoma (Fautor) boucheti- Vilvens, 2005: 2, figs New Caledonia, 505-550 m.
7-8. Calliostoma (Fautor) paradigmatum - Vilvens, 2005:
2.
Material examined. Norfolk Ridge. NORFOLK 2: Material examined. Norfolk Ridge. NORFOLK 2:
Ssttnn DDWW22005670., 2244°°4400''SS,, 116688°°3398''EE,, 555852--566050mm,, 71 llvv.. -- dstdn. C-P2S0tn47D,W2230°6439',S,251°6280°'0S2,'E,16785°958-'8E0,77m9,5-18l5v2, 1m,juv
CSSttOnnNCDDAWW22L00I88S74:,,st22n44°°D55W62''2SS9,,7911,6688°°122822°''1EE6,,'S,55181866--257°835604'mmE,,, 4335ll0vv..m,-- llvv..--SSttnnDDWW22111006,,2233°°4584''SS,, 116687°°1472''EE,, 469835--875507mm,, 111
dd.
1 dd.
Distribution. Off Ile Surprise, northern New
Distribution. South of Ile des Pins, southern New Caledonia, 585 m (living); South of Ile des Pins,
Caledonia and northern Norfolk Ridge, 233-650 m, southern New Caledonia, northern Norfolk Ridge,
living at 503-650 m (range computed from Marshall, 470-795 m, living at 550-795 m (range computed
1995); northern New Caledonia, 350 m; Vanuatu, using data of Marshall, 1995); Tonga, 342-500 m
585 m. (dead).
Figures 1-14. Scale bars: 5 mm.
1-2. Calliostoma (Calliostoma) cochlias n. sp., holotype MNHN (22073), Solomon Islands, 464-819 m
[SALOMONBOA 3, stn CP2816], 17.0 x 15.8 mm; 3-4. Calliostoma (Benthastelena) margaritissimus (Habe &
Okutani, 1968), holotypeNSMT 38669, Midway Island, 400-460 m, 15.9 x 15.8 mm; 5-6. Bathyfautorrapuhia
Marshall, 1995, paratype MNHN, New Caledonia, Wanganella Bank, 660-757 m [NZOI, stn P8], 20.1 x 15.4
mm; 7-8. Calliostoma trotini Poppe, Tagaro & Dekker, 2006, holotype NMP, Philippines, Palawan Is., 1 1.8 x
1 1.3 mm (© Conchology, Inc.); 9-10. Calliostoma (Calliostoma) occidentale(Mighel & A.Adams, 1842),
eastern Canada (Digby, Nova Scotia), C.Vilvens coll., 12.5 x 1 1.5 mm; 11-12. Calliostoma (Benthastelena)
uranipponensis Okutani, 1969, holotype NSMT (Mo 69544), Japan, depth unknown, 10.1 x 8.8 mm;
13-14. Calliostoma (Fautor) takujii Kosuge, 1986, holotype IMT (IMT-86-1), Japan, Ogasawara (Bonin)
Islands, 12.0 x 9.7 mm.
132
C. VlLVENS Novapex 10 (4): 125-163, 10 décembre 2009
133
C \ Il M \s Calliostomatidae from New Caledonia and Solomon [slands
Calliostoma (Fautor)periglyptum Marshall. 1995 Distribution. Northern New Caledonia, 255-300 m
Colour Figs01-02, Figs39-44 (living)(rangecomputed from Marshall, 1995).
Calliostoma (Fautor) periglyptum Marshall. 1995: Calliostoma (Fautor) aprosceptum n. sp.
401-402, figs 25-27. 123. 157. Type loeality: southem Colour Figs G1-G2, Figs 29-32, Table 1
Non Caledonia. 230 m.
Type material. Holotype (12.5 x 9.9 mm) MNHN
Material examined. Norfolk Ridge. NORFOLK 2: (MNHN 20074). Paratypes: 3 MNHN (MNHN
stn DW2093, 24 44'S, l68o09'E, 230 m, 1 dd. - Stn 20075). 1 coll. C.Vilvens.
SDoWl2o1m2o3n, l2s3la1n8d'sS., S16A8L1O5'ME,ON1871-:197stmn, DIW1d7d5.8, Type loeality. Northern New Caledonia, southem
08 49'S, 159°52'E, 180-1X7 m. 2 dd sub. slope of Grand Passage, CONCALIS, stn DW2943,
SALOMON 2: stn CP2234, 06°51'S, 156°24'E, 18°56.8'S, 163°22.7'E, 380-430 m.
182-277 m. 2 dd sub.
Material examined. Northern New Caledonia.
CONCALIS: stn DW2943, 18°57'S, 163°23'E, 380-
Distribution. Loyalty lslands (off Lifou), 280-520 430 m, dd (holotype). - Stn DW2979, 18°16'S,
(fRdiredoagmde),;Mas1ro9sn7hta-hl2el7r,4n mN1,9e9w5l)i;vCianSlgeodlaotonm2ioa3n0anmldsl(narnoadrsnt,ghee1mc8o2Nm-op1ru8f7toelmdk D1186W°231°065'40S'3,E,,1163215°805°73m'4,E',S,157l4v1-663(6°p00a8rma',tEy,p1e)d4.d50-(-p4aSr5ta4ntymp,De)W.29-8S0dt,dn
1
(dead). (paratype). - Stn DW3025, 18°57'S, 163°23'E, 396-
Remarks. This species was so far known only from 400 m, 1 dd, 1 sub lv (with paratype).
New Caledonia and the Loyalty lslands. The Distribution. Northern New Caledonia, 350-574 m,
spécimens from Solomon lslands are ail immature (up living at 350-396 m.
to 6 teleoconch whorls for the largest one), having
about 12 smooth spiral cords on the base; ail match Diagnosis. A typical Calliostoma species with a rather
the other distinctive features ofthe species (mainly S2 elevated, more or less conical spire, a strong
appearing at 4,h whorl, SI between 5th and 7lh whorl, subsutural granular spiral cord, a second strong
S3 absent). This is a significant distribution extension granular spiral cord making keel under the middle of
that should need confirmation with living spécimens. the intermediate whorls, last whorls convex with
numerous granular spiral cords, a weakly convex base
Calliostoma (Fautor) necopinatum Marshall, 1995 with about 13 granular spiral cords, without
Figs 19-20 umbilicus.
Calliostoma (Fautor) necopinatum Marshall, 1995: Description. Shell of médium size for the genus
394-395, figs 31-33, 121, 155. Type loeality: northern (height up to 14.2 mm, width up to 1 1.2 mm), higher
New Caledonia, 255 m. than wide, conical in shape; spire rather elevated,
height 1.2x to 1.3x width, 3.3x to 3.8x aperture height;
Material examined. Northern New Caledonia. angulate periphery; anomphalous.
CONCALIS: stn DW2977, 18°15'S, 163°07'E, 326- Protoconch about 250 um wide, of 1 whorl, rounded,
368 m, 1 dd. - Stn DW2978, 18°16'S, 163°04'E, 360- covered by a network of ridges producing polygonal
400 m, 2 dd. - Stn DW3003, 18°34'S, 163°08'E, 450- areas; apical fold straight with a very thin rounded
454 m, 1 dd. terminal varix.
Figures 15-28. Scale bars: 5 mm.
15-18. Calliostoma (Calliostoma) vilvensi Poppe, 2004.
15-16. Holotype, Philippines, offAliguay Island, 80-200 m, 20.3 x 19.2 (© Conchology, Inc.);
17-18. Solomon lslands, 400 m [SALOMON 2, stn CP2234], 12.0 x 1 1.4 mm.
19-20. Calliostoma (Fautor) necopinatum Marshall, 1995, Northern New Caledonia, 326-368 m [CONCALIS,
stnDW2977], 11.2x8.7 mm.
21-24. Calliostoma (Fautor) boucheti Marshall, 1995.
21-22.Norfolk Ridge, 586-730 m [NORFOLK 2, stn DW2084], 12.2 x 10.6 mm; 23-24. Northern New
Caledonia, 350 m [CONCALIS, stn DW2979], 9.9 x 8.4 mm.
25-26. Calliostoma (Fautor)paradigmatum Marshall, 1995, Norfolk Ridge, 795-852 m [NORFOLK 2, stn
DW2069], 12.5 x 10.4 mm; 27-28. Calliostoma (Fautor) houbrickiMarshall, 1995, paratype MNHN, southem
New Caledonia, 170 m [Chalcal 2, stn DW84], 10.5 x 8.6 mm.
134