New discoveries of amphibiens and reptiles from Vietnam

2010·5.7 MB·English

by  Thomas Ziegler

Bonn zoological Bulletin Volume 57 Issue 2 pp. 137-147 Bonn, November 2010
New discoveries of amphibians and reptiles from Vietnam
Thomas Ziegler1 & Truong Quang Nguyen2-3
1 Cologne Zoo, Riehler Strafie 173, D-50735 Cologne, Germany; E-mail: [email protected]
institute ofEcology and Biological Resources, 18 Hoang Quoc Viet, Hanoi, Vietnam;
E-mail: [email protected]
3 Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D-53113 Bonn, Germany
Abstract. We provide alistof21 newamphibian and reptilian species andsubspecies discoveries fromVietnam, includ-
ing one new snake genus, published after the comprehensive overview by Nguyen et al. (2009). The new herpetofauna
representatives are introduced inclusive ofthe original description, type locality, English andVietnamese names, aswell
as current distribution.
Key words. Vietnam, herpetofauna, new species.
INTRODUCTION
AlthoughVietnam has one ofthe world's richest amphib- ofIndochinese and specifically on Vietnamese herpetol-
ian and reptilian fauna, as revealed particularly through ogy. Besides the publication ofseveral identificationman-
surveys by Vietnamese scientists and their international uals, he is most famous for a substantial series ofmono-
collaborators during the past quarter century, the study of graphs ofIndochinese herpetology, which are to date the
its herpetofauna was long overshadowed by research in most importantbackground on the subject (Bourret 1936a,
India, China, and the East Indies (Adler 2009). The first b, 1941, 1942). In total, Rene Bourret reported of 177
significant summary oftheVietnamese herpetofaunawas lizard taxa (i.e., species and subspecies), 245 snake taxa,
writtenby Morice (1875), inwhich 13 amphibians and 114 45 turtle taxa and 171 amphibian taxa forthe Indochinese
species ofreptiles including the marine species were list- region (Nguyen 2006).
ed. Ten years later, Tirant (1885) published a 104-pages
book containing 166 species ofherpetofauna from Viet- Because ofthe French IndochinaWar, no remarkable her-
nam and Cambodia. Subsequently, the following staff petological studies were undertaken in the period from
members or associates of the Museum d'Histoire Na- 1946 to 1954. In 1954, when northern Vietnam attained
turelle, Paris, published specific studies ofVietnamese am- independence from France, Vietnameseherpetologists be-
phibians andreptiles: LeonVaillant (1834-1914), Francois gan to conduct herpetological field surveys predominant-
Mocquard (1834-1917), Jacques Pellegrin (1873-1944), ly in northern Vietnam, and the first lists and keys to the
Paul Chabanaud (1876-1959), Fernand Angel species ofVietnamese amphibians andreptiles were com-
(1881-1950), and Rene Bourret (1884-1957), who was piledby Dao (e.g., 1977, 1978, 1979, 1981, 1982), includ-
the only one ofthe afore mentioned zoologists who ever ing 363 species in total, but with inaccuracies. Another
set foot in Vietnam (Adler 2009). noteworthy key to snakes ofsouthern Vietnam was writ-
tenduring the Second IndochinaWar, betterknown inthe
It was Rene Bourret, originally a geologist, who became West asVietnamWar, by Campden-Main (1970), who had
the leading authority on Vietnamese herpetofauna. From served several years as amedic withAmerican forces sta-
1927 until 1947 he published a series ofpapers andbooks tioned in Vietnam.
Bonn zoological Bulletin 57 (2): 137-147 ©ZFMK
138 Thomas Ziegler & Truong Quang Nguyen
The end ofthe Vietnam War in 1975 marked the begin- LIST OFNEW SPECIESAND SUBSPECIES SINCE
ning ofanotherperiod ofbiodiversity research inVietnam NGUYEN ET AL. (2009)
(Sterling et al. 2006). Thereafter, an increasing engage-
ment in herpetological field work mainly by Russian in- Amphibia
stitutions was observable, which is still persistent. Even-
tually, in the last 30 years many international herpetolog- Anura
ical cooperations emerged, which led to an enormous in-
crease in newrecords and species descriptions from Viet- Megophryidae
nam (see overview in Nguyen 2006). As a consequence,
&
the updated checklist by Nguyen et al. (2005) comprised Leptolalax applebyi Rowley Cao, 2009
458 species, viz., 162 species of amphibians and 296
species ofreptiles, which included more than 100 addi- LeptolalaxapplebyiJ.J.L. Rowley & T.T. Cao, 2009, Zoo-
tional species comparedwith the previous checklist ofthe taxa 2198: 52.
herpetofauna ofVietnam by Nguyen & Ho (1996). Fur-
AMS
thermore, the most actual list (Nguyen et al. 2009) cov- Holotype: R171703.
ers 12 additional species ofamphibians and 64 addition-
al species ofreptiles compared withNguyen et al. (2005). Type locality: Song Thanh Proposed Nature Reserve,
Phouc Son (Phuoc Son) District, Quang Nam Province,
On the occasion ofthis Proceedings Volume addicted to Vietnam, 1,402 m a.s.l.
Professor Dr. Wolfgang Bohme, Vice Director, Head of
the Vertebrate Department and Curator ofHerpetology at English name: Appleby's Asian Toad.
the Zoological Research Museum Alexander Koenig,
Bonn, Germany, we provide a list ofnew amphibian and Vietnamese name: Coc may ap-li-bai.
reptilian discoveries from Vietnam that were published
subsequent to the comprehensive overview provided by Distribution: This species is currently known only from
Nguyen et al. (2009). We would like to dedicate this pa- the type locality.
per to Wolfgang Bohme, who supervised the PhD theses
of both authors (T. Ziegler: 1997-2000; T.Q. Nguyen
2007-2011) andthus decisivelybrought forwardherpeto- Ranidae
diversity research in Vietnam.
Odorranageminata Bain, Stuart, Nguyen, Che & Rao,
2009
MATERIALAND METHODS
Odorranageminata R.H. Bain, B.L. Stuart, T.Q. Nguyen,
We herein compiled species' descriptions that were for- J. Che & D.-Q. Rao, 2009, Copeia 2: 355.
mally published after the appearance of Nguyen et al.
(2009). We therefore followed the style and taxonomic Holotype: AMNH 163782.
arrangement provided by the latter authors.
Type locality: MountTay Con LinhII, Cao Bo Commune,
Abbreviations are as follows: AMNH =American Muse- Vi Xuyen District, Ha Giang Province, Vietnam, 1,420 m
um ofNatural History, NewYork, USA;AMS =Australian a.s.l.
Museum, Sydney, Australia; IEBR = Institute of Ecolo-
gy and Biological Resources, Hanoi, Vietnam; ITBCZ = English name: Geminated Cascade Frog.
Institute ofTropical Biology, Collection ofZoology, Ho
Chi Minh City, Vietnam; LSUHC = La Sierra Universi- Vietnamese name: Ech bam da hoa.
ty, Herpetological Collection, La Sierra University, River-
side, California, USA; UNS = Zoological Collection of Distribution: This species (Fig. la) is currentlyknown on-
the University of Natural Sciences, Ho Chi Minh City, ly from montane areas in northeastern Vietnam (Ha Gi-
Vietnam; VNMN =VietnamNational Museum ofNature, ang and Cao Bang provinces) and southeastern Yunnan
Hanoi, Vietnam; ZFMK = Zoologisches Forschungsmu- Province, China.
seum Alexander Koenig, Bonn, Germany; ZISP = Zoo-
logical Institute, St. Petersburg, Russia; a.s.l. = above sea
level.
Bonn zoological Bulletin 57 (2): 137-147 ©ZFMK
New amphibians and reptiles from Vietnam 139
Fig. 1. a) Odorranageminata from Ha Giang Province, Photo T.Q. Nguyen; b) Theloderma lateriticum from Lao Cai Province,
Photo T.Q. Nguyen; c) Leiolepis ngovantrii from Ba Ria-Vung Tau Province, Photo L.L. Grismer; and d) Pseudocalotes ziegleri
from Kon Turn Province, Photo C.T. Ho.
Rhacophoridae Reptilia
Theloderma lateriticum Bain, Nguyen & Doan, 2009 Squamata
Theloderma lateriticum R.H. Bain, T.Q. Nguyen & K.V. Sauria
Doan, 2009, Zootaxa 2191: 60.
Agamidae
AMNH
Holotype: 168757/IEBR A. 0860.
&
Leiolepis ngovantrii Grismer Grismer, 2010
Type locality: NamTha Commune, VanBan District, Lao
m &
Cai Province, Vietnam, 1,300-1,400 a.s.l. Leiolepis ngovantrii J.L. Grismer L.L. Grismer, 2010,
Zootaxa 2433: 52.
English name: Brick-red Bug-eyed Frog.
LSUHC
Holotype: 9234.
Vietnamese name: Ech cay san do.
Type locality: Binh Chau-Phuoc Buu Nature Reserve,
Distribution: This species (Fig. lb) is currently known on- Xuyen Moc District, BaRia-VungTauProvince,Vietnam,
m
ly from the type locality. 30 a.s.l.
Bonn zoological Bulletin 57 (2): 137-147 ©ZFMK
140 Thomas Ziegler & Truong Quang Nguyen
Fig. 2. a) Cnemaspispsychedelica from Ca Mau Province, Photo L.L. Grismer; b) Cyrtodactylus cattienensis from Dong Nai
Province, Photo P. Geissler; c) Cyrtodactylus roesleri from Quang Binh Province, Photo T. Ziegler; and d) Dixonius aaronbaueri
from Ninh Thuan Province, Photo T.V. Ngo.
English name: Ngovantrfs Butterfly Lizard. Remarks: Specimens identified as Pseudocalotesflowed
from Kon Turn Province (Bain et al. 2007) were subse-
Vietnamese name: Nhong cat ngo van tri. quently re-identified as P. ziegleri by Hallermann et al.
(2010).
Distribution: This species (Fig. lc) is currentlyknownon-
ly from Vietnam (Ba Ria-Vung Tau Province).
Gekkonidae
Pseudocalotes ziegleri Hallermann, Nguyen, Orlov & Cnemaspispsychedelica Grismer, Ngo & Grismer, 2010
Ananjeva, 2010
Cnemaspis psychedelica L.L. Grismer, T.V. Ngo & J.L.
Pseudocalotes ziegleri J. Hallermann, T.Q. Nguyen, N. Grismer, 2010, Zootaxa 2352: 48.
&
Orlov N. Ananjeva, 2010, Russ. J. Herpetol. 17(1): 32.
UNS
Holotype: 0444.
Holotype: IEBR 330.
Type locality: Hon Khoai Island, Ngoc Hien District, Ca
Type locality: Nuoc Ka forest, near Mang Canh, Kon Mau Province, Vietnam.
Plong District, Kon Turn Province, Vietnam, ca. 1,200 m
a.s.l. English name: Psychedelic Gecko.
English name: Ziegler's Tree Lizard. Vietnamese name: Tac ke duoi vang.
Vietnamese name: Nhong zig-lo. Distribution: This species (Fig. 2a) is currently known on-
ly from the type locality.
Distribution: This species (Fig. Id) is currentlyknown on-
ly from Vietnam (Kon Turn Province).
Bonn zoological Bulletin 57 (2): 137-147 ©ZFMK
New amphibians and reptiles from Vietnam 141
Cyrtodactylus cattienensis Geissler, Nazarov, Orlov, Distribution: This species is currently known only from
Bohme, Phung, Nguyen & Ziegler, 2009 the type locality in Khanh Hoa Province.
Cyrtodactylus cattienensis P. Geissler, R. Nazarov, N.L. Dixonius aaronbaueri Ngo & Ziegler, 2009
&
Orlov, W. Bohme, T.M. Phung, T.Q. Nguyen T. Ziegler,
2009, Zootaxa2161, 21. Dixonius aaronbaueri TV. Ngo & T. Ziegler, 2009,
Zoosyst. Evol., 85(1): 119.
Holotype: IEBRA.0856.
UNS
Holotype: 0284.
Type locality: CatTienNational Park, DongNai Province,
m
Vietnam, 120 a.s.l. Type locality: BinhTien Forest Station,Ninh Hai District,
Nui Chua National Park, Ninh Thuan Province, southern
m
English name: Cattien Bent-toed Gecko. Vietnam, 4—5 a.s.l.
Vietnamese name: Thach sung ngon cat tien. English name: Aaron Bauer's Leaf-toed Gecko.
Distribution: This species (Fig. 2b) is currently known on- Vietnamese name: Than lan chan la a-ron-bau-o.
ly from Vietnam (Ba Ria-Vung Tau and Dong Nai
provinces). Distribution: This species (Fig. 2d) is currently known on-
ly from the type locality.
CyrtodactylusroesleriZiegler, Nazarov, Orlov, Nguyen,
Vu, Dang, Dinh & Schmitz, 2010 Gekko canhi Rosier, Nguyen, Doan, Ho, Nguyen &
Ziegler, 2010
Cyrtodactylus roesleri T. Ziegler, R. Nazarov, N. Orlov,
T.Q. Nguyen, T.N. Vu, K.N. Dang, T.H. Dinh & A. Gekko canhi H. Rosier, T.Q. Nguyen, K.V Doan, C.T. Ho,
Schmitz, 2010, Zootaxa, 2413: 24. T.T. Nguyen & T. Ziegler, 2009, Zootaxa 2329: 57.
Holotype: ZFMK 89377. Holotype: IEBRA.0910.
Type locality: Phong Nha-Ke Bang National Park, Minh Type locality: Huu Lien, Huu Lung, Lang Son Province,
Hoa District, Quang Binh Province, Vietnam. North Vietnam.
English name: Roesler's Bent-toed Gecko. English name: Canh's Gecko.
Vietnamese name: Thach sung ngon ro-x-lo. Vietnamese name: Tac ke canh.
Distribution: This species (Fig. 2c) is currentlyknown on- Distribution: This species (Fig. 3a) is currently known on-
ly from the type locality. ly from northern Vietnam (Lang Son and Lao Cai
provinces).
Cyrtodactylusyangbayensis Ngo & Chan, 2010
Gekko russelltrainiNgo, Bauer, Wood & Grismer, 2009
Cyrtodactylus yangbayensis TV. Ngo & K.O. Chan,
Zootaxa, 2504: 48. Gekko russelltraini TV. Ngo, A.M. Bauer, PL. Jr. Wood
&
J.L. Grismer, 2009, Zootaxa 2238: 34.
UNS
Holotype: 0476.
UNS
Holotype: 0293.
Type locality: Yang Bay Waterfall, Dien Khanh District,
m
Khanh HoaProvince, southernVietnam, 500-600 a.s.l. Type locality: Chua Chan Mountain, Suoi CatCommune,
Xuan Loc District, Dong Nai Province, Vietnam, ca. 100
m
English name: Yangbay Bent-toed Gecko. a.s.l.
Vietnamese name: Than lan chan ngon yang bay. English name: Russell Train's Marble Gecko.
Bonn zoological Bulletin 57 (2): 137-147 ©ZFMK
142 Thomas Ziegler & Truong Quang Nguyen
Fig. 3. a) Portrait ofpreserved Gekko canhi from Lang Son Province, Photo T. Ziegler; b) Gekko russelltraini from Dong Nai
Province, Photo T.V. Ngo; c) Gekko takouensis from Binh Thuan Province, Photo S.N. Nguyen; and d) Gekko vietnamensis from
An Giang Province, Photo S.N. Nguyen.
Vietnamese name: Than lan da ru-xen-tren. Gekko vietnamensis Nguyen, 2010
Distribution: This species (Fig. 3b) is currentlyknownon- Gekko vietnamensis S.N. Nguyen, 2010, Zootaxa 2501:
ly from Vietnam (Dong Nai Province). 55.
Holotype: ITBCZ 667.
Gekko takouensis Ngo & Gamble, 2010
Type locality: Tuc Dup Hill, An Giang Province, south-
Gekko takouensis T.V. Ngo & T. Gamble, 2010, Zootaxa ern Vietnam, 43 m a.s.l.
2346: 18.
English name: Vietnam Gecko
Holotype: UNS 0491.
Vietnamese name: Tac ke viet nam.
Type locality: Ta Kou Mountain, Ham Thuan Nam Dis-
m
trict, Binh Thuan Province, Vietnam, 425 a.s.l. Distribution: This species (Fig. 3d) is currentlyknownon-
ly from the type locality.
English name: Takou Marbled Gecko.
Vietnamese name: Than lan da ta kou. Scincidae
Distribution: This species (Fig. 3c) is currentlyknown on- Scincella apraefrontalis Nguyen, Nguyen, Bohme &
ly from the type locality. Ziegler 2010
Scincella apraefrontalis T.Q. Nguyen, S.V Nguyen, W.
Bohme & T. Ziegler, 2010, Folia. Zool, 59(2): 116.
Bonn zoological Bulletin 57 (2): 137-147 ©ZFMK
New amphibians and reptiles from Vietnam 143
Holotype: IEBRA.0832. CalamariasangiNguyen, Koch & Ziegler, 2010 (2009)
Type locality: Huu Lien Nature Reserve, Huu Lung Dis- CalamariasangiT.Q. Nguyen,A. Koch &T. Ziegler, 2010
m
trict, Lang Son Province, Vietnam, ca. 200 a.s.l. "2009", Hamadryad 34(1): 2.
English name: Huulien Ground Skink. Holotype: IEBR 360.
Vietnamese name: Than lan co huu lien. Type locality: Mang Canh Commune, Kon Plong District,
Kon Turn Province, Vietnam, 1,200 m a.s.l.
Distribution: This species is only known from Lang Son
Province, northeastern Vietnam. English name: Sang's Reed Snake.
Vietnamese name: Ran mai gam sang.
Tropidophorus boehmei Nguyen, Nguyen, Schmitz,
Orlov & Ziegler, 2010 Distribution: This species is currently known only from
Vietnam.
Tropidophorus boehmei T.Q. Nguyen, T.T. Nguyen, A.
Schmitz, NX. Orlov & T. Ziegler, 2010, Zootaxa 2439:
57. Colubroelaps nguyenvansangiOrlov, Kharin, Ananje-
va, Nguyen & Nguyen, 2009
VNMN
Holotype: 822.
Colubroelaps nguyenvansangi N.L. Orlov, V.E. Kharin,
Type locality: Hoang Lien Mountain, near Ban Khoang, N.B. Ananjeva, T.T. Nguyen & T.Q. Nguyen, 2009, Russ.
Sa Pa District, Lao Cai Province, northern Vietnam, J- Herpetol. 16(3): 235.
m
1,200-1,300 a.s.l.
Holotype: ZISP/IEBR 25682.
English name: Boehme's Water Skink
Type locality: Loc Bac Forest Enterprise, Lam Dong
m
Vietnamese name: Than lan tai boe-me Province, Vietnam, ca. 720 a.s.l.
Distribution: This species (Fig. 4a) is currentlyknownon- English name: Nguyenvansang's Snake.
ly from Hoang Lien Mountain in Sa Pa and Van Ban dis-
tricts, Lao Cai Province, Vietnam. Vietnamese name: Ran nguyen van sang.
Distribution: The secondrecord ofthis species wasreport-
Serpentes edbyN. Poyarkov from Bu Gia MapNational Park, Binh
Phuoc Province (Fig. 5). Therefore Colubroelaps nguyen-
Colubridae vansangi is currently known from Lam Dong and Binh
Phuoc provinces, Vietnam.
&
Calamariagialaiensis Ziegler, Nguyen Nguyen, 2008
Calamaria gialaiensis T. Ziegler, S.V. Nguyen & T.Q. Lycodon ruhstrati abditus Vogel, David, Pauwels,
Nguyen, 2008, Current Herpetol., 27(2): 72. Sumontha, Norval, Hendrix, Vu & Ziegler, 2009
Holotype: IEBRA.0714. Lycodon ruhstrati abditus G. Vogel, P. David, O.S.G.
Pauwels, M. Sumontha, G. Norval, R. Hendrix, T.N. Vu
Type locality: Kon Ka Kinh, K Bang District, Gia Lai & T. Ziegler, 2009, Tropical Zoology 22(2): 144.
m
Province, Vietnam, 1300 a.s.l.
ZFMK
Holotype: 86451.
English name: Gialai Reed Snake.
Type locality: U Bo region, Phong Nha - Ke Bang Na-
Vietnamese name: Ran mai gam gia lai. tional Park, Quang Binh Province, Vietnam.
Distribution: This species (Fig. 4b) is currentlyknown on- English name: Hidden Mountain WolfSnake.
ly from Gia Lai Province, Vietnam.
Vietnamese name: Ran khuyet an.
Bonn zoological Bulletin 57 (2): 137-147 ©ZFMK
144 Thomas Ziegler & Truong Quang Nguyen
Fig. 4. a) Tropidophorus boehmei from Lao Cai Province, Photo T.T. Nguyen; b) Portrait ofpreserved Calamaria gialaiensis
from Gia Lai Province, Photo T. Ziegler; and c) Protobothvops trungkhanhensis from Cao Bang Province, Photo T.T. Nguyen.
Fig. 5. Colubroelaps nguyenvansangi from Binh Phuoc Province, Photo N. Poyarkov.
Bonn zoological Bulletin 57 (2): 137-147 ©ZFMK
New amphibians and reptiles from Vietnam 145
450
385
400 368-
350
295
276
300 ^5§"
250
Fteptiles
177 181
200 ±62- Aupliibiais
150
87 82
100
id
50
0
Eto(1977- Naiyai& I^nmet al Miiyaiet aL July 3010
1982) H>(1996") (3005) (2009)
Fig. 6. Species diversity ofthe herpetofauna ofVietnam.
Distribution: This subspecies is currently known from DISCUSSION
Quang Binh and Vinh Phuc provinces in Vietnam, and
from Fujian, Anhui, Zhejiang, Guangdong, Yunnan, After the publication of the "Herpetofauna ofVietnam,,
Sichuan andGansuprovinces in China. Because it has of- by Nguyen et al. (2009) 20 new amphibian and reptilian
ten been confused with other taxa, its range is probably species, one new subspecies, and a new snake genus have
much wider, both in northern Vietnam and China (prob- been described fromVietnam byJune 2010. Among them
ably also present in Guangxi Province). there were three new amphibians (1 Megophryidae, 1
Ranidae, 1 Rhacophoridae) and 18 new reptilian taxa (2
Agamidae, 9 Gekkonidae, 2 Scincidae, 4 Colubridae, and
Viperidae 1 Viperidae). In contrast, two species which were listed
as valid and occurring inVietnam in Nguyen etal. (2009)
Protobothrops trungkhanhensis Orlov, Ryabov & were synonymizedmeanwhile: Gekko ulikovskiiDarevsky
Nguyen, 2009 & Orlov, 1994 was regardedas ajuniorsynonym ofGekko
badenii Szczerbak & Nekrasova, 1994 by Nguyen et al.
Protobothrops trungkhanhensisNX. Orlov, S.A. Ryabov (2010d) and the specimen previously identified as
& T.T. Nguyen, 2009, Russ. J. Herpetol. 16(1): 71. Pseudocalotesfloweri from Kon Turn Province was re-
identifiedasP. ziegleriby Hallermann et al. (2010). Most
Holotype: ZISP 25351. of the recent species' discoveries affected lizards, with
geckos clearly being the predominant group. In addition
Type locality: Trung Khanh Nature Reserve, Trung to these new species descriptions, three new country
m
Khanh District, Cao Bang Province, Vietnam, 600 a.s.l. records were published after the book of Nguyen et al.
(2009): one amphibian species, the megophryid anuran
English name: Trungkhanh Pitviper. Leptobrachiumpromustache, the scincid lizard Scincella
monticola, and the colubrid snakeAmphiesmoides ornat-
Vietnamese name: Ran luc trung khanh. iceps (Bain et al. 2009b, Nguyen et al. 2010a, b).
Distribution: This species (Fig. 4c) is currently knownon- The results ofthis paper clearly exemplify that even af-
ly from the type locality. ter the comprehensive book provided by Nguyen et al.
Bonn zoological Bulletin 57 (2): 137-147 ©ZFMK
146 Thomas Ziegler & Truong Quang Nguyen
GeisslerP,NazarovR,OrlovNL, BohmeW, PhungMT,Nguyen
(2009) much research is needed to describeVietnam'srich QT, Ziegler T (2009)Anew species ofthe Cyrtodactylus ir-
herpetodiversity. Inparticularbecause not only cryptic or regularis complex (Squamata: Gekkonidae) from southern
inconspicuous species were discovered and formally de- Vietnam. Zootaxa 2161, 20-32
scribed in the past months, but also striking and colour- GrismerJL, GrismerLL(2010) Who's yourmommy? Identify-
ful species like Cnemaspis psychedelica (Grismer et al. ing maternal ancestors ofasexual species ofLeiolepis Cuvi-
2010) or even new genera, as was recently shown by the er, 1829 andthedescriptionofanewendemic speciesofasex-
ual Leiolepis Cuvier, 1829 from Southern Vietnam Zootaxa
description ofColubroelaps (Orlov etal. 2009a). Current- 2433: 47-61
ly, the herpetofauna ofVietnam comprises 181 species of Grismer LL, Ngo VT, Grismer JL (2010) A new colorful new
amphibians and 385 species ofreptiles (Fig. 6). Howev- species ofinsularrockgecko (Cnemapsis Strauch 1887) from
er, diversity research and species inventories are only the southern Vietnam. Zootaxa 2352: 46-58
first steps, which must be followed by investigations of Hallermann J, Nguyen QT, OrlovN,AnanjevaN (2010)Anew
species of the genus Pseudocalotes (Squamata: Agamidae)
the natural history and specific adaptations, which final-
from Vietnam. Russ. J. Herpetol. 17(1): 31^40
ly are prerequisites for adequate conservation measures. Morice A (1875) Coup d'oeil sur la faune de la Cochinchine
francaise. Ass. Lyon. A. Sci. Nat, Lyon: 25-121
Acknowledgements. We are indebted to our colleagues Peter NgsopeVcTie,sBoafueGrekAkMo,LWauoroendtiP,L1J7r6,8Gr(iSsqmuearmaJtLa:(2G0e0k9k)onAidnaeew)
Geissler (Bonn), L. Lee Grismer (Riverside), Ngo Van Tri, from Dong Nai Province, Southeastern Vietnam. Zootaxa
Nguyen Ngoc Sang (Ho Chi Minh City), Nguyen Thien Tao 2238: 33^2
(Hanoi),Nikolay Poyarkov(Moscow), andNikolaiL. Orlov (St. Ngo VT, GambleT (2010)Anew species ofGekko (Squamata:
Petersburg) who kindly provided photographs. Gekkonidae) from Ta Kou Nature Reserve, Binh Thuan
Province, southern Vietnam. Zootaxa 2346: 17-28
Ngo VT, Onn CK(2010)Anew species ofCyrtodactylus Gray,
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