New records of Flabellinidae (Opisthobranchia: Aeolidacea) from the tropical Americas, with descriptions of two new species

13 Pages·1994·4.9 MB·English

PROCEEDINGS
OF THE
CALIFORNIA ACADEMY OF SCIENCES
Vol. 48, No. 9, pp. 171-183, 10 figs. August 16, 1994
NEW RECORDS OF FLABELLINIDAE
(OPISTHOBRANCHIA: AEOLIDACEA) FROM THE
TROPICAL AMERICAS, WITH DESCRIPTIONS OF TWO
NEW SPEOH^-gguj*^
By
Terrence M. Gosliner
CaliforniaAcademyofSciences, Golden GatePark,SanFrancisco, California94118
Abstract: TwonewspeciesofFlabellinaaredescribedfromthetropicalAmericas.Flabellinavansyocisp.
nov.iswidelydistributedfromBahiaMagdalenaonthePacificcoastofBajaCaliforniatotheGulfofChiriqui.
Panama.Itiscomparedtoitscloserelative,F.pedata(Montagu, 1815),whichisfirstrecordedherefromthe
AzoresIslands.Flabellinahamannisp.nov.hasbeenfoundfromtheBahamastoVenezuelaintheCaribbean
and is mostcloselyrelatedtoF. marcusorumGoslinerand Kuzirian, 1990.FlabellinabertschiGoslinerand
Kuzirian, 1990,previouslyrecordedonlyfromtheGulfofCalifornia,isalsofoundalongthePacificcoastof
Panama.Flabellinamarcusorum,was knownpreviouslyfromtheGulfofCaliforniaandtheCaribbeanand
isherereportedforthefirsttimefromPanama.FlabellinastohleriBertschandFerreira, 1974,isconsidered
ajuniorsynonymofF. telja Marcusand Marcus, 1967.TherangeofF. teljaisextendedsouthward to the
IslasRevillagigedos,Panama,and theGalapagos.
ReceivedOctober 13, 1993.AcceptedDecember3, 1993.
INTRODUCTION Material.—CASIZ088179,onEudendriumsp.,22mdepth,
HillRocks(7°17'20"N,8P40'65"W),SWofPuntaAnegada,
Recently, Gosliner and Kuzirian (1990) re- Is"laCoiba,GulfofChiriqui,PacificcoastofPanama, 18April,
viewedtheFlabellinidaefromthetropicalcoasts ' 3-
oftheAmericas.BoththisworkandthatofGos- Distribution.-Flabellina bertschi is known
liner and Willan (1991) examined the phyloge- wimin the GulfofCalifornia from Puerto Pen-
neticrelationshipswithinFlabellina. Sincethen, asc0 soum t0 Isla San Dieg0 (Gosiiner and Ku-
additional collections ofmatenal from both the zirian {990) and from the Pacific coast ofPan.
easternPacificandCaribbeanhaveyieldedspec- ama
riemceonrsdiosffFtowrousnodmeescprreivbie•odusslp,yecineas,measd,wte.alxla.asT-yn.h,ei•ws ^Ex'te^rnalM.o,rmpmhology.—LTi.vi.ng&animal,t,h.in,
paaddpserndeewscbrioigbeeosgtrhaepmhiocraplhodaltoagyroegfatrhdeisnegttarxopaiacnadl lfulcoenngtatwe^antde 4™th ovienrlleynigntgh.oBpoaqduyecowlhoirtetrapnisg--
American Flabellinidae. mentondfaltwothirdsoftherhinophoresand
oraltentacles.Eachtranslucentceraswithcentral
reddigestivediverticulum,terminatinginalarge,
Descriptions
opaque white cnidosac. Rhinophores smooth.
Flabellinabertschi Goslinerand Kuzirian, 1990 Oral tentacles approximately as long as rhino-
[171]
172 PROCEEDINGSOFTHECALIFORNIAACADEMYOFSCIENCES,Vol. 48, No. 9
phores. Cerata ofuniform diameter throughout ata arranged in five paired clusters. Anterior 2-
most oftheir length. Many cerata damaged and 4 postcardiac groups containing complete arch
partially regenerating. Cerata arranged in dis- ofcerata while posterior 1-3 groups with only
creteclusters.Anteriormostclusteroneitherside single row ofcerata. Gonopore situated on right
containing four rows of cerata with 1-3 cerata side ofbody, ventral to third and fourth ceratal
per row. Posterior digestive branch with four rows. Pleuroprocticanussituatedatposteriorend
clusters ofcerata, each with 1-3 rows ofcerata. of interhepatic space. Foot with elongate, ten-
Each row with 1-3 cerata. Genital apertures lo- tacular corners.
cated on right side of body, ventral to second Discussion.—Theexternal morphologyofthe
and third ceratal rows. Pleuroproctic anus situ- present materialagreesentirelywiththeoriginal
atedwithininterhepaticspace,belownotum,im- description.
mediatelyanteriortoanteriormostcerataofright
posterior digestive branch. Anterior border of Flabellina telja Marcus and Marcus, 1967
foot with elongate corners. (Figs. 1, 2)
Discussion.—There isnodoubtthatthepres- FlabellinateljaMarcusandMarcus, 1967:223(inpart), Figs.
entspecimenisFlabellinabertschi. Itagreeswith 76-82.
the original description of this species, but is FlabellinastohleriBertschandFerreira, 1974:347,Figs.3, 16-
slightly smaller than any ofthe type specimens. 21. syn. nov.
Nevertheless, it is sexually maturewith well-de- Material Examined.—CASIZ 019047, five paratypes of
veloped gonads and reproductive organs visible Flabellinastohleri, 2-3 m depth, lowerend ofbay, Bahiade
through the transparent body wall. SanFrancisquito,SofPuntaSanFrancisquito,BajaCalifornia,
Mexico, 24 August 1971, H. Bertsch, E. Coan, R. Holiday.
CASIZ088160,onespecimen, 16 mdepth,NsideIslaMon-
Flabellina marcusorum Gosliner and Kuzirian, tuosa.GulfofChiriqui,PacificcoastofPanama,15April1993,
1990 T. Gosliner. CASIZ 088154, one specimen, 13 m depth, N
sideofIslaMontuosa,GulfofChiriqui, PacificcoastofPan-
Material.—CASIZ 088212, two specimens, 15 m depth, ama, 15 April 1993,T. Gosliner.CASIZ079164, 3 mdepth,
IslasSecas,GulfofChiriqui,PacificcoastofPanama,21April WendGardnerBay,IslaEspanola,IslasGalapagos,7Septem-
1993, T. M. Gosliner. CASIZ 088180, one specimen, 22 m ber 1991, T. Gosliner. CASIZ 071303, one specimen, 5 m
depth,SpanishWaters,Curacao, April 1990,JeffHamann. depth, Spoint,AcademyBay, IslaSocorro, IslasRevillagige-
dos, Mexico.
Distribution.—Flabellina marcusorum has
beencollectedfromtheAtlanticcoastofCuracao Distribution.—Flabellinateljawasoriginally
(present study), Venezuela (Jeff Hamann, pers. describedfrom BahiaCholla, PuertoPenasco,in
comm.), and Brazil (Marcus and Marcus 1961). the northern extreme ofthe GulfofCalifornia
Flabellina marcusorum is known from the Pa- andsubsequentlyrecordedfromBahiaSanFran-
cific coast ofMexico, from Isla Cedros south to cisquito (Bertsch and Ferreira 1974, as F. stoh-
Sayulita,Nayarit,andfromSanAgustin,Sonora, leri)onthewesternsideoftheGulfofCalifornia.
south to Los Islotes, north ofLa Paz, within the Specimens examined in this study extend the
GulfofCalifornia(GoslinerandKuzirian 1990). range ofthis species to the Islas Revillagigedos,
Thepresentspecimensextendtherangesouthto GulfofChiriqui, and the Galapagos Islands.
—
Panama in the Pacific and north to Venezuela External Morphology. Livinganimals 3-
mm
and Curacao in the Caribbean. 9 in length. General body colortranslucent
External Morphology.—Living animals 4 white, sometimeswithbluishpurplecast. Rhino-
and 10 mm in length. General body colortrans- phores translucent whitish with opaque white
lucent rose pink. Oral tentacles, foot corners, apex. Basal portion oforal tentacles same color
rhinophoresrosebasally with purple pigment in as body. Distal third of oral tentacles opaque
theirmiddleandopaquewhitepresentonapical white. Entire surface of body, including basal
portions. Oral tentacles with wide band ofpur- portionofcerata,frostedwithdenseopaquewhite
ple. Ceratawithonlynarrowpurplering. Rhino- spots. Through translucent surface of cerata,
phores elongate, bearing numerous elongate pa- salmon to red pigment of digestive diverticula
pillae on theirposterior face. Oral tentaclesthin visible. Apex of cerata opaque white. Rhino-
and elongate, longer than rhinophores. Cerata phores bulbous and densely perfoliate in their
arrangedindistinctgroups. Anterior, precardiac distal halves. Oral tentacles thin and elongate.
cluster with 2-4 rows ofcerata on either side of Cerataarearranged indistinctpedunculateclus-
animalwith 2-4cerataperrow. Postcardiaccer- ters with 3-4 ceratal rows in the anteriormost
GOSLINER:TROPICALAMERICAN FLABELLINIDAE 173
Figure 1. Flabellinastohleri. Scanningelectronmicrographsofparatype. (A) Halfrowofradularteeth, scale 60nm. (B)
Ventralviewofrachidiantooth, scale= 30nm.
cluster. Posterior peduncles with only a single distance, oviduct joining short stalked recepta-
row ofcerata. Gonopore situated on right side culum seminis. Oviduct entering female gland
ofbody,ventraltosecondandthirdceratalrows mass in region ofalbumenglandandcontinuing
ofprecardiaccluster. Pleuroprocticanussituated towardsgenitalapertures. Largebursacopulatrix
withininterhepaticspace.Anteriorcornersoffoot
tentacular.
—
Internal Morphology. Buccal mass short
andmuscular. Jawsthinandcoriaceous. Several
rows ofdenticles present along masticatory bor-
der. Radular formula 17-31 x 1.1.1., in two
specimens examined (CASIZ 079164, and the
paratype ofF. stohleri, CASIZ 019047, respec-
tively).Rachidianteeth(Fig. 1A,B)simplyarched
with 6-11 triangular denticles on either side of
elongatecentralcusp. Centralcuspslightlywider
than adjacent denticles and depressed ventrally
fromtheirlevel.Lateralteeth(Fig. 1A)triangular
inshapewithbroadbaseextendingtowardsouter
edge. Laterals with single prominent, acutely
pointed apex. Inner margin oftooth with series
of0-12 irregular denticles.
Arrangement ofreproductive organs (Fig. 2)
essentiallythe same as describedbyMarcus and
Marcus (1967). Narrow preampullary duct wid-
ensintoampullaconsistingoftwoconvolutions. buFmiegnurgela2n.d;Falambel=lianmaputellljaa.;Rbeepr=odbuucrtsiavecospyusltaetmr,ix;alm=eal=-
Postampullary duct again narrowing and divid- membranegland;mu=mucousgland;pe=penis;pr=pros-
ing into oviduct and vas deferens. After short tate;rs = receptaculumseminis,scale= 1.0mm.
174 PROCEEDINGSOFTHECALIFORNIAACADEMYOFSCIENCES,Vol.48, No. 9
Figure3. Livinganimals.(A)Flabellinavansyoci,photographofholotype.(B)Flabellinahamanni,photographofspecimen
fromTurksandCaicosIslands.
adjacent to female gonopore. Bursa exiting via uniformdiameter.Vasdeferensenteringbulbous
moderately long duct. Female gland mass con- penis. Penisornamentedwith many small, glan-
sisting of membrane, albumen, and mucous dular papillae,
glands. Vas deferens moderately short and of Discussion.—There has been considerable
GOSLINER:TROPICALAMERICAN FLABELLINIDAE 175
B
Figure4. Flabellinavansyoci. (A)Ceras, scale= 1.0mm.(B)Lateralviewofbodyshowingceratalarrangement,a=anus;
ga=genitalaperture;r=rhinophore,scale= 2.0mm.(C)Ventralviewoffoot,scale= 2.0mm.(D)Reproductivesystem,am
= ampulla; fgm = femaleglandmass,pe = penis;rs= receptaculumseminis;v = vagina,scale=0.75mm.
confusionsurroundingthesystematicsofseveral andFlabellinastohleriarelikelysynonymouswith
speciesofFlabellinaoccurringintheeasternPa- each other. Bertsch and Ferreira (1974) stated
cific. Gosliner and Kuzirian (1990) noted that thatF. stohleridiffered fromF. teljain itscolor,
Marcusand Marcus's (1967) description ofFla- number ofradular rows, and shape and dentic-
bellina telja also included specimens that were ulation ofthe radular teeth. Both taxa were de-
subsequently described as F. bertschi Gosliner scribedashavingatranslucentbodydenselycov-
and Kuzirian, 1990. Goslinerand Kuzirian also ered by opaque white spots. The orange body
indicatedthatspecimensidentifiedasF. teljaby colordescribedbyBertschandFerreiraisdueto
FerreiraandBertsch(1972)andKerstitch(1989) thecolorofthevisceraratherthantheepidermis.
were F. marcusorum Gosliner and Kuzirian, ThebluishpurplecolordescribedbyMarcusand
1990. GoslinerandWillan(1991), incomparing Marcusisevidentinsomespecimensandabsent
Flabellina rubropurpurata Gosliner and Willan, inothers,evenfromthesamelocality.Themor-
1991 toothertaxa,indicatedthatFlabellinatelja phology ofspecimens ofFlabellina telja studied
176 PROCEEDINGSOFTHECALIFORNIAACADEMYOFSCIENCES,Vol. 48, No. 9
Flabellina vansyoci sp. nov.
(Figs. 3A,4-6)
Flabellinasp. 2 Behrens, 1991:86, Fig. 173.
—
Type Material. Holotype, California Academy of Sci-
ences,SanFrancisco,CASIZ088274,SendofIslaMagdalena,
MagdalenaBay,PacificcoastBajaCalifornia, 16January, 1984,
R. Van Syoc. Paratypes, CASIZ088081, twospecimens,one
dissected, 10mdepth,anchorageIslasLadrones,GulfofChi-
riqui, PacificcoastofPanama, 13 April 1993,T. Gosliner.
Distribution.—Flabellina vansyoci has been
collectedfromMagdalenaBayonthePacificcoast
ofBajaCaliforniaandfrom IslasLadronesalong
the Pacific coast ofPanama.
Etymology.—Flabellina vansyoci (pro-
nounced "van psyche") is named for my friend
andcolleague,RobertVanSyoc,oftheCalifornia
Academy of Sciences, who collected the first
specimen ofthis species.
—
External Morphology. Living animals
mm
(Fig. 3A) 15-30 in length. General body
color translucent rose pink to purple. Cerata
translucentpurplishwithreddigestiveglandvis-
iblethrough surfaceofcerata. Distalhalfofeach
ceras bearing congested spots ofopaque white.
Whitish cnidosac visible at apex ofceras. Rhin-
ophores and oral tentacles same color as body.
mm
Figure 5. Flabellina vansyoci. Scanning electron micro- Rhinophores elongate (approximately 4 in
graphofmasticatoryborderofparatype,scale = 43 nm. length) and conical. Rhinophores each bearing
indistinct rugose markings along their entire
length. Oraltentaclesthinandelongate, tapering
herefrom IslasRevillagigedos, Panama, andthe to a rounded apex. Oral tentacles slightly longer
Galapagos is entirely consistent with morphol- than rhinophores. Cerata (Fig. 4A) cylindrical
ogydescribedbyMarcusandMarcus(1967)and and elongate with rounded apex and elongate
the holotype ofF. telja (USNM 678418) exam- cnidosac. Thick core of digestive gland filling
ined at the Smithsonian. The Galapagos speci- most of diameter of ceras. Cerata arranged in
menexaminedhere hadfewerradularrowsthan distinct clusters ofrows (Fig. 4B). Only anterior
the type material, but was ofsmaller body size. cluster slightly elevated from notum. Anterior,
The morphologyofaparatype ofF. stohleriwas precardiac cluster with five rows of cerata on
examined in this study (Fig. 1A, B). Its number either side of animal with 1-5 cerata per row.
of radular rows (31) is more similar to the 30 Postcardiac cerata arranged in 6-8 clusters per
rows described for F. telja than the 14 rows de- side ofbody. Generally, each group containing
scribed forF. stohleri. Also, the numberofden- 1-3 tightlypackedrowswith 1-5 cerataperrow.
ticles and shape ofteeth are intermediate to the Onlyposteriormost 1-2 clustersconsistofsingle
materialexaminedbyMarcusandMarcus(1967) ceratal row. Gonopore situated on right side of
and Bertsch and Ferreira (1974). The reproduc- body, ventral to second and third ceratal rows
tive system ofF. stohleri was not originally de- ofprecardiaccluster. Pleuroprocticanussituated
scribedby Bertsch and Ferreira. Examinationof nearmiddleofinterhepaticspace,anteriortofirst
two paratypes ofF. stohleri (present study) in- postcardiac ceratal row. Nephroproct immedi-
dicatesthatthesespecimenshaveapapillatepen- atelyanterodorsaltoanus. Foot(Fig. 4C)grooved
ialpapillaasdescribedforF.telja.Thereremains anteriorlyandpossessingelongate,tentacularfoot
nobasisforseparationofthetaxa,andF. stohleri corners. Foot tapering gradually to narrow tail
is here regarded as ajunior synonym ofF. telja. posteriorly.
GOSLINER:TROPICALAMERICANFLABELLINIDAE 177
Figure 6. Flabellina vansyoci. Scanningelectron micrographs ofparatype. (A) Dorsal view ofradula, scale = 60 ^m. (B)
Rachidianteeth, scale = 20i*m. (C)Lateral viewofradula, scale= 25 Mm. (D)Lateraltooth, scale= 15^m.
Internal Morphology.—Buccal mass short Jawsovoidwithelongatemasticatorymargin(Fig.
and muscular. Large digitate oral gland extend- 5). Margin bearing five or six rows ofdenticles
ing from anteroventral portion ofeither side of with about 34 denticles on outer row. Radular
buccalmassandcontinuingintowidenedportion formula 36 x 1.1.1., in one paratype examined.
ofnotum in region of precardiac cerata. Large Rachidianteeth(Figs. 6A-C), simplyarchedwith
pairofchitinousjawssituatedwithinbuccalmass. 5-6 triangular denticles on either side of more
178 PROCEEDINGSOFTHECALIFORNIAACADEMYOFSCIENCES,Vol. 48, No. 9
Figure7. Flabellinapedata. Scanningelectronmicrographs.(A)Dorsalviewofrachidianteeth, scale 20Mm.(B)Lateral
viewofradula, scale = 20^m.
elongate central cusp. Central cusp approxi- and penial apertures. Vas deferens narrow, but
matelysamewidthasadjacentdenticles, butde- appearing to contain prostatic cells throughout
pressed ventrally from theirlevel (Fig. 6C). Lat- most of its length. Vas deferens entering short
eral teeth (Fig. 6A, D) triangular with a broad penial sac, wider than the vas deferens. Penial
baseextendingtowardsouteredge. Singleprom- papilla simple and unarmed.
—
inent, acutely pointed apex present on lateral. Discussion. Flabellina vansyoci differs
Inner margin oftooth with series of 15-20 tri- markedly from other eastern Pacific species of
angular denticles. Laterals on right side of the Flabellina.Itistheonlyspecieswithrugoserhin-
radula ofparatype entirely lack denticles (likely ophores.TheothereasternPacificspeciesallhave
aberration rather than characteristic feature of annulate,perfoliate,orsmoothrhinophores.Fla-
species). Pairofelongate salivaryglandsthatex- bellina vansyoci has a reddish-purple body sim-
tendposteriorlyondorsolateral surface ofstom- ilar to that ofF. marcusorum. It can be readily
achpresentatposteriorlimitofbuccalmass,near distinguished by the opaque white spots and a
itsjunction with esophagus. lack of purple bands on the cerata. Flabellina
Arrangement of reproductive organs essen- telja may also have a purplish body color and
tiallytriaulic(Fig.4D).Narrowpreampullaryduct opaquewhite spots. However, its spotsare scat-
widening into simply saccate ampulla. Ampulla teredoverthebody surface, not restrictedto the
narrowingintoelongatehermaphroditicductand cerata.Thisspeciesalsohasperfoliateratherthan
dividinginto vas deferens and oviduct. Oviduct rugose rhinophores.
curving andjoining pair ofequally sized recep- Flabellina vansyoci is related to other flabel-
tacula seminorum and vaginal duct. More dis- linidswith a bilobed receptaculum seminis. It is
tally, vaginal ductenteringfemaleglandmass in most similar to F. pedata (Montagu, 1815) and
region ofalbumen gland. Vaginal duct continu- the sympatric F. bertschi, both of which have
ingtowardsgenitalapertures.Nidamentalglands smoothratherthanrugoserhinophores.Allother
comprisingbulkofreproductivesystem.Mucous species with a bilobed receptaculum have papil-
glandlargestportionbyfar. Albumenand mem- late rhinophores. In orderto make comparisons
brane glands smaller. Nidamental glands emp- with F. vansyoci, six specimens of F. pedata
tyingvia their own gonopore, ventral to vaginal (CASIZ 072597, one dissected, intertidal pools,
GOSLINER:TROPICALAMERICAN FLABELLINIDAE 179
Figure8. Flabellinahamanni. (A)Ceras, scale = 0.5 mm. (B)Lateral viewshowingarrangementofcerata,a =anus;ga =
genitalaperture;r= rhinophore,scale = 2.0 mm.(C)Ventralviewofanteriorendoffoot, scale = 2.0mm. (D) Reproductive
system, al = albumengland; am = ampulla; be = bursacopulatrix; me = membranegland; mu = mucousgland; pe = penis;
pr=prostate;rs= receptaculumseminis;v = vagina, scale = 1.0mm.
PontaDelgadaHarbor, IlhaSao Miguel,Acores, ofF. vansyoci,thelateralteetharemarkedlydif-
Portugal, 17 July 1988, T. Gosliner) were ex- ferent. In F. pedata, the lateral teeth bear 6-9
amined.ThespecimensofF.pedatastudiedhere coarsedenticles,whileF. vansyocihas 15-20fine
representthe firstrecord ofthis species fromthe denticles. The reproductive systems are very
Acores. The ground color ofF. pedata is a deep similar,butitappearsthattheprostateisthicker
purplevioletwhileF. vansyociisreddishpurple. in F. pedata than in F. vansyoci (Schmekel and
Flabellinapedata lacks the opaque spots on the Portmann, 1982; present study).
basal portion ofthe cerata that are present in F.
pedata. As noted above, F. pedata has smooth Flabellina hamanni sp. nov.
rhinophores,whileinF. vansyocitheyarerugose. (Figs. 3B, 8-10)
In F. pedata, the postcardiac ceratal groups are Type Material.—Holotype, California Academy of Sci-
arrangedinarcheswhileinF. vansyocithegroups ences, San Francisco, CASIZ 088275, 3 m depth, Freeport,
contain 1-3 rows ofcerata. While the rachidan GrandBahamaIsland,CaribbeanSea,June 1986,J.Hamann.
teethofF.pedata(Fig. 7A,B)aresimilartothose Paratypes, CASIZ088276, onespecimen dissected, same lo-
180 PROCEEDINGSOFTHECALIFORNIAACADEMYOFSCIENCES,Vol. 48, No. 9
Figure 9. Flabellina hamanni. Scanningelectron micrographs. (A)Jaw, scale 430 Mm. (B) Masticatory margin, scale
43tim.
cality and date as holotype. Paratype, CASIZ 072269, one portion ofcerata. Posteriorendoffoot also pur-
specimen,livingonhydroids,20mdepth,Thea'sWreck,Ba- ple with opaque yellowish line or spot on its
hamas,CaribbeanSea,April 1986,B.Rose.Paratypes,CASIZ posterodorsalend. Digestiveglandwithincerata
088882,threespecimensandeggmass,reefSofWestCaicos,
Turksand Caicos Islands, 20 m depth, December 1983, Jeff charcoal gray and visible through inner trans-
Hamann. parent portion of ceratal bases. Rhinophores
mm
elongate (approximately 4 in length) and
Distribution.—Flabellina hamannihasbeen conical. Eachrhinophorebearingapproximately
collected from theTurksandCaicosIslandsand 100 elongate papillae on its posterior face. Pa-
mm
Grand Bahama Island (Jeff Hamann, pers. pillae approximately 0.5 in length and ar-
comm.) ranged in indistinct rows, covering middle two-
Etymology.—This species is named for my thirds of rhinophores. Apex of rhinophores
friendandcolleague,JeffHamann,whofirstcol- conical and devoid of papillae. Oral tentacles
lected this species. thin and elongate, tapering to acutely pointed
—
External Morphology. Living animals apex.
(Fig. 3B)25-32mminlength. Generalbodycol- Cerata narrow at base and widening to just
ortranslucentrosepink. Oraltentacles, footcor- below their apices. Cerata terminating in acute
ners, and rhinophores rose basally with narrow apex(Fig. 8A).Thincoreofdigestiveglandfilling
band ofpurple pigment and opaque yellowish- muchofdiameterofeachceras. Cerataarranged
white present on apical portion. Oral tentacles indistinctgroups(Fig. 8B). Onlyanteriorcluster
with wide area ofopaque yellow covering two- slightly elevated from notum. Anterior, precar-
thirds of their length. Cerata and rhinophores diac cluster containing four rows of cerata on
bearing only proportionately narrower band of either side ofanimal, with 3-4 cerata per row.
yellow. Surfaceofceratacoveredwithdustingof Postcardiaccerataarrangedinsixgroupsofarch-
opaque white. Dusting absent from inner basal es per side of body. Generally, each arch con-