NEW OR LITTLE-KNOWN TIPULIDAE FROM NEW GUINEA

2016·3.4 MB·

by  Alexander, Charles P

» - NEW OR LITTLE-KNOWN T/PULIDAE FROM NEW GUINEA

5s (Fipatidae, Dipteray’ +

Port 1°
Cnanins P. Audxanppe
= {University ot Blarssetanette, Amhesst)
With 8 wasttiemes

2, Abthis the I are deacribirg four new species belonging to the fenus
rlistie Wassenuon, @1 taken by Dr M.A, Tikeaac fn the vieinlly
ci Uke Eumono Oilf'ekls, ia nortiovestern New Guices. ‘The lypes of
these species will be dezasite: in the wathor's colleetion, At the seme-time «
i amefign sing tha mele hyxopygium of five further members of the
conus. that had heen deseribed euelier Zy rigself but had nev
*iNlastrat ies were tuker by Mise L. E, Camueican in Waigen,
2 Guinea and is Papua, the typos boing proserved in
‘he Brilish Museum of Natural History (Landon). Tam very deeply in.
debted to Dr Livrrsck for tre privilege n” determining: the ahove mal
Wale ‘git other more exieusive series trom New Guinea that wil, be
digenssed in a later paper.

Gynoplistia (Paralimnophili) euryphaea Anexannet (ix. L)
198. Gyunplistia Pecetin:nophiba) eyrypionet Aursasves; Ann. Mas. Nu:
et) tai = 75s,

The type, a2, was Fram. Mount ‘Tafa, Papue, altitude 8590 feet,
taken in February 1984, by Camesman. | *

Genoplistia (Gynoplistia) dixantha ALexanpur (Zig. 2)
Giyaeptistia Uiynopliotat diconthe. ALEXSSEM; Ana. Mag. Nat. Hist, (1D)

was from Mafula, Papea, altitede 4000 fect, taker,
TERENAS,

Gyauplistia (Gyawplisiia) Eavizowa ALwxannine (ip. 8)
Ingr. Gyneplistta (Gyouplivtia) jlevizcna Auexssper; -Ana, Mag. Nab, ili, (111
19d,

+ Tho tynsfa of, was from Kokoda, Papua, elnitude 1200 feet, taken
ig Scprember-Ostober 1988, by Cibusman,
Tere, Vor. 21, 1951, Paee A. ae

-

syuoplistia (G:

iplistia) Klamanaensis sp. n. (ig. 4) :

General coloration ‘black, the pleura pele on pyopleure ani pt
pleurites legs entirely black; wingd.erayish eudhyeline. hexylly pusternet
with dark beotm, iheluding cells C and Se, with putlerior exle:

abdomen black, scemeats two to six with yellow bases; male hypopy:zinm

with a single apical lobe o2 vasislyes dististyle singlé, eapandled ‘at tips
sdeagts relatively slender, Une. uplcal third with ecversl small darker,

projuet ions. *

Male. —Tongtie bout 7 5 wiew 05 nus.

Rostrum’ very reilaced, victually dush with he: fom, testacsuus ,
brown; palpi pale brows. Anizanae wits scape and pedine uranie:
flagellum broken, relatively large, heed
eaneverse,

Peonvital ecuium bly (lun polished heswnish black. Meso
notyyn almast anifnemly blackened, the suxfsee suzopaque by & ypara
pruinosity. Leura black, ste sersely wi yale, Ukiswiolog invo:- *
ving she greplews sud the prsroplenrita. Halters veith stem pale, the
onter end, with the knob, Slick. Legs blaek, the eoxae epansely pri
Wings graye subliyalize, heavily patterned with dark brown, ine-ading
the pveuretar fie’d and cells (aad Se, wita posterior exteusiecss Uhere-
from, beoaulest at cord and wie tip, the former not invelving t]e venuer of
coll 136 Af; other Carkenings incinde the aveulus, oxigin ef Bs, outer enc
af eall (st A and ve-eus an inolntes. dack apot in exe era! celts veins
Jark Leon, Velus beyond cori with abundant macrolsichia, Venestun
Sq encing about osposite the Zork of Ry o54 1. Seo ab ils Hip; Pe lone,
strdugly arewated at origin: Roy, gy shirts aneciee branch of Rs ying
close Lo Ry 5, the outer end of the ce” narrow: velit £2, faint tr aul
oheolete; cell ©, wreseni, acboanal to ite petiole; eo: 9("R, recisingular,
with va-cw at midleasth; yeln gal A strongly simacgs.

Alitumen conspicuzusly bivdlared, le vogments blesk, the eecond to
sixch, inclusive, with conspicuzns yellow basal rings, these brvulesl ox
the accond sopment waere ue e-halt is inchded, Serem-"ng: progress
ively narrower em the onter segments; seventh und auceweding segments
no.uding she hypopygiurs, uniformly blaekaned, Mule hyporygtim ("ig, 2
with Oto lergite, 9f, “ange, transveree, the catdal musgin:traneate. Busi
style, 6, at apex produced into a single lobe or blade, iis broadest beLoue
midlength, fhenes narrawel to ie Lip, Ue ever manger with umisualiy
long setae, the longest: mors than Lwiee tae diameter of she blade eppuaily

4 above Cars brown: eyé

ns to *
5, arégin af Its, along cord and over the wing tip; cell M, presanl; va «
CLP, Araxniimi: New Piputites feom Mow Guinea. a
‘tho yoint of insertion, Dististyle, d, single, dongute, nearly twice as long
as the Jobe of the basistyle, the stem straight, he, outer fourth mors
oxparded, near the tip suddenly narrowed inte a more slender lobule,
the apex ol which is flactened o* slightly expanded. Aedvawus, «, Telati-
vely clendex, straight, on apical third with several small weal darkened
projections that ave seaxcely spinous; tip oblnse, pale and membranons,
Hsloigpe, ,oKbamonn Oilfields, Northwest New Guinea, August 18
IS (LIBPLINCK).
There are a few generally. sind

uw Papuan species, all Ciffering in

the structure of the male hyponygiim, Thavdetivtyle and acdeagus of
the present fy are distinctive cmong such ole

Gynoplistia (Gynoplistia) lulibaestlis sp. n,

General co.vrution of mesonotal 9

avsentum and seutam reddish
rn, possibly pruinose in fresh specimens; mediotergive veddish brown.
with a blick central triangle; pleura brownish black, based re ly
with silvery oa the propleura end pisroplearize: antennae (female) 16-
sopiiented, with eighl, branched segments, the longest branches
three times the segments; Zemora yellow, the tips black; w
blackened, Ue prearcular field move yellowed : a broad more whitish band
before cord, about ha'f ag wide aa the basal black band; eells Cand Se
uniformly divk brows; macrorichia of veins beyond cové very sparse
bilomen black, segments two to four, with the extreme base of five, dall
searye: genital shield oranse,

Female, — Length boul 11 mins wing 9.5 mm.

Rostrum and pal:i yellow, Antermac (femsle) L6-sexmented, the.
formla being 2-- 2+ 6+; seapo and pedicel ‘ercnindc® of
organ chie‘ly dark brow», the incisnres of she four or Live proximal
Fagellar scements yellow, very brdsdly so oti the first one, on the others
with only ‘the apleal sie: darkened; all ouler segments and. branches
uniformly darkened; branch of frst flagellen acement nearly twice the
segment, of the second about {wo and one-helf times; longest branch (abaut
flagellar semment. four or five) about. threx times the segment. Head
the extreme central part of anterior vertex 9

gs chiefly

Fronctum brownizh black, Mecouolal praestntum snd seutum chiefly
reédlish hrowa, the surface discolored, probably prainose in fresh material;
Jum dark brown; meriotergite obscure reddish brown, with a black
central triangle, Uke point bebiii Bleue brownish black, the propleurs

akly darks
4 ‘Tram, Ver. 24, 2051, Pam 4

and pievopieutite pale, heavily silvery white pruinose, Hlalteres infuscated,
Knob brownish black, basco! stoma narrowly yellow. Legs with corse
Heht gray pruinose; trockanters dark brown; femors yellow, the tine
ck, including about the anter sixth; ectreme bases.of femora nasrowly
and vaguely infuscated; tibiae dark brown, tke tips narrowly darker;
tarsi black; claws microscopically serrulate on Inwer face of ahout tke
Stoximal quarter. Wings ehietly blackened; proarcular field moive pellow.
ed; @ bread more whitish band before the cord, extending from vein &
virtually Lo the posterior border; centers of outer radial vella very narrow
ly whitened <c form. an ouler incomplete crossliand: calls C and Se
unitoymiy dark browns the dark bssal crossband is approximately twice
As wide us the majax pale area; veine brown, a trilfAlighter in the pale
dend, peer I xellawed, Mactotrichia even spares,
eccurring only om distal section of i. Venssion: Aap, , short, one
about one-third the subobsolete basal section af Ke,> cell, present, sab
Caaal to ils petiole; mew al nea midlength of cell 158 3,

Abdomen with the fire: sagrient luck: segmen-s twe ww Sour, wit
extreme baso of five, orange yellow. the surface 6
black; genital shield orange, cerci ovange-yellow

Holotype, ¢, Klariono Oilficlds, Northwest New Guinea, August 18
24, 19¢8 (Limerick), :

Tn its unusually heavy wing prltern, the present fly sugges
plistia (Canoplistia) melanckotica WALKER end @. (C) saigrithoron Atm
ANDRE, esnotially the latter. ‘his hay dhe enteunae Tisemented, with
ten branched segments, and w'ta the enti The blunt
serrulations un the claws, xs descrshed, ane fou ‘ge in other. spcvies
of the subgenus but usually to s greater degres than here

21; succeeding segments

wplisiia (Gynoplistia) Heftinel
General eclorasion 07 head and thorax orange yeliow, virtually un-
patterued; antennae (male) 16-segmented, with bwalve branched segments 5
femora orange yellow; wings light yellow on basal half, including the
veins, the veins of outer halt darkened, (ae wing tip narrowly infuseated ;
cell Wi Tncking; akdomen dul orange, the outer four segmenia Duk:
male hypopygium with two spinous points ut epex of basintylo; lististylo
single; phalloscre cansisting of twa sets of blackered roda,
Male. — Length about 11.5 mm; wing &.5 mr
Veraale.— Length shout 14 mm: wing 10 mm,
Rostriim of male yellowed at base darkened oulwiard
the dorsal surface uniformly darker

jana sp... (fig, 5).

ly. in feinale,
ed, provided with conapicumis »
CLP. Anpanuine: Now Piputtine from New Gace 5

sotact palpi Tight reddish brown, Antenmae (male) 16-segmiented, the
formula boing 242-1042; eeape and pedicel veddish yellow, the
succeeding two or taree segments wita the stem yellow, the branches
black; intermediate soprients black, the extreme hase of stera obscure
yallow, outer branched segments uniformly black, the outer two simple
ones mediumn brown; branches of flagellar scymenis one and two abort
four times the soxments longest branches (about. flagellar segmo
and seven) oe seven timed the segments; last’ branch subequal
in ‘engltr.lo the segment, T: female, the terminal semmenis broken bat
there are at least seven branched segments, the lorzest some four times
the scumont; lesal favelier sestcents with the ineisures broadly yellow,
the invermediate part of stern and the branches black, Head reddish town,
the vertex in femate more derkencil, almost blic.

Thorax polished oranve yellow, in female darker but giving the
appearanee of being Sod discolored from within. Halteres crange

yollow. Logs with the coxdd, prochunters and femora orange yellow, Ee
tips of the Inst not ar sex Jeenedl in malo, mors blackened in feraeles
tiniac aind basitarsi obscure yel.ow, the cuter tarsal segments black. Wings
wilh aoout the bagal half light yellow, inelading the veins, the cord and
the veing beyond brown, the wing tip narrowly pale brown; a vague
darkening over cord in ranle, bouiek xd moze couspiencus in Lene.
A complete series of trichia on distal section of vein I,; cu ouler madial
veirs, insiuding a few scablered Leichia on dislal seclion of ven ¥
Incking on the branches behind this. Venation: R, pale, about one-bul
as long as Ry 3 Riyy— a little shorter thar rem; cell Mf, lacking;
ariesie a sneer une-thi Teugils of cell fst My,

Abdomen of raale dull orange, the sixth and all succeeding segmeus,
inchidieg the hypepyeiam, blaek, the sixth and seventh segments with
purplish reflexions, Ie the female. thie basal live segments orense, Ue
romrindor of abdlomen polished blu-blsck, the valves of Lae ovipositor
orsnge. Ms opprium Gig, 5) with the ninth tergite, wt, trameste
across tip, Basistyle, b, at apex produced into two strong blackened: rods
or spines, the longest directed mesad, its tip acute or rn:eroseopieally
notched; shorter spine direstod cwuCad. Dististyle, d, single, abont twice
as long ay the shorter spine of the basistyle, narrowed to the Lip, the
lower margin microscopically tnbercnlate, Phallosome, », consisting of
various blackened rods and sp!nes (of whieh ore-half aro shown in figure) ;
Intorl rods beariug a strong spur near daze, with furlaer smaller spines
near its hase; inner rods drond-based, bent st 2 right angle into a very
shrenst straighs black spine,
woe bia, VoL 21, 1851, Paes 4 ad
. GP, AnaxannrR: Mew Tiplielze from Now Gainoe,

Holotype, *, Kiamono Oilficlds, Northwest New Guinea, Auguat 18 <
271943 CAEPTINCK). 7
Tam most pleased lo name this very insexesting species for the
ciilectar, Dr'B. A. LIReTINGR, to yhn T amt sreally Mndebiee for cooper
sion in Uke scady of the New Guinea Tiphifidae, The Lis pears s, marked
Superficial resemblance to Cyeontisti« (Gynopiistin) veckynae AUEXNME
Vrhich paswoll Mf,0f she wings present and differs conspieuensly in Use
oture of the male kypopyelum. The female heve associated with the
cpe uate a3 eTlotype differs i sTight regards but is elfeved Lo be con-
spec’. Pas, .

4, Geoplistix (Gymmlietid) viaihans eps. (fig: 6) .
Afesonotum brawnisa black, the plenra congoloruus, strived trans
vorsoly with pale: anternae (iaa’s) LG-zogruented, with cleyen Teng-branc!
ed segments; fenor yell, ihe tins conspicuously blackened: wings,
Wwhiigch suohyaline, wh a somewhat restrictes brown. bende palles
cell My pveserct: fizel ehiom’ne] sepment black, sezments two und thrge +
chiefly orange, the itera! borders more blackened: outer so othifal
sdioents blackened; wale hypopygiun with the basistyic produced at per
into n single setiforous inhe: phuilesome consiatings cf a pair of blackened
spines thas eubtend the centva! aedeag

‘Male, Length about 13 mun; wing 20 rem.

_Rantram cbscure avons yelon Zita seo segments ot pate 20g
the Suter (wo whitened. Antonnas (male) 16-seynrnted, the formals beinge
24: 2-9 +8; woape and pediccl orunge yellow ; sucewesting segments with

ve Incisures yellow, the intermediate mart of stern gnd the branrhes darks
Seown; ouler fone og five segments uniformy dak brown; branch plae
first tlagellar sexmont nearly Laver tines the wesment; Iomyosi« bramel
Gabont flagellar geamentt Live ox ix times bac segment
anion of the eleventh Zlageller segment shorl, less thar the segment.
(ead akovetorauge gr tzont and surrounding the basse of Ube ancenmae
fic quai on the octipat, the yostevior vertex wnd the orbits broelly in-
fuseated

Tronolign duib brownish black, the posterior angles of the geatelluso
and the preceemitee o2seure yellow. Mesoniotnm bevwnish lucky, the surface"
gorssuly piuinose, the peaceeatain withows distinst patfera, Pleura dull
Diack, erausversely sLriped with pale, the latter incladiag. a narvoss yellosy
band on the plenra extending Lo the bac of the fore coxa and & broad
sivvery white 6and extending: from the wing ract Lo above Lbe*mid coxe
Halteres brownish back, tho base of slom yellow. Legs with the coxa
® Tervina, Yoo. 21, 1

1, Pas 7
brown or 'yedtlish brown; trochanters piccous brown; femora yellow, tne
fips conspicuously blackened, with shout the outer fifth or sixth included ;
femoral bases less evidently darkeneil, ‘most extensive on the posteriye
legs: tihise brown or browzish vellow, ihe tips darker; larstblack. Wings
whitish subhyaline, the prearcular field yellowed: cells C andl Se darkened :
a somewhat restricted bown banded pattern, the most conspicuous aren
rer the cord, exterdeil narrowly to tho wing margin oyer mecusanyl distal
seclioa of vein Cu, virtually covering cell tt 32, wing tip narrowly
darkenec, the inner eile extending back alnost to tho fork of M«y ¢
basad of cord with pos.tively small isolated spots ab arenlus, origin of
ie, amdhins coll 3st A ab near lnvo-thinds the “eng-h of the vein3 veins het
drowh, darker brown in the patterned areas. Mucrotriehid of outer vein «
unssually sphrsc, even on the distal sertion of #, belug rclativey tow
aed scattered, lacking on the bagel pertions of the veine, Venation
ending jush beyond Che fork at Re, Se; near ‘its tip: section of BY
cbligtze, unusvally long, abuul Unree times 2. 3 inmer end of coli

Fel Mi, arewated, ly as col! Hy: coll A, subequal ,

m leagth to Its petfole; mecn ul near one-third the leagth of ‘ell st 3.+
vein gad A more gently sinuons thax: is commen in the gents,

First abdominal segment black; seymonts two and three ehietly
orange, the Iuteral bosders more blackened; fuurlh segment orange an
bout the prox hyvbbyaium,

Mak. Biale 1 imatgin of the ninth *
termite, 96, convexly rounded, Apex of basisty‘e, b, produced into a sinate
seievotized and provided with unusually Ione
setae, Dististyle, d, aingle, relauively brond, especially ut near mnidlenzth
“uisnie> nurrowed to the obtase Lip, which beans fur or five bleckened

leeth; Thner margin of outer half with microseopie thbercles: surface

be

wilh sestiered microscopic: pale puretures. Pha‘losome, », consisting of
# pei of blickened spines and the central acdeagus, the Igtler with a,

uuadvete dilation al near midlength .
Holotype, 7, Klamod Oilfields, Northwost 2
24, 1843 (Lrsruivor)
Quite distinc trom other regional sy ings Gynoplintia
“Gumoplistin) Memonsensiv sp... in the coloration and, osperially, the
structure of the male hypopyyium, sperticrlariy the prelosome,

» August 1g

Gynoplistia (Gynovlistia) walgeuensis Auexanper (fig. 7).
1s, Gem
(ity a5: 79

ivtia (ymopliabel waigevtusis Avexasomn; Ann. Mog, Nat. Tis, |
760, t
COP, Auiaxoan: New Tipntiive From New Guinea 8

‘The lype, a cy was from Camp Nok, Waigen, altitude 2500, feet:
taken ia April 1988, by. CUBESMAN. The detailed account of the collegtor’s,
expedition to Waigen and. Japen in 1988-1929 has been most. entertainly”
ceseribed in hér revent book, "Six-loggé.i snakes in New Guinea,” 281 Pp,
_ plates, toxt-figapes; 1909; Garona G. Hannay and Company, Les.

Gynoplaatia (Gynoplistia) xenlhocera AUEXANDER (fiz, 8).

1048, Gunephetia’ (Cyaoptistin) eunthocera ALwsanneny Ann. MagiNet, 16
ye QM) 99: 951-106,

The lype, ae’ was from Sabron, Cyeluuy (Cyclops) Mountains, North,

‘rn Datwh New Guineu, altitude $00 feet, taken in May 1936, hy CHUaSMAK:

,