New Late Cretaceous (Santonian and Campanian) gastropods from California and Baja California, Mexico

2005·8.2 MB·

THE NAUTILUS 119(4):133-14S. :Z005 Page 133
New
Late Cretaceous (Santonian and Campanian) gastropods
from California and Baja California, Mexico
Richard L. Squires LouElla R. Saul
Depiutiiu'iit (it Cieological Sciences ln\crtcliratc PalrdutoioUA Section
California State Uni\eisit\ Natural Miston Museum oi Lcis Angeles
Northridge, CA 91330-S266 USA Counts-
[email protected] 900 Exposition Boulevard
Los Angeles, CA 90007 USA
[email protected]
ABSTRACT Angeles Comity, allowed us to incoiporate two additional
species into Bullamirifica. These are Bullamirifica
Three new genera and six new species of shallow-maiine Late verruca new genus and species from the Coniacian
Cretaceous gastropods are reported from various formations in Member IV ofthe Redding Formation in the Oak Run
Cahfomia and from one formation in Baja California, Mexico.
Tegula jcanac new species, of early Campanian age, is the area, northern California, and Bullamirifica anikitos
Dailey and Popenoe (1966) new combination from the
earhest kno\\ai species of this trochid genus. Neiitn (sub-
genus?) oivvillensis new species is the second knowai Earl\- middle Campanian Pigeon Point Formation soutliwest of
Campanian neritid from California. The cerithioid Bullarnir- San Francisco, northern California; the middle upper
ifica new genus is represented bv diree species: Biilhiiiiiiifica Campanian Punta Baja Formation, Baja Cahfomia,
cernica new species of Coniacian age; BuUninirifica cicgaiis Mexico; and the upper Campanian Jalama Formation,
newspecies ofearlyCampanian age; and BuUinnirificiiainiktos soudiem Caliiornia (Figiu-e 1). "Cimolithium miija-
(Dailev and Popenoe, 1966) ofmiddle to late (.Campanian age. kocnse" (Nagao, 1934) and "Vican/a (Shoshiroia) i/abei"
The latter species has the most widespread distribution, with Kamada, 1960, reported- by Periilhat-Montoya (1968)
ofocrcnuira.reMnicneisjtrionpissoumdeliielrontaCnaleiwfogrenniausaannddnsoprectileisemofBSaajnatoCnailain- Irom Baja California, Mexico (see Figure 1, fomiabon 6),
age, and Paxitropis dicriota new genus and species of Late are judgedb\-ustobe syiiommis ofBullamirifica ainiktos.
Santonian to early Campanian age are high-spired trichotro- Also included in this present study are new triclio-
pids. As presently known, Bullamirifica, Minijtropis, and tropid gastropods found in the collections at die Natural
Paxitropi.s were endemic to the stud\- area. Histon' Vluseiun of Los Angeles Counts'. Thev are
Mini/fropis melilota new genus and species from the
Santonian part ofthe Redding and Chico formations of
INTRODUCTION nordiern California, and Paxitropis dicriota, new genus
and species from die of upper Santonian part of the
Tills stud\- is based largeK' on specimens collected h\ Redding Formation, northern California; the lower
Eric Gohre of Oroville, California. Over the years, he ("ampanian part of the Chico Formation; and the lower
has amassed a sizeable collection of shallow-marine Campanian part ofthe upper Holz Member ofthe Ladd
moUusks from the lower Campanian Pentz Road Formation, southern California (Figure 1).
member of the Chico Formation near Pentz, Biitte The geologic age ofeach newspecies describt-d in this
Countv, northern Cahfomia (Figure 1). His collection paper is showai in Figure 2. The entire inter\'al oftime
has \ielded several new species ofgastropods, and some that encompasses all these species is Coniacian to late
of diese were described bv Groves (2004) and Squires Campanian, or about 19 million years. The new .species
and Saul (2004). In part, this present studv concerns are locally common, except for Tegula jcanac, Ncrita
three adchtional new species and a new genus of (subgenus?) orovillcnsis, and B. verruca.
gastropods found in his collection. They are die trochid The classification system used here generally follows
Tegula jeanae new species, die neritid Ncrita (sub- that of Hickman and McLean (1990) for the teguhnes.
genus?) orovillensis new species, and the ceritliioid Ponder (1988) for the triehotropids, and Ponder and
Bullamirifica clcgans new genus and species. Waren (1988) for the other taxa.
Inspection ofdie hteratru'e, as well as exanimation of Studv localities are listed in Appendix 1, Abbrexiations
the collections at the Natural Histoi"v Museinn of Los usedinthete.xtare: CAS: CaliforniaAcademyofSciences,
Page 1.34 THE NAUTILUS, Vol. 119, No. 4
1 -Redding Formation AGE(m.y.) 85 80 75 111
1 1 1 1 1 1 1 1 1 1 1 1 1 1 1
2-Cliico Formation UPPER CRETACEOUS
Coniacian Sant- Campanian
3-Pigeon Point
Formation lower ] middle ', upper
4-Jalama Formation cpholraornisty «— €34 I ^"" 1 ----- ^
jeanae
Tegula \ \
Nerita orovitlensis i
1 1
verruca
elegans
[
Bullamirifica
ainiktos
; :
' 1
Minytropis melilo
; 1
Paxitropis dicriota
P
1 1 1
Figure 2. Chronostratigraphic positions of the new taxa.
Ages of stage boundaries and magnetostratigraphy from
Gradstein etal. (2004, fig. 19.1).
and Saul (1997); Ten Mile Member of the Chico
Formation, Squires and Saul (2003b); upper Holz Shale
ofthe Laclcl Formation, Squires and Saul (2001); Pigeon
Point Formation, Elder and Saul (1993) and Squires and
Saul (2003b); and Jalama Formation, Squires and Saul
(2003b). The locales of these formations are showai in
Figm"e 1. Stratigraphic information mentioned below
concerns additional pertinent biostratigraphic details.
The age of the jalama Formation used here is slightly
younger than used in our previous papers because we
had to adjust its chronostratigraphic position based on
the latest pnblislied (Gradstein et al., 2004) absolute-
time and global-paleomagnetic data correlations.
Figure 1. Location offonnation.s hearing the new tixa.
Punta B.aja Formation
San Francisco; IGM: Mexico Museodel Paleontologiadel Perrilliat-Montoya (1968) reported specimens ofgastro-
Institutede Geologa; LACMIP: Natural Histoiy Museum pods, herein assigned to Bullamirifica ainiktos. from the
of Los Angeles County, Invertebrate Paleontolog\' "Rosaiio Formation" at Punta Baja, near El Rosario,
Section; UCLA: UniversiU' of California, Los Angeles northern Baja California, Mexico. The 5-140 m thick
(collections nowhoused at LACMIP); UCMP: University Punta Baja Formation (Figure 1) overlies fluvial deposits
ofCalifornia Museum ofPaleontology (Berkeley); LISGS: of the La Bocana Roja Formation, and the angular
United States Geological Survey. unconformit)' lietween these two formations is canyon-
shaped (Boehlke and Abbott, 1986). This canyon is filled
STRATIGRAPHY with conglomerate, sandstone, and siltstone reported by
Kilmer (1963) to have been deposited in shallow-marine
E.xcept for the Punta Baja Formation, which is discussed depths not exceeding 60 meters. Boehlke and Abbott
below, the ages and depositional environments ofall tlie (1986) have a differing viewpoint and reported that the
formations and members containing the new taxa deposits representturbidites thataccumulated in bathyal
discussed in this paper can be fovmd in the folkwing depths. Thev reported, furthermore, that shallow-marine
papers: Member IV ofthe Redding Formation, Squires mollusks are common, but Kilmer's collection at I'CMP
and Saul (2003a); Must)' Buck Member of the Chico does not contain \en' mam- specimens. The Puiita Baja
Formation, Saul and Squires (2003); Pentz Road Formation is iinconformabK' overlain by terrestrial
member (informal) of the Chico Formation, Squires deposits <il llir La Escarpa Member of die El Gallo
R. L. Squires and L. R. Saul. 2005 Page 135
Formation, wliifli, in turn, is oxcrlain 1)\ llic Rosai'in Subfamilv Tegulinae Kuroda. Habe and Ovama. 1971
Formation. (ienns Tetania Lesson. 1S35
SauBlase(1d98o3n:mo2l1li-i2s2t,vuifilgo.s.si9l)s turlelpcocrltceddIi\th!'e. IaI.mmKoilnmietre, Type .Species: Tcgula clegaiis Lesson. 18.35, by
Metaplaccnticeros cf. pacificiiin (Smith. 1900) and the monotypy; Recent, west coast of Central America to
the GulfofCalifornia, Mexico.
gastropod Tiirritclla chicoensis pescaclerocnsis Arnold.
1908, from the siltstone in the Punta Baja Formation. Di.scus.sion: Although Wenz (19.38). Keen (1960). and
Although diese t^\'o mollusks were reported in Saul l)a\ies (1971) reported the geologic range o( Tcgiila to
(1983: 65-66) to be oflate Campauian age. more reeent be Miocene to Recent, Baiidel and Stinnesbeck (2000)
biostratigraphie studies (Elder and Saul. 1996: fig. 1) reported a species of Tegtila of Late Cretaceous
depicted both of tliese ta.\a as ranging in age from late (Maastrichtian) age from central Cliile. Kiel and Bandel
middle Campauian to earliest late Campauian. Adju.st- (2001) reported a tentativelv identified Tcgula from
ments for the most recentlv published (Gradstein et al.. upper Campanian strata in northern Spain. The early
2004) absolute-time and global-paleomagnetic data Campanian new species described below represents the
correlations place these anuuouite and turritellid zones confirmed earliest record we knowoffor Tesiila. Forthe
in the middle late Campauian. Recent examination by Pacific slope of North America, the prexaous earliest
the junior author of additional Punta Baja Formation record of Tcgula was given bv Addicott (1973: 17. pi, S,
mollusks revealed three specimens ofthe bivalve Calva. figs. 2, 4), who reported it from the Wygal Sandstone
The best preser\'ed specimen is irom LACMIP loc. Member oi the Temblor Formation, southwestern
12582 and is Calva (Egclicalva) cras.sa Saul and margin of the San Joaquin Valley, Kern County, central
Popenoe. 1992. whose geologic range is earK late California, Squires'(2003: table 2.1, fig, 2.1) placed this
Campauian to earl\- Maastrichtian elsewhere on the member in the lower Oligocene Matlockian Stage.
Pacific slope of North America (Saul and Popenoe,
1992). The otlier two Calva specimens are worn and
broken, from UCMP loc. B-3388. These two specimens Tcgula jcaiKic new species
(Figures 3-5)
are similar to Calva {Calva) peninsularis (Anderson and
Hanna. 1935). whose geologic range is latest Campauian Diagnosi.s: A Tcgula with low to moderate spire.
to earl\- Maastrichtian elsewhere on the Paciiic slope ol Whorls convex, smooth, and bearing one spiral groove
North America (Saul and Popenoe. 1992). on posterior third of last whorl. Anomphalous. Last
Boehlke and Abbott (1986: fig. 4) assigned the age of whorl with raised lip along basal edge, base sunken
the Pmita Baja Formation to the early Campauian based between this lip and coliimellar lip. wliicli bears at least
entire1\- on calcareous nannofossils. They also reported one denticle and one much smaller denticle udapicalK-,
that the bentliic foraminifera in tliis formation corre-
spond to the F2-lower E foraminiferazones ofGoudkofl Description: Shell medium (up to 16,6 mm height
mm
(1945). but thevdid not relvon the foraminifei'a fortheir and 21.7 diameter, same specimen), Turliiniform
age call. .Almgren (1986: table 2) reported that die F2- with spire low to moderatel)' elevated, Protoconch
louer E foraminifera zones are essentially correlative to unknowTi, Teleoconcli consisting ofthree whorls. Suture
die early Campauian to late Campanian. It is important impressed, slightly channeled. Whorls convex, some-
to mention diat the Alcalde Shale in the Coaliuga area times slightly concave short distance anterior of suture;
along the west side of die San Joaquin Valley, central blunt angulation anterior of medial part of last whorl.
California, is correlative to the E zone (Almgreii, 1986: Whorls smooth; spiral groove posterior of medial part
table 3). As depicted in Saul (1983: fig. 10), die Alcalde of last whorl. Aperture oblique, peristome discontinu-
Shale contains Mctaplacenticeras cf M. pacificum, and ous, Anomphalous. umbilical area covered bv broad
.\lmgren (1986) assigned the Alcalde Shale to the early callus. Base wide and smooth, peripheral (abirxial)
late Campanian. margin coincident with raised lip along edge of last
In summary, die Metaploccnticeras Tunitvlla, Calva, whorl; area depressed between this lip and columella.
,
and benthic foraminifera data strongly support a middle Outerlip stronglvprosocline. Columellarlipwith at least
late Campanian age for the Punta Baja Formation. The one oblique denticle; much weaker second denticle
calcareous nannofossils. however, support an early sometimes present immediately posterior of main
Campanian age. It seems probable tluit the older denticle. Growth lines stronglv prosocline, formingwide
calcareous nannofossils are reworked, and this would bands,
be consistent with the depositional environment of the Holotype: LACMIP 13.322, 18,7 mm in height.
formation. „ nnii m diameter
SYSTEM.\T1C PALEONTOLOGY ParatyiJe: LACMIP 13323,
Type Locality: LACMIP loc, 24337
Superfaniilv Trochoidea Rafinesque, 1815
Faniilv Trochidae Rafinesque, 1815 Geologic Age: Early Campanian,
Paee 136 THE NAUTILUS, Vol. 1 19, No. 4
/i-::^
Figures 3-8. New tegulid and neritid gastropods. Specimens coated with ammonium chloride. 3-5. Tegulajeanae new species,
LACMIP loc, 24337. 3."Parat>iie LACMIP 13323, aperturalview, height 13.9 mm, diameter 17.9 mm. 4-5. Holotype 13322, height
18.7 mm, diameter 22 mm. 6-7. Nerita (subgenus?) orovillensis new species. 6. Holotype LACMIP 13324, apertural xaew, height
11.6 mm, diameter 16.8 mm. 7-8. Parat\pe LACMIP 13325, total shell height 10 mm, diameter 13.7 mm. 7. Aliapertural view. 8.
Apical view.
Distribution: Cliico Formation, Pentz Road iiioiiiber strata in northern Spain. The new species differs greatly
(informal), near Pentz, Butte County, northern Califor- from T.?simplex by having a less elevated spire, smoodi
nia. shell, blunt I'atherthan a shaqi angulation anteriorofthe
medial part of the last whorl, broad callus covering the
Etymology: Named for Jean Gohre, mother of Eric umbilical region, wider aperture, ven' much stronger
Ccilire, who collected and donated the tyjDe material to denticles on the columella, raised lip along the basal
LACMIP. edge of the last whorl, and sunken base between diis
raised lip and the eolnmellar lip.
Discussion: Tlii.s new species is known from two
specimens, both showint; good presei"vation. The new
Familv Neritidae Ratinesqne, 1815
species is remarkably similar to Tegula (Chloro.stoma) Genus Nerita Linnaeus, 1758
funebraUs (Adams, 1855), from Pliocene and Pleistocene
strata ofsouthern California (Grant and Gale, 1931) and Type Species: Nerita pelorouta Linnaeus, 1758, by
from the Recent ofVancouver Island, British Columbia, subse(juent designation (Montfort, 1810); Recent, soudi
to central Baja California, Mexico (McLean, 1978). The Florida, West Indies, and Bermuda.
new species differs from T. (C.) fiinebralis by having
Discussion: Nerita sensu lato ranges from Early
a smootli shell rather than being ornamented by weak
spiral libs. The similarity' between the new species and fCrreotmacetohues O(nHaouterMievmiabne),r aonld tthhee eBaurhdedsetnreCcaorndyoins
T. (C)fiinebralis is even strongerifthe specimens oftlie Formation, Trinity County, northern California (Saul
latter are worn. and Scpires, 1997). The new species described below
Tegula ovallei (Philippi, 1887: pi. 12, fig. 4; Bandel represents the first record of an early Campanian Nerita
and Stinnesbeck, 2000: 761, pi. 1, B), the only other Irom tlie stndv area.
positively identified Cretaceous Tegula that we know of
is from Maastrichtian strata in central Chile. The new
Subgenus?
species differs from T. ovallei by having a smooth shell
Nerita (subgenus?) orovillensis new species
rather than biiiig ornamented by granulated spiral
(Figures 6-8)
ridges.
Kiel and Bandel (2001: 139, pi. 1, fig. 1) reported Diagnosis: A Nerila wilh appiiixiniately 18 to 19
a tentatively identified Tegula? simplex (Qnintero and beatled spiral ribs. (Johmicl ar hp with l(ini" or five
Revilla, 1966: 49, pi. 8, fig. 3) from upper Campanian obscure teeth.
R. L. Squires and L. R. Saul. 2005 Page 137
Description: Shell medium small (up to 11.6 mm in beads that are not elongate, wider interspaces between
height and 16.3 mm in diameter, same specimen), the ribs, and fewer, stronger, and wider teeth on the
broader than high, globose. Last whorl rapidK' e.xpand- colinnellar lip.
ing. Protoconch unknovni. Teleoconch consisting o( 2.5 The only other e;Lrly Campanian neritid known from
to 2.75 whorls. Uppermost spire \'eiy low. Suture the Pacific slope of North America is Neritina (Dostia)
obscure. Earliest 1.5 teleoconch whorls apparently cnneata (Gabb, 1864: 137, pi. 21, fig. 97) from lower
smoodi, rest of teleoconch covered with appro>dniately Campanian strata atTuscan Springs on Little Salt Creek,
IS narrow spiral ribs bearing small beads; interspaces Tehanui County, northern California. Gabb's species
between ribs approximateK' as wide as interspaces. might also be present in 1) upper Campanian and/or
Beads on ribs becoming smalk'r and slightly elongate lower Maastrichtian strata in the Pozo area, San Luis
on base oilast whorl, especialK' in parietal region. Spiral Obispo County (Vedder, 1977) and 2) Maastrichtian
rib adjacent to suture can be slightK' stronger than other strata along tlie western edge ofdie San Joaquin Valley,
ribs. Aperture large, nearly circular. Outer lip flared, California (Woods and Saul, 1986). The new species is
interior smooth. Columellar lip with tiye somewhat vastlydifferent from Neritina (Dostia) and does not have
obscure teetli, most posterior tooth widest and longest. its patelliform shape nor its distinctive collabral sculp-
Deck area broad, sloping, and shaipK' demarcated from ture.
base of last whorl. Growth lines prosocline.
HoIoUpe: L.^CMIP 13324, 11.6 mm ni height, Superfamily Cerithioidea Fenissac, 1819
mm Family Indeterminate
16.S in diameter.
Paratope: LACMIP 13325. Discussion: The new genus described below is most
likely a cerithioid, on the basis of its sigmoidal growth
T^pe LocaliK: L.\CMIP loc. 24337. lines, high spire, sculpture, short siphonal canal (shghtly
twisted), smooth columella, and smooth interior of die
Geologic Age: Early Campanian.
outer lip. Some specimens of the new genus have
Di,sti-ibution: Chico Formation, Pentz Road member a narrow spire, like that found in ceritiiioids, but other
unft)rmal), near Pentz, Butte Count)'", noitliern CaUfor- specimens of the new genus have a buccinid-like shell.
nia. The strongly sigmoidal growth hues of the new genus,
however, are unlike that found on buccinid shells. It is
EUTiiology: Named for Oroville, Calilornia. possible that the new genus belongs to a new ceridiioid
Discu,s,sion: The new species is based on two speci- family.
mens. The external surfaces are moderately well pre-
ser^-ed, but the columellar lip and especially tlie deck Genus Bulhiinirifiea new genus
area are poorly presei"yed.
Type Species: Bidlamirifica elegans, new species;
The new species is remarkably similar to Nerila Early Camp;uiian, Pentz area, Butte Coimt\', northern
(Thelioshjla) crooki Clark (1938: 700, pi. 4, figs. 1, 2)
California.
from die Markley Formation east of San Francisco,
Solano County, northern Cahfornia. Squires (2003: table Description: Shell medium (up to 83 mm heiglit and
2.1. fig. 2.1) assigned this formation to the middle 37 nmi diameter, same specimen), fusiform to tini'eted.
Eocene ("Tejon Stage"). The newspecies differs from N. Height to diameter ratio 2 to 2.7. Spire high, comprising
(T.) crooki by having fewer and wider teeth on the 41 to 55% oftotal shell height. Pleural angle 33 to 42°.
columellarhp, fewerribs on the lastwhorlwith relatiyely Protoconch imknown. Teleoconch whorls six to eight.
wider interspaces, and ribs near die middle of the last Spire whorls with shoulder angulate; last whorl with
whorl not noticeably broader than adjacent ribs. peiiphery' angulate. Ramp short to moderately long,
The new species is also very similar to Nerita concave to rarely straight-sloped. Suture slightly un-
umzamhiensis Woods (1906: 311, pi. 37, figs. 14-15; dulatoiy, weakly impressed. Collabral sculpture consist-
Bandel and Kiel, 2003: 51-52, pi. 1, figs. 4-5) from ing of many narrow ribs, closely to moderately widely
die Santonian/Campanian Umzamba Formation in spaced; interspaces smooth. Collabral ribs slightly
southeastern Soutli Africa. The new species differs opisthocline to opisthocyit, usually extending from
from A", umzamhiensis by having fewer teeth on the suture to suture. Collabral ribs present on base or
columella Hp, ribs on the base of the last whorl, and obsolete; if present, swollen and elongate. Spiral
a deck area sharply demarcated from the base ofthe last sculpture consisting of several .spiral ribs with variable
whorl. width and spacing, especially on last whorl. Spire whorls
The new species somewhat resembles Nerita (The- with strongest spiral rib on whorl shoulder, severalweak
Uostijh':') kennediji Squires and Saul (2002: 185-187, or moderately strong spiral ribs occasionally near
figs. 31-34) from tiie upper lower to lower middle anterior suture, and suture coincident widi weak spiral
Eocene ("Domengine Stage") Santiago Formation, rib either bearing weak nodes or without nodes. Last
northern San Diego County, southern California. The whorl sculpture with three to four widely spaced, strong
new species differs from A'. (T.?) kennediji by having spiral ribs on peripheiyand one ortwoweakerspiral ribs
.
Pase 138 THE NAUTILUS, Vol. 119, No. 4
or se\eral .spiral riblet.s on base. IntersectitMis ot collabral Pficitdmapa Holzaptel. 1888, a mouot\'pic genus from
and spiral ribs producing many nodes or strongly the Vaals Greensand in the Nedierlands and Germany.
projecting tubercles, either rounded (knob-like) to The age of these strata was determined to be early
spiiiose or narrowly elongate. Nodes and knobs most Campanian by Albers (1976). Wenz (1941: 1083,
pronounced on whorl shoulder, especially on last whorl. fig. 3079) illustrated Pseudorapa. Btdlamirifica differs
Intersections also strongon anteriorportion oflastwhorl from it by having a less twisted siphonal canal and
periphery. Aperture short but moderatelywide, comma- more variable sculpture, including the possibilities of
shaped; small, narrow arch (canal-like) present where having spiral ribs and a relatively narrow shell. In
outer lip meets most posterior pait of aperture. addition, Pseudorapa has an outer lip that is very
Columellar lip smooth. Siphonal canal short and spout- crenulate and a spiral band near the base of the last
like or well developed, moderately short, and can be whorl that pi-oduccs a tooth-like projection on tlie outer
twisted to left. Outer lip thin, markedly sinuous, interior lip.
smooth. Growth lines sigmoidal between suture and
shoulder; antispiral sinus coincident with tuberculate Bullamirifica vernica new species
spiral rib. (Figures 9-12)
Geologic Age: Coniacian to earlv late Cainpanian. Diagnosis: Small Bultaiiiirifica with prominent round-
Etymology: Combination of the Latin bulla, mean- ed knobs, nine on shoulder of last whorl. Suture
ing knob, and the Latin iiiirifiis, meaning to cause coincident with noded spiral rib. Base of last whorl
wonder. without elongate collabral ridges. Siphonal canal short
and straight.
Discussion: Three species can be herein assigned to
mm
this new genus. Two of these species, Biillamirifica Description: Small (up to 34 estimated height
vernica and Biillamirifica clegans, are based on entirely and 16.8 rrun diameter, same specimen), fusiform,
new material. The third species was originallytentatively moderatelv slender. Height to diameter ratio appro.xi-
assigned by Dailey and Popenoe (1966) to Pscudoglaii- mately 2. Spire high, approximately 44% ofshell height.
conia Douville, 1921. Dailey and Popenoe (1966) stated Pleiu'al angle approximately 35 to 37°. Protoconch and
that tliis particular species belongs in a new genus, but upper spire unknown. Teleoconch v^'horls approximately
they withheld dieir description until better specimens six (estimated). Spire whorls widi shoulder angulate, last
were obtained. Although representatives of the new whorl with peripheiy angulate. Ramp short, concave.
genus have sigmoidal growth lines (see Wen/. 1940: 764, Suture slightK- impressed, possiblv undulaton-. Collabral
fig. 2214) similar to that of Pscudoglaiiconia. Bullamir- sculpture consisting of man\' ribs, widcK' spaced;
ijicd differs considerablv from Pscndoglauconia hv interspaces smooth. Collabral ribs slightK opisthochne
having a bucciniform rather than a tapered/conical and extending from suture to suture. Collabral ribs
shape, angulate rather than flat-sided whorls, and mainly prevalent on ramp. Spiral sculpture consisting of
tubercles on die sides of die whorls instead ofonly near several ribs with variable strength and spacing, especially
the suture, and tlie suture between the penultimate and on last whorl. Intersections of collabral and spiral ribs
last whorl is not extremely deep and widcl\- sunken. pj-oducingmanystronglyprojecting roundedtuberculate
Unfortimatelv, the aperture of Pscudoglaucoma is not knobs or, less commonlv, nodes; knobs and nodes most
kno\\^l and none ol the apertures on the available prominent on shoulder and usually extendingposteriorK*
specimens of Bulhiinirificii is complete. In spite of the across ramp and become narrow lidges. Spire whorls
absence of knov^'ledge about the aperture of Pseudo- with strongest spiral lib on whorl shoulder, very faint
glauconia, Wenz (1940) believed this gastropod genus to spual liblets present between whorl shoulder and
belong in the Cerithiidae. anterior suture. Sutiu"e coincidentwith moderatelyweak
The shape of the growth lines in Biillninirifica is spiral rib bearing small nodes. Penultimate whorl widi
similar to that ot Batillaria cchinaidis clavatulata spiral rib on shoulder bearing nine, moderately closely
(Lamarck, 1804) from the middle Eocene (Lutetian) of spaced knobs. Last whorl with three spiral ribs on
the Paris Basin, France. The siphonal canal of this periphen, strength of ribs progressively decreasing
Eocene species is longer than normally found in anteriorly: strong rib on shoulder and bearing nine veiy
Batillaria Benson, 1842. Ponder and VVaren (1988) and prominent tuberculate knobs; middle lib moderately
Iloubrick (1988) placed Batillaria in superfamily Cer- strong and bearing more numerous nodes (about half
ithioidea, family Batillariidae Thiele, 1929. The similarity sized ofthose on shoulder) that tend to become smaller
in growtli-hne shape between Bullamirifica and Batil- and even obsolete adaxiallv; and most anterior peripheiy
laria suggests to us that the new genus might be rib weakest and weak nodes tending to be obsolete
a cerithioid. The high turreted spire of Batillaria, adaxiallv. Base of last whorl usuallv with one or tvvo
however, is quite unlike the lower, more paucispiral spiral ribs, both bearing nodes (best developed near
spire o{Bullamirifica outer lip) or bearing no nodes, and with strengdi of
Bullamirifica has the growth-line shape, shell shape, spiral ribs decreasing anterior in direction; occasionally,
sculpture, and twisted siphonal canal similar to that of anterior region ofbase with onlyveryweak spiral riblets.
R. L. Squires and L. R. Saul. 2005 Pao-e 139
'^W'
Figures9-21. Newcerithioid?gastropods. Specimenscoatedwitliammoniumchloride. 9-12. Biilhnnirificavcrnica newgenusand
species.LACMIPloc.8133.9.ParatopeLACMIP 13327.aperturalview,heiglit27 mm,diameter19.2 mm.'10-11. HoIotvpe^LACMIP
13326. height 29.7 mm. diameter 17.8 nnii. 10. Apertural \iew. 11. Abapertural view, 12. Paratvpe LACMIP 13327. basal view,
diameter 18.9 mm. 13-17. Bidlamirifica elegans new genus and species. LAfJMIP loc. 24337. 13-14. Holotvpe LACMIP 13328,
height 66.9 mm, diameter 34.9 mm. 13. Apertural view. 14. Abapertiu-al view. 15-16. Paratype LACMIP 13329, height 60.1mm.
chameter25.7 mm. 15. Apertural view. 16. Abapertural view. 17. Holotyise LACMIP 13328, basal view, diameter 35.7 mm. 18-21.
BnUamirificaainiktos (DailevandPopenoe. 1966)newgenus. 18-19. Hvpotvpe LACMIP 13330. LACMIPloc. 10691. height60 mm,
diameter24.1 mm. 18. .-\pertnralview, 19. Abapertiu'alview. 20. Plasto-hulntvpe LACMIP40435. LACMIP loc. 24125. right-lateral
view, height58.2 mm, diameter29,1 nun, 21. HvptoUpe LACMIP 13331, LACMIP loc. 24124, basal view, diameter 16.7 mm.
Page 140 THE NAUTILUS, \oI. 119, No. 4
Aperture round with veiy small posterior "arch." Collabral ribs sigmoidal between posterior suture and
Aperture eUiptical, cokimellar lip smooth, outer lip thin; shoulder, opisthocline between shoulder and anterior
siphonal canal short and spout-like. suture. Collabral ribs somewhat swollen on base of last
Holotype: LACMIP 13326, incomplete specimen whorl. Spiral sculpture consisting of several ribs with
mm variable strength and spacing. Intersections of collabral
with mtwmo whorls (upper spire missing), 29.7 lieight, and spiral ribs producing many swollen, axially elongate
17.8 diameter.
nodes or, in some cases, rounded tuberculate knobs;
Paratype: LACMIP 13327. nodes and knobs most prominent on shoulder. Spire
whorls with strongest spiral rib on whorl shoulder and
Type Locality: LACMIP loc. 8133. occasionalK' two moderateK' strong, noded spiral ribs
Geologic Age: Coniacian. between whorl shoulder and anterior suture. Suture
coincident with weak spiral riblet. Penultimate whorl
Distribution: Redding Formation, Member I\', Oak with spiral rib on shoulder bearing 11 widely spaced
Run ai'ca, northern C^alilornia. nodes. Last whorl with three or four spiral ribs on
peripher)', strongest rib on shoulder and bearing 11
EtMnology: Latin icrnicus. wart.
nodes. Spiral ribs on peripheiyoflastwhorl usuallyshow
Discussion: This new species is based on three strength progressively decreasing anteriorly, occasionally
specimens. It differs Irom Bullaiiiiiifica elcgans new rib immediately anterior ofshoulder weaker tlian other
species below by smaller size and presence of rounded rilis on peripheiy. Growth lines sigmoidal, with antispiral
rather than elongate knobs, fewer knobs on shoulder of sinus coincident M.ith shoulder. Aperture elhptical,
last whorl, no elongate collabral ridges on base of last columellar lip smooth, outer lip thin; siphonal canal
whorl, a noded spiral rib coincident with the suture, and short but well developed and slightl)- twisted to left.
a straight siphonal canal. BiiUamirifica vcniica differs Holotype: LACMIP 13328; nearlvcomplete specimen
from BiiUamirifica ainiktos by being smaller wdth mm
a shortei" spire and ha\ing rounded and much more of five whorls (upper spire missing), 66.9 height,
34.9 nnn diameter.
projecting nodes, many fewer nodes on shoulder oflast
whorl, much less tendency for elongate collabral ridges Paratype: LACMIP 13329.
on ramp, and muchweaker spiral ribs between shoulder
and anterior suture. Type Locality: LACMIP loc. 24337.
Bullamirifica vcrnica is \'en' similar to tlu' ceiithioid Geologic Age: EarK' Campanian to possiblv middle
Ti/mpanoloniis {Tipiii>aii(ifonns) robtistus Dockeiy Campanian.
(1993: 47, pi. 7, fig. 1) in the shape of the spire, strong
nodes on the spire, and shape ofthe growth lines next to Distribution: Earlv Campanian: Chico Formation,
the outer lip. Tijmpanotonus (T.) robustiis, which is of Pentz Road member (informal), near Pentz, Butte
Campanian age and from Mississippi, however, has no County, northern California; Possibly middle Campa-
siphonal canal. nian: Pigeon Point Formation, San Mateo County,
nordiern Calilornia.
Bullaininfica dedans new species Etymology: Latin clegans. meaning veiy fine or
(Figures 13-17) choice.
[?] Pscudoghiuci^nia? aff P. ainikfos Daile\- and Pope- Discussion: The new species is based on six speci-
noe.—Elder and Saul, 1993: pi. 2, fig. 11. mens from the Pentz area, and presenation is moder-
Diagnosis: Large BuUiiDiirifica with prominent ately good on all of them. Nearly all the specimens are
opisthocline collabral ridges on upper spire and base moderateK'wide, but a few are somewhat slender, as tlie
specimen illustrated in Figures 15-16.
of whorl. Shoulder of last whorl with 11 nodes. Suture
coincident witli unnoded weak spiral rib. Siphonal canal There might be one specimen of the new species from
the Pigeon Point Formation. This museum specimen,
short but well developed and twisted to left.
which cannot be located, was figured by Elder and Saul
Description: Medinin large (up to 83 mm estimated (1993: pi. 2, fig. 11). It was"identified by them as
mm
height and 37 diameter, same specimen), fusiform, Pseiidoglaiiconia? sp. aff P. ainiktos Dailey and Pope-
moderately wide, rarelv slender. Height to diameter noe, but it has a profile somewhat more like Bullamir-
ratio approximatelv 2.2. Spire high, approximatelx 40% ifica elcgans and basal nodes like B. clegaus. As will be
of shell height. Pleural angle approximately 33 to 42", discussed later, there are specimens of B. ainiktos from
rarely approximately 30°. Protoconch and uppermost the Pigeon Point Formation. It seems likelv that some of
spire unknown. Teleoconch whorls approximatelv eight the Bullamirifica specimens from the Pigeon Point
(estimated). Whorls with angulate shoulder. Ramp short Formation represent transitional forms between B.
and slightly concave. Collabral sculpture consisting of elegans and B. ainiktos.
many moderately strong, narrow ribs; interspaces Bullamirifica elcgans differs from Bullamirifica ver-
smooth. Collabral ribs extending from suture to suture. ruca in having larger size, prominent collabral ribs
—
R. L. Squires and L. R. S;i 200.5 Page 141
extending from suture to suture (iu the spire whorls, shoulder and live moderately strong, non-noded spiral
swollen et)llabral lihs on llic neek. more nodes on ribs betxveen slioulder and anterior suture; most anterior
shoulder ol the penultimate and last whorls, and one ol these ribs coincident with suture. Penultimate
siphonal canal longer and slightK twisted to the left. In and last whorls witli appro.ximately 14 nodes on
addition, B. clegans occasionalK has strong unnoded shoulder. Last whorl with three spiral ribs on periphen',
spiral ribs het\\'een suture and shonkk-r. progressixeK'weaker anteriorly, and bearing nodes. Base
Bulldiuirifica ch'<^(iiis diiiers from Biilldiiiiiificd aiiiik- ol last whorl bearing fine spiral riblets. Siphonal canal
tos bv usnalK- ha\ing a wider pleural angle, usually a less short and twisted to left. Columellar lip smoolh. Outer
elexated spire, wider and stronger nodes on shoulder, lip sinuous; interior smooth.
Bfsiweiowllell/rneunniroidfceiocslalaocblnreaglsahnosruilbadslseorihnoasfstel\aaasdrtiawbohifloirtlf\,inieannmdsopiibrapashleolwroiigbtswh. oHfolsoptiyrpee:andLsAiCphMoInaPl4c0a4n3a5l, iminscsoimnpgl)e,te39spmemcimheenig(htti,p
29 nun diameter.
whereas B. ainiktos does not.
Btilhiiiuiifica clegans is xen- similar to the cerithioid Type Locality: LACMIP loc. 4125.
Tt/mpanotonus {Tt/mpanotoniis) binodosns Docken*
(1993: 47, pi. 7, fig. 2) in the shape ofthe spire, strong Geologic Age: Middle Campanian to Late Campa-
nodes on the spire, and shape ofthe growth lines next to nian.
the outer lip. Ti/mpanotonns [T.) hinoclosns of Campa- Distribution: Middle Campanian: Pigeon Point For-
nian age and from Mississippi, howcxcr, has no siphonal mation, near Pigeon Point, San Mateo Count\', northern
canal. ("alifornia. Middle late Campanian: Punta Baja Forma-
tion, southwest of El Rosario, Baja Cahfornia, Mexico.
Bnllamirifiea ainiktos (Dailex and Popenoe, 1966) new Late Campanian: [alama Formation, Santa Barbara
combination County, southern California.
(Figures 18-25)
Discussion: Description ol the new species is based
Pscucloghuiconia? ainiktos Dailey and Popenoe, 1966: on 25 specimens: 18 Irom the jalama Formation, five
21-22, pi. 6, figs. 3, 5. 6. from the Pigeon Point Formation, and \^\'o from the
Pscndoglauconia? aff P. ainiktos Dailev and Popenoe. Punta Baja Formation. The Jalama specimens ha\'e poor
Elder and Saul, 1993: pi. 2, fig. 10. — to moderately good presenation, although the apices are
Cimolithiiun mit/akocnse (Nagao, 1934). Perrilliat- broken off and the siphonal canal missing. The Pigeon
Monto\a, 1968: 20, pi. 4, fig. 2. Point specimens also have poor to moderateK' good
—
Vicanja (Shoshiroia) tjalwi Kaniada. 1960. Perrilliat- presei-vation, but the matilx is veiy hard to remove. The
Montova, 1968: 21. pi. 6. fig. 1. Punta Baja specimens have poor preservation.
The Pigeon Point Formation specimen illustrated by
sDlieangdneorsitsu:rretMeeddiwuhmo-rlssi.zeRaBmiipllaiointiriufpipcaerwistphireussluiaolrtK, lEilkdeerthaatndfoSuanudl (o1n99B3.: apii.ni2k,tofsig.bu1t0)itlihaassawwihdoerrlpplreoufrialel
with narrow collabral ribs crossed h\ weaker spiral ribs; angle, similar to that found on B. elcgans. Unfortunately
ramp on later whorls much longer and smooth and
this specimen is missing. As mentioned earlier, it seems
conca\'e. Spire whorls concave on posterior half but
likely that some ofthe BuUamirifica specimens from the
angulate medialK. with moderately strong tubercles.
Pigeon Point Formation represent transitional forms
Shoulder oflast whorl with 14 nodes. Base oflast whorl between B. elcgans and /x ainiktos.
with man\', closeK' spaced spiral riblets. Siphonal canal
BuUamirifica ainiktos differs from BuUamirifica ver-
short and possibK straight. nica In larger size and having less roimded and much
Description: Mediurn, up to 67.5 mm height estimat- lower nodes, more nodes on shoulder of last whorl,
ed and 25.4 mm diameter, same specimen. Shell usualK much more tendency for elongate collabral ridges on
slender, turreted, occasionally wide, fusiform. Height to ramp, and much stronger spiral ribs between shoulder
diameter ratio approximately 2.7. Spire high, appro.xi- and anterior sutvu'e.
mateK' 55% of total shell height. Pleural angle approx- BuUamirifica ainiktos tlillers from BiiUiiiiiirifica elc-
imately 33 to 37". Protoconch unknowai. Teleoconch gans by smaller size, ha\ing a narrower spii'e, usually
whorls appro.ximately eight (estimated). Spire whorls a more elevated spire, narrower and weaker nodes on
with shoulderangulate. Ramp usualK'short and concave, shoulder, more nodes on shoulderoflastwhorl, andbase
occasionalK- long and straight-sloped. Suture obscured. with fine ribs instead ofswollen collabral ribs.
Collabral sculpture consisting ot many spiral ribs closely Perrilliat-Montoya (1968) identified one of the Pmita
spaced and narrow. Collabral ribs opisthocyrt, most Baja Formation specimens (Figures 23-25 herein) ot B.
prominent on ramp. Spiral sculpture consisting of ainiktos as Vicanja (Shoshiroa) i/abci and identified die
se\eral ribs, moderately weak and closely spaced. other specimen as Cimolithiiim miijakoense. Witli Wenz
Intersections of collabral and spiral ribs producing mam' (1940: 740-741, fig. 2145) as a basis for comparison, we
nodes, most prominent and somewhat spinose on consider that BuUamirifica ainiktos diflers considerably
shoulder. Spire whorls with strongest spiral rib on wliorl from the formergastropod by ha\ing a non-conical shell.
Page 142 THE NAUTILUS, Vol. 119, No. 4
Figures 22-34. Newceritliioid?andtiichotropid gastropods. Specimens coatedwithammoniinn chloride, nnlessothei^wisenoted.
22-25. BuUamihfica ainikfos (Daile\- and Popenoe'. 1966) new genus. 22. H)i5ot)'pe LACMIP 1.33.32. USGS loc. M-S601, height
.35.8 mm, diameter 20 mm. 23-25. Plasto-h\pot}pe IGM 1325, height 80 mm, diameter 36.9 mm. 23. .4pertnral view. 24.
Abapertural view. 25. Basal view. 26-29. Minijfropis mclilota new genus and species. 26. Parahpe LACMIP 13334, LACMIP loc.
10787, apertural view, height 7 mm, diameter 2 mm. 27-29. Holot)pe LACMIP 13333, LACMIP loc. 10786, abapertural view,
height 62 mm, diameter 2.2 mm. 27. Abapertural view. 28. Oblitjue apertural view oftip; arrow indicates where protoconch ends
and teleoconch starts. 29. Pi'otoconcli, height 0.5 mm, diameter 0.8 mm; arrow indicates where protoconch ends and teleoconch
starts; SEM pliotomicrograph (uncoated). 30-34. Paxitmpis dicnota new genus and species. 30-31. P;nahpe LACMIP 13336,
LACMIP loc. 2.3643, height 12.7 mm, diameter 6.8 mm. 30. Apertural view"31. Abapertural view. 32. llolotvpe L.A.CMIP 1.3335,
LACMIP loc. 23639, apertural view, height 14.7 mm, diameter 6.6 mm. 33. Paratope LACMIP 13337, LACMIP loc. 23642,
apertural view, height 10.1 mm, diameter 4.4 mm. 34. Paratype LACMIP 13338, LACMIP loc. 24217, abapertural view, height
1 1 nun, diameter 4.3 mm.