New Perspectives on Horned Dinosaurs: The Royal Tyrrell Museum Ceratopsian Symposium

657 Pages·2010·15.137 MB·English

by  Michael J. Ryan, Brenda J. Chinnery-Allgeier, David A. Eberth

paleontology
Easily distinguished by “New Perspectives on Horned Dinosaurs records a ryan New PersPective s oN “Triceratops and its kin may hail from the dim
landmark event and makes clear that our understanding - Horned and distant past, but this new volume brings
the horns and frills on their chinnery
of this group is undergoing truly explosive growth. to give them fully into the light of today. An all-star
allgeier
just one measure, the number of ceratopsids discussed at dinosaurs and comprehensive list of authors not only
skulls, ceratopsids were one
this meeting represented a doubling of species compared eberth effectively puts horned dinosaurs in the context
of the most successful of to a comprehensive review of this clade published just of their own time and place, but also brings
three years earlier. the remarkable abundance of newly The Royal Tyrrell Museum them alive as living, breathing biological
all dinosaurs. This volume discovered forms was augmented by presentation of Ceratopsian Symposium organisms. New Perspectives on Horned
rigorous studies of stratigraphy, phylogeny, ontogeny, Dinosaurs is able proof of the vitality of
presents a broad range of biomechanics, taphonomy, paleogeography, and paleo- New modern dinosaur science, bringing to bear
environment. these results, including descriptions of PersPectives twenty-first-century ideas and approaches
cutting-edge research on
ten new taxa, are captured in this volume, which oN to ask—and answer—questions that once
the functional biology, will be a must-own for dinosaur paleontologists Horned would have been thought to be out of reach.”
and enthusiasts alike.”
—Larry Witmer, Ohio University
dinosaurs
behavior, systematics,
—Scott Sampson, University of Utah
paleoecology, and
Michael J. Ryan is Vice-Chair Curator and
“From Archaeoceratops to Zuniiceratops, from Ceratopsids, or horned dinosaurs, are a group
Head of Vertebrate Paleontology at the
paleogeography of the Alaska to Mexico, and from sediments to functional of large-bodied, quadruped herbivores that
Cleveland Museum of Natural History.
morphology, this book covers much of present-day lived roughly 65–70 million years ago. Part
horned dinosaurs, and research on ceratopsians. these horned dinosaurs of a larger group of dinosaurs that include
are rendered as living, behaving, and evolving stegosaurs, ankylosaurs, ornithopods, and
includes descriptions
organisms throughout the thirty-six chapters of pachychephalosaurs, the better known
this book. i encourage everyone interested in members of the ceratopsids include centro-
of newly identified
how a myriad of incredible fossils can inform saurs, chasmosaurs, and triceratopsians.
species. about life of the past to read it.” All are easily distinguished by the horns and
frills on their skulls; in fact, ceratopsids have
—David Weishampel, co-editor of
among the largest, most elaborate skulls
The Dinosauria and co-author of
Dinosaurs: A Concise Natural History found in vertebrates. they were one of the
most successful of all dinosaurs and their
“this book captures an explosion of new and remains are well known from many locations
exciting research on one of the most fascinating in the United States, Canada, and Mexico.
groups of dinosaurs. it will be a landmark in they died out at the end of the Cretaceous
Brenda J. Chinnery-Allgeier is Lecturer the study of ceratopsians.” along with the rest of the dinosaurs.
in the School of Biological Sciences at
—David C. Evans, University of Toronto
The University of Texas at Austin.
the outgrowth of the Royal tyrrell Museum’s
Ceratopsian Symposium, this volume presents
a broad range of cutting-edge research on
the functional biology and behavior, system-
atics, paleoecology, and paleogeography of
the horned dinosaurs, including descriptions
of newly identified species. there is also a
history of collecting these dinosaurs, plus a
supplementary Cd-ROM containing a history
of ceratopsian discoveries in Canada and
EditEd by michael j. ryan,
a list of the specimens recovered to date.
brenda j. chinnery-allgeier,
indiana and david a. eberth
INDIANA
University Press
David A. Eberth is a senior research
Bloomington & Indianapolis
scientist at the Royal Tyrrell Museum in www.iupress.indiana.edu Life of the Past
Drumheller, Alberta, Canada. 1-800-842-6796 James O. Farlow, editor supplementary cd-rom included Cover illustration: ©2009 donna Sloan
NEW PERSPECTIVES ON
HORNED
DINOSAURS
LIFE OF THE PAST
James O. Farlow, editor
NEW PERSPECTIVES ON
HORNED
DINOSAURS
The Royal Tyrrell Museum
Ceratopsian Symposium
EDITED BY
MICHAEL J. RYAN
BRENDA J. CHINNERY-ALLGEIER
DAVID A. EBERTH
Indiana University Press PATRICIA E. RALRICK
Bloomington and Indianapolis EDITORIAL ASSISTANT
This book is a publication of
Indiana University Press
601 North Morton Street
Bloomington, IN 47404-3797 USA
www.iupress.indiana.edu
Telephone orders 800-842-6796
Fax orders 812-855-7931
Orders by e-mail [email protected]
∫ 2010 by Indiana University Press
All rights reserved
No part of this book may be reproduced or utilized in any
form or by any means, electronic or mechanical, including
photocopying and recording, or by any information storage
and retrieval system, without permission in writing from
the publisher. The Association of American University
Presses’ Resolution on Permissions constitutes the only
exception to this prohibition.
! The paper used in this publication meets the minimum
requirements of the American National Standard for
Information Sciences—Permanence of Paper for Printed
Library Materials, ANSI Z39.48-1992.
Manufactured in the United States of America
Library of Congress Cataloging-in-Publication Data
Royal Tyrrell Museum Ceratopsian Symposium (2007 :
Drumheller, Alta.)
New perspectives on horned dinosaurs : the Royal Tyrrell
Museum Ceratopsian Symposium / edited by Michael J. Ryan,
Brenda J. Chinnery-Allgeier, and David A. Eberth ; Patricia E.
Ralrick, editorial assistant.
p. cm.
Includes bibliographical references and index.
ISBN 978-0-253-35358-0 (cloth : alk. paper)
1. Ceratopsidae—Congresses. I. Ryan, Michael J., [date]-
II. Chinnery-Allgeier, Brenda J. III. Eberth, David A. IV. Title.
QE862.O65R695 2010
567.915—dc22
2009019913
1 2 3 4 5 15 14 13 12 11 10
this volume is dedicated to the late
Halska Osmólska,
whose work on basal neoceratopsians
set a standard for excellence,
and to
Wann Langston, Jr.,
for his work on horned dinosaurs that
now spans more than five decades and
continues to inspire new research.

What seest thou else in the dark backward
and abysm of time?
Shakespeare, The Tempest, act I, scene 2

CONTENTS
Preface xiii
Acknowledgments xv
List of Contributors xvii
List of Reviewers xxi
PART ONE § OVERVIEW
1. Forty Years of Ceratophilia / PETER DODSON 3
PART TWO § SYSTEMATICS AND NEW CERATOPSIANS
2. Taxonomy, Cranial Morphology, and Relationships of Parrot-Beaked Dinosaurs
(Ceratopsia: Psittacosaurus) / PAUL C. SERENO 21
3. A New Species of Archaeoceratops (Dinosauria: Neoceratopsia) from the Early Cretaceous of the
Mazongshan Area, Northwestern China / HAI-LU YOU, KYO TANOUE, AND PETER DODSON 59
4. A Redescription of the Montanoceratops cerorhynchus Holotype with a Review of
Referred Material / PETER J. MAKOVICKY 68
5. First Basal Neoceratopsian from the Oldman Formation (Belly River Group), Southern
Alberta / TETSUTO MIYASHITA, PHILIP J. CURRIE, AND BRENDA J. CHINNERY-ALLGEIER 83
6. Zuniceratops christopheri: The North American Ceratopsid Sister Taxon Reconstructed on the
Basis of New Data / DOUGLAS G. WOLFE, JAMES I. KIRKLAND, DAVID SMITH, KAREN POOLE,
BRENDA J. CHINNERY-ALLGEIER, AND ANDREW MCDONALD 91
7. Horned Dinosaurs (Ornithischia: Ceratopsidae) from the Upper Cretaceous (Campanian)
Cerro del Pueblo Formation, Coahuila, Mexico / MARK A. LOEWEN, SCOTT D. SAMPSON,
ERIC K. LUND, ANDREW A. FARKE, MARTHA C. AGUILLÓN-MARTÍNEZ, CLAUDIO A. DE LEON,
RUBÉN A. RODRÍGUEZ-DE LA ROSA, MICHAEL A. GETTY, AND DAVID A. EBERTH 99
8. New Basal Centrosaurine Ceratopsian Skulls from the Wahweap Formation (Middle Campanian),
Grand Staircase–Escalante National Monument, Southern Utah / JAMES I. KIRKLAND AND
DONALD D. DEBLIEUX 117
9. A New Pachyrhinosaurus-Like Ceratopsid from the Upper Dinosaur Park Formation
(Late Campanian) of Southern Alberta, Canada / MICHAEL J. RYAN, DAVID A. EBERTH,
DONALD B. BRINKMAN, PHILIP J. CURRIE, AND DARREN H. TANKE 141
10. New Material of ‘‘Styracosaurus’’ ovatus from the Two Medicine Formation of Montana /
ANDREW T. MCDONALD AND JOHN R. HORNER 156
11. A New Chasmosaurine (Ceratopsidae, Dinosauria) from the Upper Cretaceous Ojo
Alamo Formation (Naashoibito Member), San Juan Basin, New Mexico / ROBERT M. SULLIVAN
AND SPENCER G. LUCAS 169