New or little known Chrysomelidae (Coleoptera) from Nepal, Bhutan and the northern territories of Indian subcontinent

2004·2.4 MB·

Bull.KitakyushuMus.Nat.Hist.Hum. Hist.,Ser.A, 2:47-63, March31, 2004
New or little known Chrysomelidae (Coleoptera) from Nepal,
Bhutan and the northern territories of Indian subcontinent
Shinsaku Kimoto
7-97, Mikunigaoka, Ogori City, Fukuoka, 838-0103, Japan
(ReceivedOctober2,2003;acceptedFebruary 19,2004)
ABSTRACT—Sixteen species of the Chrysomelidae from Nepal, Bhutan, N. India and Pakistan are
described asnewspecies, viz. Cryptocephalus eberti, Pseudostonopaflaviventris, Doryxena minor and
Paridea quadrimaculata from Nepal, Monolepta tatemizo from Nepal andBhutan, Arthrotus kalimpon-
ganus, Calomicrus nigriceps, Cneorane wittmeri, Japonitata ochracea, Luperus variabilis, Monolepta
atosuji, Monolepta gantokensis, Monolepta labiatella, Sastroides nigriceps andChalaenosoma megala-
yanum from N. India, and Taumacera pakistanica from Pakistan. Number of taxonomic new treatment
areproposed andnumberofspeciesarenewlyrecorded fromtheareaorfromparticularcountries.
timesaslongawide; firstsegmentlong,robust,somewhat
PREFACE
club-shaped;secondshort,nearlyhalfaslongasfirst; third
In this paper, new or little known species of Chry slender,nearly 1 1/2 times as long as second ; fourth sub-
somelidae from Nepal, Bhutan and the northern territories equal tothird inlength and shape ; fifth nearly 1 1/2 times
ofIndiansubcontinent are treated. Sixteen species are de aslongasfourth,andfifthtotenthsubequal toeachother in
scribed as new species. Number of taxonomic new treat length but stouter than preceding ; eleventh slightly longer
ment are proposed and a number of species are newly re thantenthanditsapexpointed. Pronotum stronglyconvex,
cordedfromtheareaorfrom particular countries. transverse, nearly 1 2/3 times as wide as long, distinctly
The following abbreviations are used herein for the narrowedanteriorly,lateralmarginnearlystraight,andante
typedepository museums. rioranglerounded, posteriorangleslightlyproduced; dor
salsurfaceverysparselyand indistinctlypunctate. Scutel-
BASEL NaturhistorischesMuseum,Basel. lumsubtriangular,rounded at apex, surface nearly impunc-
MUNCHEN Zoologishe Staatsammlung, Miinchen. tate. Elytron elongate, subparallel-sidedwith apexrounded;
surface strongly convex, distinctly punctate-striate, with in
Subfamily Cryptocephalinae tersticesveryfinelyandremotelypunctate.
Length :5.2-5.6mm.
Cryptocephaluseberti sp.nov.Fig.la Distribution: Nepal.
Subcylindrical. Generallyreddishtoyellowishbrown; Holotype : Sun Khosi Tal, 2150 m, Nepal, 2. v. 1962,
pronotum witha pairofdistinctblackishmarkings,andba G.Ebert(MUNCHEN)
salmarginnarrowlyblackish; elytronwithfairlylargehu Paratype:RiKholaTal,2300-2800m,Nepal,1ex.,13.
meralandpostmedian markings,together with lateral mar v. 1962,G. Ebert.
ginblackish;antennapitchy black with two or three basal This new species resembles Cryptocephalus oppositus
segments brownish ; ventral surfaces with third to fifth ab Jacoby,1908,from N.W.Himalayas (Fig. lb), but separable
dominal segments generally pitchy black ; legs with tibiae fromthisspecies inhaving theelytron with thehumeral and
andtarsigenerallypitchyblack. thepostmedian markings, thelegswithtibiaeandtarsigen
Headwithvertexsparselyimpressedby minutepunc erallyblackish and the ventral surfaces with the third to the
tures, andwithadistinctlongitudinalimpressionat middle, fifth segmentsblackish.
starting fromanteriormargin; fronto-clypeus sparselyim
pressedby fine punctures. Antenna slender, nearly 2/3 as
longas body length ; in preapical segments nearly 2 1/2
48 Shinsaku Kimoto
a
Fig. 1.a-b.dorsal surfaces, c-d,prothorax. e-f,male genitalia, a.Cryptocephalus eberti sp.nov.: b.Cryptocephalus oppositus (Jacoby): c. Pseudo-
stonopaflaviventris sp.nov.;d.Dercetina viridicyanea Kimoto:e.Luperus variabilis sp.nov.;f.Japonitata ochracea sp.nov.
New or little known Chrysomelidae 49
SubfamilyEumolpinae tate. Elytronslightlywideratbasethanthorax,subcylindri-
cal ; surface strongly punctate-striate, interstices of punc
Aulexis nepalensis MedvedevetSprecher, 1997 turessmooth, shining ;humerus costiform, and with aridge
Coleoptera,Schwanfeld, 1:230(Nepal,India). below humerusrunningposteriorly.
tibialis : Kimoto & Takizawa, 1973, Kontyu, Tokyo, Length :4.7-4.8 mm.
41 : 176(Nepal). Distribution :Nepal.
Distribution :Nepal, India Holotype : Rapti Tal, 300 m, Nepal, 27. iii. 1962, G.
Material examined. Mene Bhanjang, Darjeeling, India, Ebert(MUNCHEN).
1ex.,9.vii. 1975,W.Wittmer. Lopchu, Darjeeling, India, 1 Paratype : 1ex.,same data asthe holotype.
ex., 9. v. 1975,W. Wittmer. This new species resembles Pseudostonopa picea Ja
The record by Kimoto& Takizawa (1973) from Nepal coby, 1908, from Sikkim, but differs from this species, in
should becorrectedasthis species. havingthe ventral surface reddish brown, and sides of pro
Aulexistibialis Jacoby, 1889 notumstronglyangulateandwidestat 1/3 frombasalmar
Ann. Mus. Civ. Genova,27 : 167 (Burma). gin.
Distribution :India,Burma Xanthomaoblonga Takizawa et Basu, 1987
Material examined. Daugiri, 450 m, Garo Hills, Kontyu,Tokyo, 55 :275 (Darjeeling).
Megalaya,India,1ex.,19.v. 1976,W.Wittmer&C.Baroni This species isvariable on the coloration of dorsal sur
Urbani. faces : 1) Generally yellowish brown with lateral and su-
The body length ofthis species is shorter than Aulexis tural margins, together with scutellum, pitchy black,2)gen
nepalensis Medvedevet Sprecher. erally yellowishbrown with lateral and sutural margins, to
Colaspoides bengalensis Duvivier, 1892 getherwithscutellumandmiddle ofprontoumpitchyblack,
Ann.Soc.EnLBelg.,36 :418 (Sikkim). 3)pronotumyellowishbrownandelytrondarkbrownwith
subrugosa Jacoby, 1908, Fauna India Col, Chrys., 1 : the apical and basi-sutural areas yellowish brown, 4) en
515 (Himalaya). New Synonym. tirely yellowishbrown.
Distribution: India, Nepal. Distribution :India, Nepal.
InthecollectionofNaturalHistory Museum, London, a Material examined. Landrung, 1645 m, Nepal, 3 exs.,
cotype of Colaspoides bengalensis Duvivierand the holo- 16. v. 1983, Y. Hama. Late, Nepal, 2438 m, 1 ex., 12. v.
typeof ColaspoidessubrugosaJacobyare preserved. As a 1983,Y. Hama.
result of study of these type series, it is known that these
two species are same, and subrugosa Jacoby becomes a SubfamilyGalerucinae
synonymofbengalensis.
Pseudostonopafulviventris sp.nov. Fig. lc Agetoceraflaviventris Jacoby,1879Fig.4a
Subquadrate-oval ; generally reddish brown, elytron Proc. Zool. Soc. London, 1879 :788 (India).
yellowishbrown ; antenna pitchy brown with four or five Distribution :Nepal, Bhutan, India, Burma
basalsegments paler. Material examined. Chasilakha, Darjee Khandu D.,
Head with fronto-clypeus and vertex distinctly and Bhutan, 1ex., vi. 1979,1 ex., 1978.
rather closely punctate and their surfaces smooth, shining. This species is here recorded for the first time from
Antennafiliform, nearly 2/5 as long as length of body ; in Bhutan.
preapicalsegmentsnearlythreetimesaslongaswide;first Apophyliaassamensis (Jacoby, 1891)
segment robust, somewhat club-shaped ; second slender, Entomolog., 24(Suppl.): 34 (Assam) (Malaxia).
nearly 11/4 times as long as first; third nearly halfas long pallipes :Kimoto,1977, EnLBasiliensia, 2 : 353 (Bhu
as second ; fourth nearly 1 1/2 times as long as third and tan).
fourth to tenth subequal to each other in length but stouter Distribution :Nepal, Bhutan, India, Burma
thanpreceding; eleventh 1 1/4 times as long as tenth and Materialexamined. Beguna, DorjeeKhandu, Bhutan, 1
itsapexpointed. Pronotumtransverse,nearly 11/3 times ex.,23. viii. 1976.
as wideas long,sides strongly angulate and widest at 1/3 Kimoto(1977) recorded Apophylia pallipes from Bhu
from basal margin, and strongly narrowed anteriorly and tan. However, this record should be corrected asApophylia
posteriorly ; anterior margin distinctly rounded anteriorly assamensis. This species is here recorded for the first time
and posterior margin less strongly so posteriorly ; surface from Bhutan.
closely impressed by strong punctures and their interstices Apophyliaflavovirens (Fairmaire, 1878)
smooth and shining. Scutellum subtriangular with apex Ann. Soc. Ent. France, sen 5, 8 : 139 (S. China) (Mal
rounded ;surfaceslightlyconvex, smooth, shining, impunc- axia).
50 Shinsaku Kimoto
a
Fig.2. a-b,prothorax. c-f,malegenitalia, a,Doryxena minor sp. nov.; b,Doryxena grossaHope ;c,Monolepta gantokensis sp.nov. d, Taumacera
pakistanicasp.nov.;e,Stenoluperusbhutanensis Kimoto;f,Chalaenosomamegalayanum sp.nov.
New or little known Chrysomelidae 51
Distribution : Bhutan, Thailand, Laos, Vietnam, China, Elytron broader than prothorax at base, slightly widened
Hainan,Taiwan. posteriorly and rounded at apex ; subbasal area not dis
Material examined. Ungar - Lhuntsi, 2500-1800 m, tinctly raised, surface smooth, shining, distinctlyand closely
Bhutan, 1ex.,6. vi. 1976,L. Caminada. punctate.
This species is here recorded for the first time from Length :5.5-6.2mm.
Bhutan. Distribution: N. India
Apophylia maculata Kimoto, 1977 Fig. 4b Holotype : Punchung, 450 m, Kalimpong, Darjeeling,
EnL Basiliensia,2 :352(Bhutan). India,5. iv. 1977(BASEL).
Distribution :Nepal, Bhutan, India Paratype : 1ex., samedataas the holotype.
Material examined. Mane Bhanjang, Sukhia Pokri, This new species resembles the brownish type ofAr
Darjeeling,India, 1ex.,9. vi. 1975, W.Wittmer. throtus phaseoli Laboissiere, 1932, from Sumatra, Java,
This species is here recorded for the first time from N. Tonkin,but differsfrom this speciesin havingthe pronotum
India with apairofdistinctdepressionslaterally.
Arthrotusnigripennis (Jacoby, 1889) Arthrotusphaseoli Laboissiere, 1932 Fig. 3a-d
Ann. Mus. Civ. Genova,27 :224(Burma)(Taphinella). Mem. Mus. Hist. Nat Belg., 4 : 174, fig. (Sumatra Java
Distribution : Nepal, Burma Thailand, Laos, Vietnam, Tonkin).
China antennalis Laboissiere, 1932, Ann. Ass. Nat Levallois-
Material examined. Darugiri, Garo Hills, 450 m, Perret, 20 : 134(Tonkin). New Synonym
Megalaya, India, 1ex., 19.v. 1976,W.Wittmer&C. Baroni Distribution : Nepal, Bhutan, Thailand, Laos, Vietnam,
Urbani. Sumatra,Java
This species is here recorded for the first time from N. Calomicrusannapurnae Medvedevet Sprecher, 1999
India Ent Basiliensia, 21 :367 (Nepal).
Arthrotuscyeaneus (Chujo,1966) Distribution: Nepal, Pakistan.
Mem. Fac. Educ. KagawaUniv., II, 145 :8 (Nepal) (An- Material examined. Kawai, Khagan V., 1450-1880 m,
astena). Pakistan, 1ex., 14.vi. 1977,W. Wittmer& M. Brancucci.
hauseri Kimoto, 1967, Esakia, Kyushu Univ., 6 : 73 This material slightly differs from the nominate form,
(Simla). New Synonym. in having the dorsal surfaces entirely bluish black, instead
Distribution: Nepal, India ofcupreousblack.
Arthrotuskalimponganus sp.nov. This species is here recorded for the first time from
Body oblong-oval. Generallyochraceous, antennadark Pakistan.
brownwith four or five basalsegmentspaler. Calomicrusnigriceps sp.nov.
Head with vertex convex, smooth, shining, nearly im- Generally ochraceous ; head pitchy black with vertex
punctate, and with adistinctdepression in front ofmiddle ; mostlydarkbrown.
frontal tubercles distinctly raised, transverse, contiguous, Headwith vertexnot very convex, smooth,shining, im-
distinctly delimited posteriorly. Antenna slender, slightly punctate ; inter-antennal space longitudinally raised ; inter-
shorterthan body length, inpreapical segments nearly 3 1/2 antennal space narrow, slightly narrowerthan width ofeye ;
times as long as wide ; first segment long, robust, club- frontal tubercles well developed, contiguous, subtriangular,
shaped;secondshort,slightly longer than wide,and nearly and their anterior angles inserted between antennal inser
halfas long as first; third short, nearly as long as second ; tions. Antenna slender, slightly shorter than body length ;
fourth long,nearly 11/2 times as long as the length ofsec in preapical segments nearly seven times as long as wide ;
ond and third combined ; fifth slightly shorter than fourth, first segment long, robust, somewhat club-shaped ; second
and fifth to tenth subequal to each other in length and short,nearly 1/3aslong asfirst; third slender,nearlytwice
shape ; eleventh slightly longer than tenth and its apex as long as second ; fourth nearly 1 1/2 times as long as
pointed. Pronotumtransverse,nearly 13/4timesaswideas third, and fourth to ninth subequal to each other in length
long ; anterior margin nearly straight, posterior margin and shape ; tenth slightly shorter than ninth ; eleventh
widely rounded posteriorly ; lateral margin slightly slightly longer than tenth and its apex pointed. Pronotum
rounded, widest almost at middle, slightly narrowed anteri transverse, convex from side to side, nearly 1 1/2 times as
orly and more strongly so posteriorly ; anterior and poste wide as long, widest almost at middle, and gradually nar
rior corners thickened, each with a setigerous pore ; dorsal rowed anteriorly and posteriorly ; anterior margin slightly
surface convex, smooth, shining, sparsely impressed by rounded anteriorly and posterior margin widely rounded
minutepunctures, and with apair ofdistinctdepressions lat posteriorly ; anterior and posterior comers thickened, and
erally. Scutellum subtriangular, surface smooth, shining. each with a setigerous pore ; dorsal surface sparsely im-
52 Shinsaku Kimoto
< *
Fig. 3.a-il.Arthrotusphaseoli LABOISSIERE.
pressed by minute punctures. Scutellum small, sublriangu- ex., 2. vi. 1976.W. Wittmer & C Baroni Urhani.
lar. surface smooth, nearly impunctate. Elytron hardly This species ishere recorded for the first time from Ne
broaderat base than prothorax, slightly widened posteriorly, pal.
and apex rounded ;dorsal surface closely impressed by dis
tinct punctures. Genus Charaea Baly
Length :5.0-6.0 mm.
Distribution : N. India. Charaea Baly, 1878, Cist. Ent.. 2 : 376 (type species :
Holotype : Umtygar, Chedrropunjee. Magalaya, Assam, Charaeaflaviventre Baly, 1879:monobasic).
India. 16. vi. 1976. W. Wittmer & C. Baroni Urbanl Malacosoma Chevrolat, 1845, Diet. Univ. d'Hist. Nat..
Paratype : 1ex.,same data as the Holotype. 6 : 5 (nee Huber, 1820) (type species : Chrysomela lusi-
This new species resembles Calomicrusflavus (Jacoby, tanica Linnaeus, 1769).
1892), from Burma, but differs from this species in having Exosoma Jacoby, 1903.Trans. Ent. Soc. London, 1903 :
the pronotum less transverse and nearly 1 1/2 times as wide 25 (new name for Malacosoma Chevrolat, 1845). New
as long, and the head partly blackish. Also, from Calomi Synonym.
cruspersimilis Kimoto, 1989,from Laos and Vietnam,dif In the type species, Charaea flaviventre Baly, 1878,
fers in having the head partly blackish. the prosternal process iselevated between the coxal cavities
and separating them. The type of this species is preserved
Cassenacollaris (Bai.y, 1879) Fig. 6a in the Natural History Museum, London. Thus, Exosoma
Cist. Ent..2:457 (Assam) (Euphyma). Jacoby becames asynonym ofCharaea Baly.
Distribution : Nepal. India. Burma. Thailand. Vietnam. Charaeabalyi (Medvedeet Sprecher. 1998)
China. Malaya, Sumatra, Java. Medvedev et Sprecher, 1998, Spixiana, 21 : 32 (Nepal :
Material examined. 5 km E. Manhari. 350 m. Nepal. 1 new name for Charaeaflaviventre Baly, 1878. nee Calomi-
NeworlittleknownChrysomelidae 53
cmsflaviventreMotschulsky, 1860)(Calomicrus). Dercetinabretinghami (Baly, 1879)
flaviventreBaly, 1878,Cist.Ent.,2 : 376 (neeCalomi Ann.Mag.Nat Hist, sen5,4:118(India) (Antipha).
crusflaviventre Motschulsky,1860)(Murree)(Charaea). pallidus Laboissiere, 1932, Ann. Ass. Nat Levallois-
balyi: Kimoto,2001,Bull. Kitakyushu Mus.Nat Hist, Perret, 20 : 133 (Tonkin) (Arthrotus). - Kimoto, 1989,
20:36(Nepal)(Charaea). Esakia,Kyushu Univ.,27:227(=bretinghami).
Distribution:India,Nepal. hainanaGressitt & Kimoto, 1963,Pac.Ins.Mon., IB :
Materialexamined. Phulchoki, 2600m,Nepal, 17exs., 707(Hainan). - Kimoto, 1989, Esakia, Kyushu Univ., 27 :
11-14.vi.1976,W.Wittmer&C.BaroniUrbani. Godavari, 227(=bretinghami).
1500-1700m,Nepal, 1ex.,21. v. 1977,W. Wittmer & M. bhutanensis Kimoto, 1977, EnL Basiliensia, 2 : 385
Brancucci. Pam,S.Gantok,980m,Sikkim,India,4 exs., 17. (Bhutan). New Synonym.
iv. 1977,Bakta Bahadur. Mussorie, 1300-2000m, U. P., Distribution : Nepal, Bhutan, India, Burma, Thailand,
India, 1ex.,27. vi. 1976, W. Wittmer. Cambodia,Laos,Vietnam.
Cneoranewittmerisp.nov. Dercetinaminiaticollis (Hope, 1831)
Generally yellowish to reddish brown ; scutellum InGray,Zool.Misc.: 29(Nepal)(Haltica).
pitchy black ; elytron bluish black; antenna pitchy black Distribution :Nepal, Bhutan, N.India
withtwoor threebasalsegments partlybrownish ; meso- Material examined. Pedong, 800-1200 m, Darjeeling,
and metathorax and abdomen bluish black ; legs pitchy India, 1ex., 14. viii. 1978, Bhakta Bahadur. Sherpa Gau,
blackwithfemoralargelyyellowish toreddish brown. nr. Kalimpong, 1380m,Darjeeling, India, 1ex.,9.v. 1977.
Head with vertex smooth, shining, impunctate ; frontal This species ishere recorded for the first time from N.
tubercles well developed, subtriangular, contiguous, surface India
smooth,shining,clearlyseparatedposteriorlybyadeepfur Dercetinaviridicyanea Kimoto,1977 Fig. Id
row. Antenna slender, slightly shorterthan body length ; in Ent Basiliensia,2 :387 (Bhutan, Nepal).
preapical segmentsnearly3 1/2 timesalongas wide; first Distribution :Nepal,Bhutan, N.India
segment long, robust, club-shaped ; second short, slightly Material examined. Sherpa Gau, nr. Kalimpong, 1380
longerthanwide,nearly 1/3 as longasfirst; thirdslender, m, Darjeeling, India 9 exs., 9. v. 1977, Bhakta Bahadur.
long,nearlythreetimesas longas second; fourthslender, Lebong, 1600-1860 m,Darjeeling, India 2exs.,8.vi.1975,
long,nearly 11/2timesaslongasthird;fifthnearly 3/4 as W.Wittmer. Jhepi, 1300-1400m, Darjeeling,India, 1ex.,
longasfourth, andfifth totenthsubequal toeachotherin 17.v. 1975, 1ex., 22-23. v. 1975, W. Wittmer. Ladhama-
lengthandshape;eleventhslightlylongerthantenth andits Jhepi, 1100-1350m,Darjeeling,India, 1ex., 22.v.1975, W.
apex pointed. Pronotum transverse, nearly 12/5 times as Wittmer. Chim-Khona (Ghum), 2200 m, Darjeeling, India,
wideaslong;lateralmarginrounded, widestalmostatmid 1ex.,28. v. 1975,W.Wittmer. Lopchu, Darjeeling, India, 1
dle, and stronglynarrowed anteriorly and less strongly so ex., 7. v. 1975, 1ex., 9. v. 1975,W. Wittmer.
posteriorly ; anterior margin slightly rounded anteriorly, Thisspeciesishererecordedfor thefirsttimefromN.
posterior margin distinctly rounded posteriorly and with India
slightemarginationbeforescutellum ;anteriorandposterior Dercetisomaconcolor (Jacoby, 1889)
comerseach witha setigerous pore ; dorsal surfaceconvex Ann. Mus. Civ. Genova sen 2, 7 : 222 (Burma) (Anti
fromside toside,without any depression, and sparsely im pha).
pressedbyminute puncturess. Scutellum convex, subtrian persimilis Kimoto, 1977,EnL Basiliensia,2:389(Bhu
gular,withapexrounded;surfacesmooth,shining,impunc tan) (Arthortus). New Synonym.
tate. Elytronbroaderthan prothoraxat base,slightlywid nepalica Medvedev &Sprecher, 1998,Spixiana,21:35
enedposteriorly andapexrounded;dorsalsurfacestrongly (Nepal). New Synonym.
and closely punctate, interstices of punctures distinctly Distribution : Nepal, Bhutan, India Burma,Thailand,
granulateandnarrowerthanaveragediameterofpunctures. Cambodia,Vietnam.
Length :7.8-8.5 mm. Thisspeciesisvariableonthecolorationofdorsalsur
faces : 1)entirely yellowish to reddish brown, 2)pronotum
Distribution: N. India
Holotype:Mawphlanga 1850m,Magalaya,India, 15. yellowishtoreddishbrown,elytronblack,3)entirelyblack.
v. 1976,W.Wittmer& C.BaroniUrbani (BASEL). Doryxenaminor sp.nov.Fig.2a
Paratype: UpperShillong, 1900m,Magalaya,India, 1 Generallyyellowishbrown; antennapitchyblackwith
ex., 13.v. 1976, W. Wittmer& C. BaroniUrban. first segmentin part brownish ; legs with tibiae and trarsi
This new species closely resembles Cneorane ruguli- pitchyblack.
pennis Baly, 1886,from Northern India but differs from Head withvertex convex, nearly impunctate,with adis
thisinhaving theposteriorfumur brownish. tinct longitudinal depressionat middle,startingfrom ante-
54 Shinsaku Kimoto
rior margin; frontal tuberclesdistinctlyraised,transverse, m,Magalaya India,2 exs., 14.v. 1976,W. Wittmer & C.
subquadrate, contiguous, separated posteriorly by distinct BaroniUrbani. UpperShillong,1900m,Meghalaya, India,
furrow; surface smooth, shining ; inter-antennal space ex 2 exs., 13. v. 1976, W. Wittmer & C. Baroni Urbani. U-
cavated; fronto-clypeus triangularlyraised. Antennaslen matyngar - Cherrapunjee, Meghalaya, India, 4 exs., 16.v.
der,long,nearly2/3 as longas bodylength; firstsegment 1965, W. Wittmer& C.BaroniUrbani.
long, robusL somewhatclub-shaped;secondshort,slender, Thisspecies ishererecorded forthefirsttimefromN.
nearlyhalf as long as first ; third nearly twice as long as India
second,andthird toseventh subequal toeach other inlength Japonitataochracea sp.nov.Fig. If
andshape; eighthslightlyshorterthan seventh,andeighth Generallyochraceous; antenna pitchy black with two
to tenth subequal to each other in length and shape ; elev orthree basal segmentsbrownish.
enth nearly 1 1/4 times as long as tenth and its apex Headwithvertexnotveryconvex,smooth,shining,im
pointed. Pronotumtransverse,nearly11/5timesaswideas punctate, withasmallbutfairlydeepdepressioninmiddle
long ;anteriormargin nearly straight with alittle emargina- ofanteriormargin ;frontal tubercles distinctly raised,elon
tionatmiddle, posteriormargin widely rounded posteriorly; gate,subtriangular, contiguous,delimitedfrom behindbya
lateral margin rounded and distinctly produced laterally at distincLstrong transverse furrow ; inter-antennal space nar
middle; anteriorcomer thickened, and anteriorand poste row, a longitudinal ridge in middle relatively sharp. An
riorcomers eachwithasetigerouspore;dorsalsurfacewith tenna slender, long, slightly shorter than body length, in
a pair of large and deep depressions laterally, and smooth, preapicalsegmentsnearlythree timesaslong as wide;first
shining,entirely covered by distinct punctures. Scutellum segment long, robust club-shaped ; second short, slender,
broaderatbasethanapex,whichiswidelyrounded;surface nearly2/5aslongasfirst; thirdnearlytwiceaslongassec
impunctateatmiddleandsidescoveredwithpunctures. E- ond ; fourth 1 1/4 times as long as third ; fifth slightly
lytronconvexfromsidetoapex,withsideslightlywidened shorterthanfourth,andfifthtotenthsubequaltoeachother
posteriorly and apexrounded ; surface closely covered by inlengthandshape;eleventhslightlylongerthantenthand
distinctpunctures. itsapexpointed. Pronotumconvex,transverse,nearly12/3
Length :8.7-9.5 mm. times as wide as long, widest nearly at 1/3 from anterior
Distribution :Nepal. margin,anteriorandposteriormarginsnearlystraight; ante
Holotype:Godavari,Nepal,25.v.1976,W.Wittmer& riorcomerslightly thickened andposteriorcomerrectangu
C.BaroniUrbani(BASEL). larly emarginate nearangle, which is displaced anteriorly,
Paratype:1ex.,samedataastheholotype. andthesecomerseachwithaseta-bearingpore;dorsalsur
This new species resembles Doryxena grossa Hope, face smooth, shining,nearly impunctate, and with a broad
1831, from Nepal (Fig. 2b), butdiffers from thisspecies in transverseimpressionbehind middle,the impressioninter
being the body length shorter, and having the pronotum ruptedatmiddleanddeeperatsides. Scutellumsubtriangu
wider,andtheantennaandthelegsmuchdarken lar,withitsapexrounded, surfaceconvex, andimpunctate.
Hyphaenia convexicollis MedvedevetSprecher, 1997 Elytron muchbroaderatbasethanpronotum, slightly wid
Coleoptera, Schwanfeld, 1:233(Nepal). ened posteriorly, rounded at apex, surface obsolescently
Distribution:Nepal,N.India punctate, with a distinct costa starting from humerus and
Materialexamined. Chongari,2200-2500 m,U.P.,In reaching almost apical area
dia, 26 exs., 23. v. 1978, W. Wittmer. Chim-Khona Length: 5.0-6.0mm.
(Ghum),2200m, Darjeeling, India 1ex.,W.Wittmer. Distribution :N. India
This species is here recorded for the first time from N. Holotype : Chim-Khona(Ghum), 2200 m, Darjeeling,
India India, 28. v. 1975, W.Wittmer(BASEL).
HyphaeniayasudaiTakizawa, 1985 Fig.5a-b Paratypes: 1ex.,samedataastheholotype. Singmori
EnLRev.Japan,25: 100(Nepal). - Bharapatea-Bug, Darjeeling, India, 1 ex., W. Wittmer.
Distribution:India,Nepal. Jhepi, 1300m, Darjeeling, India 1 ex., 22. v. 1975, W.
Material examined. Chim-Khona (Ghum), 2200 m, Wittmer.
Darjeeling, India, 2exs.,28. v. 1975,3exs.,4. vi. 1975,W. This new species resembles Japanitata eberti(Kimoto,
Wittmer. Lopchu, 1500m, Darjeeling,India,2 exs.,3. v. 1970),fromNepal(Fig. 4c),butseparablefromthisspecies
1976, 4 exs., 9. v. 1975, 2 exs., 31. v. 1975, W. Wittmer. inhavingtheelytronwithoutanylateralcostastaringfrom
Jepi, Darjeeling, India, 4 exs., 22. v. 1975, W. Wittmer. Bi- humerus.
janbari, 800 m, Darjeeling, India, 5 exs., 12. v. 1975, W. Luperus variabilis sp.nov. Fig. le
Wittmer. ManeBhanjang, SukhiaPokri,Darjeeling, India, Generally pitchy black ; pronotum ochraceous ; in
2exs.,9. vi. 1975,W. Wittmer. BarapaniOld Road, 1000 somespecimenselytronwithill-definedyellowishmarking
Newor liule knownChrysomelidae 55
Fig. 4. a.Agetoceraflaviventris Jacoby ; b,Apophyliamaculata Kimoto :c. Ja-
ponilataeberti(Kimoto) :d,Parideaqitadrimaculata sp. nov.
laterally and in most pale colored specimens head, prono Scutellum small, subtriangular, smooth, impunctate. Ely
tum and elytron generally yellowish brown. tron hardly broader at base than prothorax, slightly widened
Head with vertex not very convex, surface smooth, posteriorly and apex rounded ; dorsal surface closely im
shining, sparsely impressed by minute punctures : frontal pressed by distinct punctures and their interstices smooth,
tubercles well developed, subtriangular, contiguous, their shining.
anterior angles inserted between antennal insertions, and Length :3.7-4.0mm.
distinctly separated posteriorly, surface smooth, shining ; Distribution : Pakistan.
fronto-clypeustriangularly raised, surface smooth, shining. Holotype : Miandam, 1800-2300 m. Swat, Pakistan, 2.
Antenna slender, slightly shorter than body length ; in vi. 1978,W. Wittmer.
preapicalsegments nearly three times as long as wide : first Paratypes :3exs., same data as the holotype, and 1ex.,
segment long, robust, somewhat club-shaped ;second short, same data as the holotype, but 3. vi. 1978. Cabral, 230-245
nearly 1/3 as long as first ; third slender, nearly twice as in.Swat.Pakistan. 10ex.,8.vi. 1978,W.Wittmer. Shogan
longassecond :fourth slightly longer than third, and fourth V., Pakistan, 4 exs., 17. vi. 1977. W. WrrrMER& C. Baroni
to tenth subequal to each other in length and shape ; elev Urbani. Kawai, Khagan V., 1450-1800 m, Pakistan. 1ex.,
enth slightly longerthan tenth and its apex pointed. Prono 1977,W. WrrrMER & M. Brancucci.
tumtransverse, nearly 12/5 times as wideas long,widest This new species resembles Luperusflavimanus Weise,
almostat 1/3 fromanterior margin and gradually narrowed 1889, from Sze-tschuan, but differs from this species, in
anteriorly and posteriorly, anterior margin nearly straight having the antenna robuster, the third antennal segment
and posterior margin widely rounded posteriorly ; dorsal shorter than the first, and the coloration of dorsal surfaces
surfaceconvex from side to side, and closely impressed by pitchy black to yellowish brown.
distinct punctures and their interstice smooth, shining. Mimastrafulva KimotoetTakizawa, 1983 Fig. 5c
56 Shinsaku Kimoto
Bull.Natn. Sci. Mus., Tokyo,ser. A,6 :87 (Nepal). sam, India, 16. v. 1976. W. Wittmer & C. Baroni Urbani
Distribution :Nepal, N.India (BASEL).
Material examined. Jhepi, 1300-1400 m, Darjeeling, Paratypes:6exs., samedataas the holotype.
India, 7exs., 22-23. v. 1975, W. Wittmer. Pom, S.Gangtok, This new species resemnbles Monolepta trifasciata Ja
Sikkim, India, 4 exs., 17. iv. 1977, Bhakta Bahadur. Sing- coby, 1896, from India, but differs from this species in hav
mari - Barapatea Bung, Darjeeling, India, 1 ex., W. ingtheelytralblackishtransverse bandsituatedat 1/3from
theapicalmargin.
Wittmer.
This species is here recorded for the first time from N. Monolepta gantokensis spec.nov. Fig.2c
Generally reddish brown ; head and pronotum gener
India
Monolepataanalis Weise,1915 ally yellowish brown ; scutellum pitchy black ; antenna
Ergebnis 2,Dtsche Zentr.-Afr.-Exp.,Zool., 1: 117(new pitchybrownwiththreebasal segments paler.
name for Candezea humeralis Jacoby, 1903, nee Galleruca Head with vertex not very convex, surface smooth,
humeralis Weber, 1801). shining, sparsely impressed by minute punctures ; frontal
humeralisJacoby, 1903, Ann. Soc. EnLBelg., 47 : 125 tubercleswelldeveloped,contiguous, subtriangular,surface
(Anamalais) (Candezea). smoothandshining. Antennaslender,slightlyshorterthan
Distribution :IndiaBhutan. bodylength,inpreapicalsegmentsnearlyfivetimesaslong
Material examined. 87 km von Phuntsholing, Bhutan, aswide;firstsegmentlong,robust,somewhatclub-shaped;
1ex., 22. 5. 1972. second short, nearly 1/3 as long as first ; third subequal to
This species is here recorded for the first time from second in length ;fourth nearly three times as long as third,
and fourth to tenth subequal to each other in length and
Bhutan.
Monolepta atosuji sp.nov. Fig.5d shape ; eleventh subequal to tenth in length but its apex
Generally pitchy black ; head with fronto-clypeus red pointed. Pronotum transverse, convex from side to side,
dish brown ; pronotum yellowish brown with latero- nearly twice as wide as long, widest almost at middle and
marginal area pitchy black ; elytron yellowish brown with gradually narrowed anteriorly and posteriorly ; anterior
basal, lateral, apical and sutural margins broadly pitchy margin slightly rounded anteriorly and posterior margin
black, and together with a transverse band situated at 1/3 widely rounded posteriorly ; dorsal surface sparsely im
fromapicalmarginpitchyblack;antennapitchyblackwith pressed by minute punctures. Scutellum subtriangular, sur
basal two or three segmentspaler;legs reddish brown with face smooth, shining, nearly impunctate. Elytron hardly
tarsidarker. broaderatbase than prothorax,slightly widened posteriorly,
Headwithvertexnotveryconvex,smooth,shining,im and apex rounded, surface closely impressed by distinct
punctate ; inter-ocular space longitudinally raised, and punctures.
nearlyaswideaswidthofeye;frontal tubercles welldevel Inmale,elytron withanelongate largefovea laterally.
oped, contiguous, subtriangular. Antenna slender, slightly Length: 5.3-6.8 mm.
shorter than body length, in preapcial segments nearly five Distribution :N. India
times aslong as wide ;first segmentlong,robust, somewhat Holotype :Rani Pull, 910 m, S. Gantok, Sikkim, India,
club-shaped; second robust, nearly 2/5 as long as first ; 22. iv. 1977,BhaktaBahadur(BASEL).
third subequal to second in length but slenderer ; fourth Paratypes : 9 exs., same data as the holotype. Ahu
nearly2 1/2 timesaslongasthird,andfourthtotenthsub- Khola, nr. Rani Pull, Sikkim, India, 1 ex., 23. iv. 1977,
equaltoeachotherinlengthandshape;eleventhsubequal BhaktaBahadur.
totenth inlengthbut itsapex pointed. Pronotumtransverse, This new species resembles Monolepta labiata (Ja
convex,nearly 12/3 times aswide aslong,widestalmostat coby),1900, from Bengal, but differs in having the prono
anterior margin, and gradually narrowed posteriorly, ante tum convex from side to side and without a pair ofdistinct
rior margin slightlyroundedanteriorly and posteriormargin depressions, and the dorsal surfaces with pronotum
widely rounded posteriorly ; dorsal surface with a pair of yelolowishbrownandelytronreddishbrown.
shallowdepressions laterally,andclosely impressedbyfine Monolepta labiatella sp.nov.
punctures. Scutellumsmall,subtriangular, surfacesmooth, Generally reddish to yellowish brown ; antenna with
nearly impunctate. Elytron hardly broader at base than fourorfive apical segmentsinfuscate.
prothorax, slightlywidenedposteiroly, and apexrounded; Head with vertex not very convex, surface finely sha-
dorsal surfacecloselyimpressedwith distinctpunctures. greened,sparselyimpressedbyfinepunctures; inter-ocular
Length: 3.5 mm. space distinctly narrower than width of eye ; frontal tuber
Distribution: N. India cles well developed, subtriangular, surface finely sha-
Holotype :Umatyngar, Cherranpunjee, Meghalaya,As greened. Antenna slender,slightly shorter than bodylength,