NEW RECORDS IN PSEUDOGNAPHALIUM (ASTERACEAE: GNAPHALIEAE) FROM NEBRASKA AND CALIFORNIA

2004·1.7 MB·English

by  Steven B Rolfsmeier

NEWRECORDSINPSEUDOGNAPHALIUM(ASTERACEAE:
GNAPHALIEAE)EROMNEBRASKAANDCALIEORNIA
Steven B.Rolfsmeier GuyL.Nesom
HighPlainsHerbanum BotanicalResearchInstituteofTexas
ChadronStateCollege 509PecanStreet
ChadronNebrasl<a69337,U.S.A. FortWorth,Texas76102-4060,U.S.A.
ABSTRACT
PseudognaphaliumjciliscenseisfirstreportedfromNebraska,documentedhereIromtwelvecoun-
ties;anearlierreportofthisspeciesfromCaliforniawasbasedoncollectionsolaformofP.
stramineum.PseudognaphaliummseumisfirstdocumentedasoccurrmgintheU.S.A.,whereithas
beencollectedmsouthernCalifornia.
RESUMEN
PseudognaphaliumjalisccnscsecitaporprnneravezdcNebraska,sedocumentaaquidedoce
condados;unacitapreviadeestaespeciesdeCaliforniaestababasadaencoleccionesdcunatormade
P.stramineum.PseudognaphaliumroseumsedocumentaporprimeravezdeU.S.A.,dondehasido
colectadoenelSurdeCalilornia.
InconnectionwithtaxonomicstudiesofGnaphalieae,preparationoftreat-
mentsfortheFloraofNorthAmericavolumesofAsteraceae,andfloristicstud^
iesinNebraska, we report newdistribution recordsfortwospeciesof
PseudognaphaliumKirp.
PseMdogMaphaIiunijflJisccn.seinNebraska
PseudognaphaliumjaUscense(Greenm.)Anderb.wasrecentlyreportedforthe
U.S.A.,basedoncollectionsfromArizona,NewMexico,Texas,andColorado
(Nesom2001,includingdetailsoftypificationanddescription).Thecurrent
reportdocumentsitsoccurrenceintwelvecountiesofNebraska.Itwasfirst
collectedinthestatein1968bySteveStephens(CusterCo.)butwasnotrecol-
lecteduntil1991byHerbKarcher(HayesCo.).Ithassincebeenrecollectedin
bothcountiesinadditiontonumerousothersitesinthesouthwestandtheNe-
braskaSandhillsinthenorthcentralpartofthestate.Therecencyofthecol-
lectionsandthecommonroadsidehabitatsuggestrecentintroduction,butother
collectionsarefromapparentlynaturalhabitats,andthenativityofP.jaUscense
inNebraskaisnotclear
DistributionofthespeciesisshowninFigure1fornorthwesternMexico
andtheU.S.A.,basedontheNebraskacitationsbelowandthoseinNesom(2001).
AdditionalrecordsforNewMexicoalsoarecitedbelowandsuggestthatthe
speciesmaybeevenmorecommoninthatstate,wherehabitatsdonotsuggest
thatthespeciesisarecentcolonizer.IfthedistributionofPseudognaphalium
SIDA21(2):1205-1208.2004
1206 BRIT.ORG/SIDA21(2)
Fig.1.DistributionofPseudognaphaliumjaliscenseintheU.S.A.andnorthwesternMexico.Thespeciesrangesfurther
southwardintoJalisco,Nayarit,andAguascalientes,andlocalitiesinsouth-centralNuevoLeonarenotshown.Therecords
forMexicoarefromcollectionsstudiedbyNesomfromvariousherbaria.
jalisccnscisnativeintheU.S.A.,itapparentlyshouldbeexpectedincountiesof
northeasternColoradoandnorthwesternKansas.
With furtherperspective,collectionsreportedasPseudognaphaUum
jalisccnscfromMendocino,Marin,andDelNortecounties,California(Nesom
2001),provetobetheformofPseudognaphaliumstramineum(Kunth)A.Anderb.
sometimesidentifiedasGnaphahumstramineumvarconfertifoliumGreene.
Thesepfantshavelargerheadswithmorenumerous,moreopaquephyllariesand
oftenmorefloretsthantypicalPstramineuni,whichisabundantinCalifornia.
ROLFSMEIERANDNESOM,NEWRECORDSOFPSEUDOGNAPHAUUM 1207
U.S.A.NEBRASKA.BlaineCo.:T24NR25WSec26,1Aug1998,Morris899(CSCN).BrownCo.:Long
Pine,12miS,6niiW,prairieminterdunalSandhillsvalley,2Aug2000,Roljsmeier15780&R.
Steinauer(NEB).CherryCo.:2.5miNofHookerCo.line,sandyroadditchesalongNebr.97,13Sep
1995,Roljsmeier12053(BRIT,CSCN,KANU,NEB);SolMerrittReservoir,sandyroadside,13Sep1995,
Roljsmeier12056(BRIT,CSCN,KANU,NEB);nearturnofltoMarshRanchonNebr.97,ca,3.5miNol
HookerCo,line,sandyroadsideditchandadjacentpasture,22Aug1996,Roljsmeier12911(BRIT,CSCN,
KANU,NEB).CusterCo.:Anselmo,4miNW,marginolprairieslough,moist,sandysoil,several
plants,28Aug1968,Stephens28200(KANU);Oconto,0.25miN,3.5miE,ravinebottominmixed-
grassprairie,22Jul1999.Roljsmeier14903&iParker(BRITNEB).FrontierCo.:T7NR30WSec18,
hSeoafviCluyrt-igsr,azTe7dNsaRn2d8prWaiSrieecn1e1,arprHaaiyrieespCaos.thunree,,629SeSpep199129,92R,oJlojfsimnesi.en.r(1C11S1C7N()B.RIGTr,aKnAtNCUo.,:N0E.1B,miCSWCN)o;l
HookerCo.line,scatteredatbaseofrailroadembankment,22Aug1996,Roljsmeier12913(BRIT,CSCN,
KANU).HayesCo.:13.5miSofjunctionofNebr.Hwys23&25,roadbank,21Jul1991,Karcher820
(CSCN,KANU);HayesCenterWildlileManagementArea,uplandprairieoneastsideoflake,6Sep
1992,Roljsmeier11109(BRITKANU,NEB,CSCN).HookerCo.:6.25miEofGrantCo.line,baseof
railroadembankment,22Aug1996,Roljsmeier12912(BRITCSCN,KANU,NEB),KeithCo.:Paxton,
3.3miW,sandyroadditchbetweenU.S.30andrailroad,25Sep1992,Roljsmeier11282(CSCN,KANU,
NEB).LincolnCo.:severalmilesnorthofHayesCo.line,sandyprairiealongNebr25,6Sep1992,
1R1(1)9/9/s(mBeRIiTe,rJClSJC3N3(,CKSCANN,UN,EBN)E;BD)i;ckTe1n2sN,9Rm3i4EW,sSaencd2y6,rosaadnbdaynbkaanlkonogfNdietbcrh,23W,1s1iSdeepol19h92w,yRo2l5,/s7meAzuegr
1997,G.Steinauer253(NEB);Wallace,6miN,sandyslopealonghighway23,20Sep1998;Davis51
(CSCN).LoupCo.:Taylor,7.5miESE,sandyroadbank,28Jul1999,Ro//smeierJ4974&Pflr)2er(CSCN,
NEB).PerkinsCo.:Elsie,0.25miE,sandyprairieinroadsideditch,11Sep1992,Roljsmeier11193(BRIT,
CSCN,KANU).NewMexico.GrantCo.:Ft.Bayard,openwoodland,6300ft,18Sep1967,Gierisch
3284(UNM).HidalgoCo.:Cloverdale,grassyhillside,22Aug1955,Castetter9959(UNM);GrayRanch,
HoweCamp,cienega,20Aug1993,iveys.n.(UNM).LincolnCo.:Sacramento(White)Mts.,Gavilan
RidgeattrailtoBlueLake,burnedareathatwasoriginallymostlyponderosapine,7300ft,30Sep
2001,Worthington30728(UNM).
PseudognaphaliumroseuminCalifornia
Pseudognaphalium roseum(Kunth)A.Anderb.iswidespread inCentral
AmericaandMexico,fromPanamanorthwardthroughChiapastoChihuahua,
Veracruz,NuevoLeon,Tamauhpas,andsoutheasternCoahuila.Itoccursmostly
above1000melevationinavarietyofhabitats.TheclosestcollectionsofP.
roseumsouthwardfromCaliforniaarefromSinaloaandsouthernChihuahua,
butthespeciesisabundantonlyineasternandsouthernMexico.Itisrecog-
nizedbyitspersistentlytomentosestemsandleaves,theleavessubclaspingand
non-decurrent,weaklybicoloredandsessile-glandularbeneaththetomentum,
oftenthickstems,largecapitulawithnumerous,opaque-whiteorsometimes
pinkphyllaries,numerousflorets,andsmooth-surfacedcypselae.Ithasbeen
confusedwiththesmaller-headedPcanescens(DC.)A.Anderb.,butplantsofP
roseumwithrelativelyfewbisexualfloretscanbedistinguishedfromP.
canescensbytheirsubclaspingleavescommonlywithcloselywavymargins,
broaderandmorenumerousphyllaries,andsmooth-surfacedcypselae.Akey
withdiagnosticcharactersforthesetaxaisprovidedbelow.CollectionsofP
roseumcitedfromSanMateoCo.andSantaBarbaraCo.maybeatypicalintheir
slightlysmallercapitula.
1208 BRIT.ORG/SIDA21(2)
U.S.A.CALIFORNIA.OrangeCo.:CatalinaIsland,16Jun1Q20,Ms/uts.n.(SMU).SanDiegoCo.:Bor-
derFieldStatePark,southwesternmostacreofcontinentalU.5,A.,16N4ar1989,Turner15925(TEX);
TiaJuana,24Apr1913,tlaslwood2918(GH),SanMaieoCo.:PebbleBeach,Pescadero,29June1927,
Wolf563(TEX).SantaBarbaraCo.:PurissimaCanyon,2bMarI92'5,Munz-^7.37(TFX).VenturaCo.:
NorthSignalStreet,Ojai,8Oct1946.Pollard203M(SN-llI).
a. Mid-caulineleavesmostly3-7cmlong,(3-)6-15(-20)mmwide,subclasping,the
marginsoftencloselysinuate-wavy;pistillateflorets45~90(-l10);bisexualflorets
(5-)6-12(-18);cypselasurfacessmooth(nonpapillate) Pseudognaphaliumroseum
a. Mid-caulmeleavesmostly1-4cmlong,2-8(-l5)mmwide,notclasping,themar-
ginsusuallystraight;pistillateflorets(16-)24-44;bisexualflorets(1-)2-5(-6);cyp5ela
surfacesdenselyandminutelypapillate Pseudognaphaliumcanescens
Pseudognaphalium roseum(Kunth)A.Anderb.,Opera Bot.104:148.1991.
CnaphalinmroscinnKunihmH.B.K.,NovGen.&rSp.4|roliol:63.1818;4[quartol;81.1820.TYPE:
K4PX1CO,Gi)ANA|U.<vro:HumhohltandHoiiphiitds.i\A\\oiOT\VV:.Pliche!).
Taprootcdannualtoshort-livedperennialherbs,thestems5-20dmtall,per-
sistentlywoolly-tomentose.Leavesoblong-lanceolatetooblanceolate,themid-
cauline3-7cmlong,(3-)6-]5(-20)mmwide,subclaspingtoclaspingbutnot
basallyampliate,notdccurrent,themarginsoftencloselysinuate-wavy,per-
sistentlywoolly-tomentoseorsometiinestardilyglabrescentabove,withshort-
stipitatctosessileglandsbeneaththetomcntum. Involucres4-4.5mmhigh;
phyllariesovate,in5-6graduateseries,usuallyopaque-whitebutsometimes
pink.Pistillateflorets45-90(-110).Bisexualflorets(5-)6-12(-18),thecorollas
oftenpinkonthelowerhalf.Cypselaeshallowlylongitudinallyridged,smooth.
ACKNOWLF.nGMF.NTS
WcgratefullyacknowledgeloansfromUNM,thestaffatTEXforhelpsinvisits
there,staffatNHBandKANUlorconfirmingcollectiondatalorspecimensde-
positedthere,andCalebMorseforareviewolthemanuscript.
rhferfnc:e
NtsoM,G.L2001.NewrecordsinPseudognophalium(Gnaphalieae:Asteraceae)forthe
UnitedStates.Sida19:1185-1190.