New or Little Known Carabus and Cychrus (Coleoptera,Carabidae)from the Qinling Mountains in Shaanxi Province,Central China

15 Pages·1993·9.7 MB·

by  Imura, Y

Elyt,a,T1okyo 21(2):363-377,Nov.l5,1993
Newor Little KnownCarabusandCychrus(Coleoptera,
Carabidae) from theQinlingMountainsin
Shaanxi Province, Central China
Yuki IMURA
Shinohara-chol249-8, Kohoku-ku,Yokohama,222Japan
Abstract Twonewspeciesand fournewsubspeciesofthegenus(:a,・abus(s.lat.)
andtwonewsubspeciesofthegenusCy(・1/,1-1saredescribedfromtheQinlingMountains
in Shaanxi Province, Central China: Ca/・1(加.l, (K/11g()(・a/・aりf!Is)z/Ill/)altosp. nov., C
s) ( lpsoca,-a/ﾂ11s /af・, of/l,1/111geMsl subs nov.,C (「01;l()(・c11・a/)11s)s/1a1/e.l1/a,7gsp nov., C
(Pagoa/'a/)ll.l) 1-asses l'/ ills(/line/1,Igloo/o,・subsp nov., C(P.l,e1l1oc1-a川0,1)raf฀als/1川a _
/ﾆI subsp nov., C (9(・co/,ro/a0/・!Is)e,、-1gll1lsfiM1a,111ssubsp nov., CJ,(・/11-1ls lsplnos1l1.
actio,7s/ssubsp nov.,and C lsp1,1os1l.・l 1/1-ale,Isissubsp nov.
TheQinlingMountainsliefromeast towest forabout 400kminthesouthern
partofShaanxiProvince,CentralChina,withthehighestpointmarkedbyMt.Taibai
Shanatthecentralpart,whichattainstoaheightof 3,767m. Sincethecarabidfauna
ofthismountainrangehasnot been intensivelyinvestigatedasyet, our knowledgeis
nomorethanfragmentaryevenonthecomparativelywell-knowngeneraCa1abus(s.
lat )and (:),(・h/・11s.
Recently, Ihadanopportunitytoexaminealongseriesofexampleso「carabid
beetlescollectedfromseveral localitiesontheQinlingMountainsincludingratherhigh
altitudinal areas. Thecollectioncontainssomeunnamed formsofthegeneraCal・abus
(s lat )andC、、、(/11'its,and I amgoingtodescribeeightnewtaxainthepresentpaper.
Theabbreviationsemployedhereinarethesameasthoseexplainedinmypre-
viouspapers,andtheho1otypesofallthenewlydescribedtaxaarepreservedinthecol-
lectionoftheDepartmentofZoology, NationalScienceMuseum(Nat.Hist ), Tokyo,
andtheparatypesaredepositedinmycollection.
Beforegoingfurther,Iwishtoexpressmyheartythanksto Dr.Shun-lobiUENoof
theNationalScienceMuseum(Nat.Hist),Tokyo,forcriticallyreadingthemanuscript
of thispaper. ThanksarealsoduetoDrs. 01afJAGERandRudiger KRAUSEof the
StaatlichesMuseumfiirTierkunde,Dresden,whokindlytookthetroublefor1oanjng
metheholotypeOfCal'abus''ugoslsslmusBREUNING,andtoMessrs. Helmut ScHUTzE,
Gleichen,Satoshi KoIwAYA,Tokyo,andPin-JunFAN,Xi'an,、l,.hokindlysupported
thisstudy invariousways.
e, 1. Cara加s(KM/'gocara加s) M฀ajl' IMURA s nov.
(Figs.1,7)
Length:19.4-21,l mm(fromapicalmarginofclypeustoapicesofelytra)
364 YukiIMURA
Copperywithfaintred-purplishlustre. Elevatedpartsofelytraandventerblack.
Appendagesalsoblack, though themandibles, buccal appendagesand basal partsof
antennaearereddishbrownaswellastibiaeandtarsi.
Headexcept for mandiblessubquadrate, alittlewider than long, withlargeand
strongly protrudent eyes; apicalmargin of labrum slightlyemarginate;clypeussub-
quadrate, a little narrower than labrum; frontal furrowsshallow, with thesurface
weaklyandirregularlyrugu1ose; fronsslightlyconvexabove,withthesurfaceweakly
rugoseandsparselypunctulate;mandiblesshort andstout,stronglyarcuateinwards,
andsharplypointed at theapices; retinaculumof mandiblebidentate, theright one
beingalittlesmallerthantheleft;apicalsegmentsofpalpinotsowidelydilatedinboth
sexes; penultimatesegment of labial palpusbisetose;median toothofmentumtrian-
gularlyprotrudentanteriadthoughalittleshorterthanlateral lobes,withtheapexnot
sharply pointed; submentum bisetose, with thesurfacesmooth; antennae filiform,
reaching themiddleofelytrainmaleandreachingbasal two-fifthsofthemin female,
pubescent fromsegment5, withhairlessventraldepressionsfromsegment 7to9in
male; relativelengthsofscapeandsegments2-4asfollows: - 1:0.56:1:0.68.
Pronotumsubcordate,widerthanlong,widestataboutthemiddle,morestrongly
narrowedtowardsapexthantowardsbase;PW/HW l.35-1.38(M I.37),PW/PL128-
1.31 (M I 29), PW/PAW 156-1.66 (MI 60), PW/PBW 1.42-1.52 (M I 45), PBW/
PAW 1.09-1.12(M1.10); apicalmargin feeblyemarginate, front anglesobtuseand
barelyproducedanteriad;sidesgentlyroundedinfrontandweaklysinuatebehindthe
middle; hind angles subtriangularly protrudent posteriad though short, with the
apicesroundedandbarelybentventrad;discslightlyconvexaboveandratherdensely
scatteredwithshallowpunctures;medianlongitudinallineevidentlyimpressedthrough-
outthoughverynarrow;basal foveaesmallandveryshallow;fourmarginalsetaein-
sertedoneithersideofpronotaldisc,threemedic-anteriorlyandonenearhindangle.
Elytraelongateoval,widestbehindthemiddle,moregraduallynarrowedtowards
bases than towardsapiceswith theshoulders rather effaced; EW/PW l43-1.54 (M
1.49),EL/EW153-1.60(MI57);sculpturetriploidorpartlypentap1oid, heterody-
name;primariesthewidestandmoststronglyraised,rather frequentlyandirregularly
segmentedbylargefoveolestoformrowsofcostaeorofcallositiesinvariouslengths;
primaryfoveolesshallowlyconcaveandusuallyinvadingtheadjacenttertiaries;second-
ariesalittleweakerandnarrowerthanprimaries, composedof frequentlyinterrupted
lowcostae;tertiariesalittleweakerthansecondaries,composedofirregularlysegmented
rjdgesor rowsofgranules,eachofwhichissometimessplitintotwo rowsorpartly
djsappeared;striaebetweenintervalsonlypartlyremainingasirregularlyset rowsof
punctures;umbilicateseriesindicatedbyratheranirregularlysetrowofgranules,atthe
outsideofwhicharerecognisedratherdensegranules.
pro-andmesepisternasmooth,metepisternaandsidesofsternitespartlyscattered
wjthvaguepuncturesandwrinkles;metacoxatrisetose;sternalsulciabsent;basalfour
segmentsofmaleforetarsusdilated,withhairpadsontheventralsurface.
MalegenitaliaasshowninFig 7a-c;aedeagusslenderandgentlycurvedventrad;
(a・labusandCych1・usfromtheQinlingMountains 365
Figs.1-6. Holotypes (cl)of (arabusspp fromthe QinlingMountainsinShaanxi Province,
Central China. - 1, Ca,・abus (Rhigoca・,abus)zhubajlesp nov., from Zhouzhi Xian;2,
C (Hypsocarabus) latroqinlingenslssubsp nov.,fromChang'anXian; 3, C (Tomocarabus)
shaheshangsp nov., fromChang'anXian;4,C. (Pagocarabus)crasses(ulptusqunqingico1o,
subsp nov., from Hu Xian, 5, C. (Pseudoc・・,amon) taibais,hanicus fanisubsp nov. from
,
Chang'anXian;6,C (Eccoptolabrus)eaguusfania,1ussubsp nov.,fromZhouzhiXian.
366 Yuki [MURA
apicalloberathershortwithroundedtip;ostiumlobeabsent; ligula,orbasalsclerite,
reducedtoapatchofpigmentedgranulessituatedtransverselyat alittle rightsideof
thebasal partofendophallus;medianportionofendopha1lusrather stronglyinflated
dorso-1aterad. FemalegenitaliaasshowninFig 7d-f,outerplateofvaginalapophy-
、isnarrowandvestigial, inner platelarge, almost twiceaswideaslong, transversely
putkidney-likeinshape,andlightlysclerotized.
T?pcseries. Holotype: 2,000malt.,ontheQinlingMountainsinZhouzhi
,ca
Xian. Shaanxi Province, Central China, 21~23-V-1993. Paratypes (including al-
lotype):2 1 ,samedataasfortheholotype.
,
Notes. OnlyonespeciesbelongingtoR/1lgo(・al・abus,c/foulDEuvE(1989、p 228),
hashithertobeen known from the QinlingMountains, and the present new species
becomesasecondrepresentativeof thesubgenusoccurringonthismountainrange.
Although l knowDEuvE'sspeciesonlyfrom theoriginal description, the twospecies
、eemtobereadilydiscriminated fromeachotherbydifferentlyshapedaedeagus.
Thepresent newspeciesisalsocomparablewith C (」R) to、loensls, recentlydo-
s.cribed by DEuvE(1992, p 54) fromGansu, but isdiscriminated from thelatter by
having thegularsetaeand thedifferentlyshaped aedeagus. Position of the hairless
depressionson theventral surface of male antennae and thesculptural conditions
ofpronotumandelytraarealsodiagnostic.
Zhubajleisthenameof asensualmonsterpigappearingasoneofthethree fol-
lowersofTangsenginthestoryofXiyouji.
2. Ca,abus(Hypsocarabus)latroqinlingensl'sIMuRA,sp
nov.
(Figs 2,8)
Length:17.0-l9.0mm(fromapicalmarginofclypeustoapicesofelytra).
Most closelyallied to the nominotypica1subspeciesof Cal・abus ( 、,pso(a・/abus)
titleSEMENow (1898, p367) described from ''Lun-ngan-fu'' (=Pingwu at present,
situatedatthenorthernendofSichuanneartheGansuborders), but differsfromit in
the following respects: dorsalsurfacereddishcoppery, sometimeswith red-purplish
lustre. while it is much darker in the nominotypical subspecies; apical margin of
labrummoredeeplyemarginate; penultimatesegmentoflabial palpususuallybearing
Figs 7-12. GenitalorganofCal・abtlsspp fromthe QinlingMountainsinShaanxi Province,
Central China. - 7,Ca・,abtls(1thigo(・al・abus)・/1ubajlesp nov., from Zhouzhi Xian; 8, C
(1iJlp.so(・al・abus) latl-oqln/lnge,Isls' subsp nov., fromChang'an Xian; 9, C. C「;omoca,・abus)
s1/ahesha11gsp nov.,fromChang'anXian、le,C(Pagoca1-11bus)(・1・as.les(・1llptuscf1″It/l,1g1(・ole/-
subsp nov., fromHu Xian; 11, C (Psel,decl,arno,1) 'a10a1s7/a,71(・11s fit川subsp nov., from
Chang'anXian;12, C (E(、(・)ptolab・lfis)exlgliusfanl'a,1ll.lsubsp nov., fromZhouzhi Xian.
- a-c.Malegenitalia;a,aedeaguswithevertedendophallus(rightlateralview); b,apical
partofaedeagus(rightlateral view);c,ditto (dorsal view). - d-f, Femalegenitalia1d,
vaginalapophysis(rightlateralview);e, outerplateofvaginalapophysis(ventralview); f,
innerplateofvaginalapophysis(dorsalview). Scale:1mmfora,05mmforb-f.
tea・,abusand (ychrusfromtheQinlingMountains 367
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368 Yuki lMURA
threesetae, thoughoneofthemnear thedistalendis oftenabsent;medjan toothof
mentumslightlybutobviously bilobateat theapex;submentumasetose, whjIejt js
multlsetosein thenominotypica1subspecies; pronotumalittleslenderer, wjth front
anglesobtuseand rounded,sidesratherstronglycontractedbehindthemiddle, shat_
lowlysinuateataboutone-fourthfrombase,andthenslightlybutevidentlydjvergent
towards hind angles, which aresubtriangularly protruded postero-1aterad wjth the
apicesrathersharplypointed1elytra1intervalsless frequentlysegmented, andstrjae
betweentheintervalsmorestronglyimpressed;apicallobeofaedeaguswjderjnlatera1
、levv.Shallowlybutobviouslyconcaveventro-Iateradat therightsjdenearthebase,
、、1endererandmuchmoreelongate indorsal view; ostiumlobealmost vestjgja1, and
rightbasallobeofendophallusbeingabsent.
F「omSubsp. 'runs/7anenslsDEUVEandsubsp. /ll,a,7g/ongenslsDEUvE, thepresent
new subspecies iseasily distinguished by thedifferent configurationof pronotum,
aedeagusand/orofelytra.
T、'peso''tes. Holotype: 2,410malt.,Chang'an Xian,26-V-1993. Paratypes
,
(including allotype): 7 17 2,410-2,650malt., Chang'an Xjan, 25~2g_v_
, ,
l993. All fromtheQinlingMountainsinShaanxiProvince, Central China.
Notes. This is presumably the most specialised subspecies of Ca,・abus /at,・o
characterisedbylossor atrophyofsuchcharactersasgularsetae, ostium lobeand
right basal lobeofendopha1lusofmalegenitalia, whichare reserved inall theother
subspecieshithertoknown. Bothpronotumand aedeagusof the present subspecjes
areconsiderablydifferentinshapefromthoseoftheothers. Inaddition,itistheunjque
characteristicoftheQinIingsubspeciestohaveinhighratiotheadditionalsetaeonthe
distalpartofpenultimatesegmentof labial palpus. It ispossiblethatthisraceisdis-
tinctspecificallyfromtheSichuaneseones,buthereIprovisionallydescribeitasasub_
speciesof thelatter.
Ca●abus(Tomocal'abus)sltahes/lang IMURA,sp nov
(Figs 3,9)
Length: 14.8-16.6mm(fromapicalmarginofclypeustoapicesofelytra).
Black, mat, except for mandiblesand tarsi which are reddish brown. Buccal
appendagesand antennae also reddishbrownespecially in thebasal part of each
segment.
Headexcept formandiblessubquadrate, alittlewider thanlong, withlargeand
strongly protrudent eyes; apical margin of labrum moderatelyemarginate, clypeus
subquadrate, alittledilatedposteriad, nearlyaswideasoralittlewiderthanlabrum;
frontalfurrowsshallowlyconcave,withthesurfacestronglyandirregularlyrugu1ose;
frons gently convex above, with the surface weakly rugose-punctate. mandibles
short andstout, rather strongly arcuateinwardsandsharply pointedat theapices;
retinaculum ofmandiblebidentate, theright onebeingalittlesmaller than theleft;
apicalsegmentsof palpislightlydilatedinbothsexes, penultimatesegment of labja1
ca・,ab1l、.andCy(・/11,usfromtheQinlingMountains 369
palpusbjsetose;medjan toothofmentumtriangularly protrudent anteriad,She「te「
thanthelaterallobes,withtheapexoftenalittlere-entrantatthemiddle;Submentum
bjsetoseorsometjmesbearingasingleadditionalsetainside,withthesurfaceirre9ula「一
1yrugu1ose;antennaefiliform,reachingbasaltwo-fifthsofelytrainmaleandreachin9
basalquarter infemale,entirelypubescent fromsegment5, withhairlessvent「alde-
pressjonsfromsegment6toleinmale,relativelengthsofscapeandsegments2-4as
follows: - 1: 0.60: 1:0.75.
pronotumsubquadrate,wider thanlong,andwidest alittlebeforethemiddle;
pw/Hw 1.43_1.48(MI45), PW/PL 1.31-1.35 (MI33), PW/PAW1.43-1.52 (M
l48),pW/pBW128_1.32(MI29),PBW/PAW1.11-1.19 (M1.15); apicalma「9in
sljghtlyemargjnate, front anglesobtuseand barelyproducedanteriad;sideswidely
andweaklyarcuatein front, shallowlysinuatebehind themiddle, thensubParalle1
towardshjndangleswhichareweaklyprotrudent posteriadwith theapicesSubt「i-
angularlypojnted: discgentlyconvexabove, ratherdenselyscattered with irre9ula「
wrjnklesandlargeshallowpuncturesonthesurface;medianlongitudinal linenarrow
andshallowlyimpressed, partlyunclear;basal foveaeratherdeeplyconcavethou9h
small;sixorsevenmarginalsetaeinsertedoneithersideofpronotaldisc, fiveorSix
medic-anteriorly,onenearhindangle.
Elytraovateandconvex,widestatabout themiddleorslightlybehindthatlevel,
morestronglycontractedtowardsapicesthantowardsbases,withtheshouldersrathe「
djstjnct;EW/pW142-1.57(MI.51),EL/EW1.35-1.41(MI.38);sculptureheptaP1oid
heterodyname;primariesthemoststronglyprominent,composedofratherfrequently
segmentedshortcostaeorcallositiesinvariouslengths;primaryfoveolesratherdeeply
concaveandusuallyinvading theadjacent quarternaries; both secondaries and ter-
tjarjesnarrower andmoreweaklyconvexthaninprimaries,composedof frequently
and jrregularly segmented rows ofcostae; quarternaries weaker than theothe「in-
tervals,andusuallyreducedtorowsofgranules;striaebetweenintervalsonlypa「tly
recognjsedassparselysetrowsoffinepunctures; umbilicateseriesindicatedbyrather
ofgranules, at theoutsideofwhicharerecognisedratherdense
aregularlyset row
granules.
pro_andmesepjsternasmooth,metepisternaand sidesofsternites vaguelypunc-
tateandfinelywrinkled;metacoxabisetose, proximalsetabein9absent;SternalSulci
completelyandprominentlycarved;basal four segmentsofmaleforetarsusdilated,
withhair padsontheventralsurface.
MalegenjtaljaasshownjnFig9a-c;aedeaguswidelyandratherstronglyarcuate
ventrad;itsapical lobeveryslender, almostparallel-sidedwithroundedtipinlateral
vjew,subtriangularlypointedandmoderatelysinuateon the right lateralmar9inin
dorsalvjew;ostiumlobelargeandrobust, withtheapexbilobate. Femalegenitalia
asshownjnFjg9d-f;outerplateofvaginalapophysisnarrowandalmosttransparent
except forthelongitudinalmid-linewhichispigmentedandalittlesclerotized; inner
platecircular inoutline,weaklysclerotizedandsomewhatcup-like.
T、,pesertes. Holotype: , ca.1,900m alt., Chang'anXian, 9-V-1993. Para-
370 Yuki iMURA
Fig.13. ThehabitatofCa・labtls(Hypsoca,・abus)/af・toqinlinge,Isissubsp nov, c (Tome(al・ab1ls)
) s/ta/7es/7a'lg sp nov., C (Pagoca,・a฀11s ・ (・l aNescll/prfl ll,lc/1Mglco/01・subsp nov, and c
ﾆ (PSelldoc'a川on)fat・ls/1a'1lclls.f m sub nov.(subalpinezoneoftheQinIingMounlajnsjn
Chang'anXian,ca 2,500minaltitude).
t、pes(includingallotype):8 3 ( , samedata as for the holotype; 3
2.000malt., Hu Xian, 11-V-199,3; I , ca. 1,850m alt., Taibai Xian, 19_v_1g, gc3a;.
7 1 , ca 2,000malt.,Zhouzhi Xian, 21-V-1993. All fromtheQinlingMoun_
,
tainsinShaanxiProvince, CentralChina.
Notes. ThisnewspeciesisverysimilarinexternalfeaturestoCa,・abus(Tome(ara_
bus)frate1'clilusREITTERdistributedfromKoreaandtheadjacent regions,butiseasjly
distinguishablefromitbylesswidelydilatedterminalsegmentsof buccal palpjand
differentconfigurationofaedeagus.
ItmayalsobecomparablewithC.(T)s/1aa,1.xle,IsisrecentlydescribedbyDEUvE
(l991,P ie5) from ''AnkangXian'' situatedinthesouthernmost part ofShaanxj
Provincenear theSichuanand theHubei borders. SincetheAnkang xianspecjes
was described on the basis of a single female specimen without accompanying
illustrations,carefulconsiderationisnecessarytodetermineitstruetaxonomjcposjtjon.
Judgingfromtheoriginaldescription,however, theQinIingspeciesseemstobedjs_
criminatedfromtheAnkangXianoneatleastbythefollowingfivecharacters:1)sjze
alittlesmaller;2)penultimatesegmentoflabialpalpusalwayswithtwosetae, whjlejt
istrisetoseinthelatter;3)antennaelonger,reachingbasalquarterofelytrainfemale,
whiletheyreachbasalone-sixthsinthelatter;4)pronotumwithfivetosixmargjna1
Setae,whereasitbearsthreeorfour inthelatter;5)sternalsulci prominentlycarved,
thoughfinelysointhelatter.
Thenameofthisnewspecies,s/1aheshang,comesfromthatofawaterspritewho
(、:al・a/)111.・and C、,(,1/1-ll.l,fromtheQinlingMountains 371
appearsasoneofthethreefollowersofTangsenginthestoryofXiyouj
4. Carabus(Pagocarabus)crassescldpt1ʻsquntlI'ngicolo'' IMURA,subsp nov.
(Figs 4,10)
Length:21.8-26.0mm(fromapicalmarginofclypeustoapicesofelytra).
Closely allied to subsp. ・,ugoslsslnnlsBREUNING (l943, p ile)'' described from
'Pelting-Chan'' (thenameof amountainrangestreching from west to east between
MinshanandQinling, at thesouthernendof Gansu), anddoubtlessbelongsto the
samesubspecies-group, but distinguished fromit by the following respects: dorsal
surface more light bluish and not so stronglypurplish as in ・/ugoslsslmus; pronotum
slenderer,withthesidesmoreweaklyroundedinfront, andhindanglesmoresharply
pointed,elytral intervalsmoreweakly raised, with theelevated partsless frequently
fused with one another: preapica1emargination of elytrashallower: aedeagusmore
strongly tumid bilateradalittlebeforethemiddle, with theapical lobeslenderer and
morestronglyarcuateventrad.
T、peso,・1es. Holotype: , ca 2,000m alt., Hu Xian, 11-V-1993. Paratypes
(jncludingallotype):2 2 o , samedata as for theholotype; 1 , ca. 1,500m
,
alt.,Baoji Xian, 13-V-1993, 11 3 o , ca 2,000malt., Zhouzhi Xian, 21~23-
,
v_1993;1 o 2,410m,alt.,Chang'anXian,26-V-I993. All fromtheQinIingMoun-
tains inShaanxi Province, Central China.
Notes. This new subspecies is named after its unique coloration, ''Qunqing'',
whichmeanspurplishblueinChinese.
I,' 5. Cara加s(PseM6foeraafon) ta''฀al'sllanl'el's/iml MURA subs nov.
(Figs 5,11)
Length:21.0-26.2mm(fromapicalmarginofclypeustoapicesofelytra).
Distinguishedfromthenominotypica1subspeciesbythefollowingcharacters: the
secondantennalsegmentobviouslyshorterthanthefourth,thoughtheformerislonger
than thelatter in thenominotypicalsubspecies, pronotuma littleslenderer (PW/PL
1.26-1.42, M I 33, whileitIs l 47in thenominotypicalsubspecies); eIytra1sculpture
nottypicallytriploidasinthenominotypica1subspecies,butis regardedtobepenta-
or heptap1oid, i.e., tertiariesnot clearly recognisableasasinglerowo「granules, but
usuallysplit intothreeto fiveirregularlyset rowsofgranules.
Malegenitaliaasshownin Fig.11a-c; aedeagusrather abruptlycurvedventrad
atthebasalthird,subcylindrica1andalmostparallel-sidedinmedianportion, obviously
1) Cal・a t,s(Pa.g・o(・al・a/)list(,1・asses('ll/ rllsl'llgoslssl'1111sBREUNINc,stat nov.
1wasabletoexaminetheholotypespecimenof ('.11'igosls1.・1'1111.l, BREUNl1、?Gin thecourseofthe
present study. Judging mainly fromclosesimilarity intheconformation ofgenitalia, BREuNINc's
speciesisconsideredtobealocal raceofC.('1'asses('ll//1tlsKRAAlz, thoughratherpeculiarinexternal
features. Itisthereforedowngradedtoasubspeciesofthelatter.
l72 Yuki IMURA
protuberant rightlateradataboutapicalquarter, thenslightlyconcavelatero-ventrad
on therightside, withtheapexshort androunded; ostiumloberather slender, with
theapexslightlybilobate. Femalegenitaliaasshown in Fig. l1d-f; outer plateof
aginalapophysisabout17timesaslongaswide, widest near theapex, obviously
sinuatebehindthemiddle,withtheanteriormarginshallowlyemarginate, theposterior
beingre-entrantat themiddle; inner platealmost round-shapedor alittlepiriform,
somewhatcup-like,andstronglypigmentedandsclerotisedalongtheanteriormargin
and thelongitudinalmid-line.
T、,peseries. Holotype: (j', 2,410-2,760m alt., on the Qinling Mountains in
(hang'an Xian, Shaanxi Province, Central China, 25~29-V-1993. Paratypes (in-
cludingallotype):42 36 samedataasfortheholotype.
,
Notes. AlthoughIregardedtheChang'anXianspecimenasbelongingtoCarabus
( seucfoc1'a fon) falOa's/7aMci,sDEUvEdescribed from the''Taibai Mountains'', which
meaneither the QinlingMountainsorapart of themnear Mt. TaibaiShan,specific
determinationoftheformerisnotdefinitiveatpresent, sincethelatter wasdescribed
onthebasisofasinglefemalespecimen. DiscoveryofmalescorrespondingtoDEuvE's
formmaysolvetheproblemonthetaxonomicrelationshipbetweenthesetworaces.
6 Carabus(Eccoptolabrus)enguusfanla'tusIMURA,subsp
nov
(Figs 6,12)
Length: 16.7-18.6mm(fromapicalmarginofclypeustoapicesofelytra).
Reddishcopperytodarkbrownexcept fortheelevated partsofelytrawhichare
black,venterandappendagesalsoblack,thoughtheantennaearealittlereddish.
Headexcept formandiblessubquadrateand longer thanwide; apicalmarginof
labrumdeeplyemarginate; frontal furrowsmoderatelyor rather deeplyconcave, with
thesurfacestronglyrugose-punctateaswellasfronsanddorsalsurfaceofheadbehind
e、es; retinaculumofrightmandiblebidentate, itsanterior toothmuchlargerthanthe
posterior; that of leftmandiblealso bidentate, though theanterior tooth issubequa1
inlength to theposterior whichfusetothebasal innertooth; apicalsegment of galea
concaveinwards, withthedorsal marginedgedandslightlyemarginateinapical half:
apical segmentsof palpi triangularlydilated, morestronglyso inmale; penultimate
segment of labial palpusbisetose; median toothofmentumshorter thanlateral lobes,
with the apex triangularly pointed, submentumasetose, with the surface finely and
transverselywrinkled; antennaenearlyextending to themiddleof elytrainmaleand
reachingbasal two-fifthsofthemin female;relativelengthsofscapeandsegment 2-4
as follows: - 1: 0.87:1: 0.80.
Pronotumstronglycordate, widestalittlebeforethemiddle, morestronglynar-
rowed towards base than towardsapex which is shallowlyemarginate, with front
anglesobtuseandbarelyproducedanteriad;sideswidelyarcuatein front, strongly
contracted posteriad,shallowlysinuate at basal two-fifths, and then almost parallel-
sided towardshindangleswhichareslightlyprotruded posteriadandsubtriangularly