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Name-bearing types of subspecies of Sylvia curruca (L.) and S. althaea Hume in the collection of the Zoological Institute, St. Petersburg (Aves: Sylviidae) PDF

11 Pages·2001·1.3 MB·
by  LoskotV M
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Preview Name-bearing types of subspecies of Sylvia curruca (L.) and S. althaea Hume in the collection of the Zoological Institute, St. Petersburg (Aves: Sylviidae)

© ZooloIgnisctSaitlt. uPteet,2e 0Ir0 s burg, Name-btyepareiosnfs g u bspoefScy ilecvusir ar (uLc.a) S an.da lthaHeuami etn h ceo lleocftt hZieoo onl ogical Institute,(.ASvStey.slP:ve itiedraseb)u rg V.M.Loskot LosVk. oMt2.,0 0N1a.m e-bteyaporesfisu n bgs poeSfyc licveuisra r( uLac.an)S. d a lthaea Humietn h ceo lloetfch tZeio oonl oIgnisctSaitlt. uPteet(,eA rvSseybslu:vr iZogio dsysa­e). temaRtoiscsa1i 0c(a21,1) 9:- 229. Typsep eciomffie vsneus b spoefScy ilcveuisra r (uS.cc h.aa limoSdec.nt. dre il,e ngitica, S.cs.n igirSe.jw axscak.ri taii,nSc dc.a c. ,h uancahnoediS.f ac lat)h maoenat iacreoexla ­ amineLde.c toatrdyeep seisg fonrSat .ce udr rhuaclai moSdeunsdrhi1k 9i0an4n,S. d c . telengSiutsihc1ka9i 2Dn5a,.to ana ll stpyepceiam rreeen fisan nesddu ppleimne­nted, cludtihnelgia rba enledsx temronraplh (opllougcmyoa lgoerd aitmieonansn,iwd oi nnsg, formuBlrai)he.if s otfoi rnyv estoifgg eaotgiroavnpa hriiicanSat .lci u ornr auncSda. althiapser ae seTnhsteue bds.p setcatoiuftes hs Te u rkmLeensiWsahenir t e(tS.hc .roat snigiSrteawsckh1ia9in2 io9csw)o , n firImitfoes ud n.td ht ahhteo lootytfph seiu sb species ikse iptnth Zeo oloIgnisctSaitlt. uPteetr,ea rttshhbeiaurtnnr h Zgeo oloMguisceoauflm thMeo scUonwi vewrhsetirwtyepo, a raatrpyerp eessNe erwv ecddoa.nt fiart mhSiac.nt. g chuancPhoeritc1ea9n i5kas5oj ,u nsiyonro onfSyc. mm. a rgelSatnoiicz1am8 a9an9rn e, presented. V.M LoskZooto.lo gIincsatlRi utsustAieca,an de omfScy i enUncievse,r sain taebIt.,s k ay St.Pet1e9r9s0Rb3usu4sr,ig a . Introduction currbulcyfoatr th hie bsierM dosso:tft hbei rds correstpoto hnpedu ebdl idsihaegdnb ouste s, Amonwgi dedliys trPiabluateeWadar rc­ttihcee rxei satliesndod ivwiiditunhat lesr mediate bleorfts hg ee nSuysl tvhiiead ,e ntiofifc tatriaoiinnt tse gtrhaseta imnpiglna es si nrgolwe. tafoxram itnhsge u persS.p ceucrir-eusSc .a H um(e1 8h7a8es)m phatshireze eaodlf di itfy­ althpareeasc eonntssi ddeirffiacbulTleht ei feerse.n bceetsw tehedene scrfoirbmaesnd td h e maifena tuorfte hsge e ogrvaaprhiiiacnt idoinffi ciuntl htaeys sitghneitmnoo g no ert hree sizaen pd lumcaogleo roaftt hiLeoe ns sedri ffesrpeencDtie esspl.ia trnege esw k miant e­ White(tS.hc ruoraratun cMdao )u ntWahiint er­iaanldd a otnal ihfei stooftr hyeW saer blers thr(So.aa tl thwaeerade)e scrbiyHb uemde collientc htpeea d1s 2ty0 e atrhsde,i scussion (1817837,w8 h)ho,a edx ami1n1se6kd i onfs o nt htea xonooftm hyge r oiuscp o ntinuing Indbiianr cdosl lecchtieeidflnt y h wei ntearl asptor esAesbn etfo.tr heo,ep .i noifto anxs­ timAem.o ntgh eHmu,m dei stintghueio snhoemddi isfftwesir d eolnt yhn eu mboefsr p e­ sma"lmli nuwliatt"hhl, ei ghutpeppseatrr tcsi,(e 1s- t3h)es,ic ro npuem,b oefsr u bspecies "manptallseea nbdryo wann""d c ropwanl e( 7-1a2n)ed,s pecitahldeli ys tornoi fb ution blugirsehay n"ad,l wsiot ahb luwnitnt go ps ubspAe vcailseitst .e dreaatwluitrtt ehh ese (2= 7 ,s omet8i)(m Heusm1 e8,71 38,7 t8h)e,p robltehmmeas i;nn o menclaanttdau xroa­l lar"gael thhaaevai""un pgsp uerrfa dcaer kinsohm iincfe rewnecrpeeus b liisnrh eevdi ews gresyl,i gthitnwlgiyet bdhr oownnt hbea cka'n'md a nuiatnlh lsea syte a4(r0Vs a u1r9i5e9,; (Hum1e8,7 8an)d,a lstohfo er ·m" affinisP"o,r te1n9k6oW0,i; l lia1m9s6Ko8on;r, e lov, cloisnpe l umcaogleo rtatoti hoEenu rope1a9n7 A2l;&i R ipl1e9y7S,3t ;e pa1n9y7a8n,, "citrrbuucwtai "t",hl aragnsedtr o ubtielrl1 "9.8 139,9W 0o;l t1e9r8sE0,t; c he&cH oilpea,r Thlea tfotremwr a ass siegrnreodn teoou 1s9l8·yW3 a;t s1o9n8,C6 h;e n1g9,8 H7a;ff er, SylavffiinaBi lsy 1t8h4i,5nt hsee caorntdi c1l9e9R 1o;s e1l9a9a2r;,B 1a9k9e7r),. byH um(e1 86708s):u; b seqTuiecnet&hl uyr sCto lleocftt hiZeoo nosl oIgnisctaSiltt .uPtee­, Whis(t1l9eh3ra3 v)ee s tabtlhinesa hmS.ee d ters(bZuIrSwgPe )ro efs peciimaplo ritna nce VM LosTkyopiteSn:sy lcvuirar auncS.daa /th•aZ eOaO SYSRTO.SC SAIV ol1.0 220 thset uodfty h gee ogravpahriiciaanttl hi iosMn a tearnimdae lt hods groPulpe.(s 1k8eh8 a9ms)a dteh fier rsetv i­ siooftn h ebma,so entd h aen aloyf6s 9si pse ciT­ypsee roifse istx a bxeal ontgoti hsneug ­ menasn,ad c cetphtseep de csiteaostfS. u c su r­perspSe.cc uirer-suS. ca al thwaeereaex am- ·· rucS.a a,l thaaneS.da m inu(la=m inuscuilnae)d.. Latoenrl, a rmgaet ewraicsao ll lbeycN t.e dT htee oxfot r iglianbiaegslli svi enqn u ota­ SeverNt.Ps rozve,w aPl.Ks okzila,on vPd . timoanr tkhsae;d dictoinocnestrn hibeni gr ds Sushmkaiinin ntl hyse o utphaeorrftnWS ibea­gel,o caalniddta yto efc olleacrpteri en­g riAal,t MatisS .a,y Matns a.n,Md i ddalned sentseqdu airne brackets. CentArsaitlah ;ce o lleocfMt .iM oenn zbieArls lt anmdeaarsdu r(elmeenongftwt sih n g, warse ceailvsIeontd. h m ei d-twoef2n 0ttihte asti alr,as nubdsi ,lw ler)ce a rroiuuetsd i an g centtuhcreoy l,l eocfst kiioonfS.ns c urroufc dai caall lwiipt0eh.mr 1m p reciBsifriloolmn . ZISbPe catmhweeo rllda'rsgS ensitg.i refewastkhiiets rh lsee nogfct uhl mfreonmt hbea se (192w9h)o,is nev estoiftg hagete ioognr aopfth hifoe­r emfeoastth oeftr hfseo rewhiedatdh; cavla riaotfti hosenp ecwiaeassm ontgh ea ndde potfbh i wlelrm ee asauttrhe lede voefl mosetx tenussieavdle r,ea abdo6yu0 ts0 k intsh freo netd goeft hnear e.s. fromt hciosl leIcnht iiosop ni.nt ihopeno ,l y­Thfoel lowaibnbgr evairauets ieiodntn hse typsipce cS.i ceusr rcuocmap r8is suebds pdee­scroifpp tliuomanansgad eg c el asjsu-evs.: ci(eS.sc c.u rrSu.cc t.ae ,l enS.g ci.cta iuc­aj,u venbiilriednsjs ,u vepnliulmea lg we ; - casiSc.ca .h, a limoS.de cn.dm riin,uS. l ca., b iridnfis r wsitn (tfreerps lhu)m aaftgpeeor s t­ jaxartS.i cc.ma a,r gelaannS.di cc.ta u,r kj­uvemnoiulllets -; b iridnfis r sstu mmer meniacnatd)h m eo rpahffian (i=sS. curru(cwao rpnl)u maagd-e a;dl utbsi,r adftset rh e blyTtihcie h&u Wrhsits t1l9e3Sr3n,)i .g irfierwcs­otm pmloeutAlelt- a ;l ufelatah e1r-sP3 ; skrie gaSr.ad letdhH auema1e 8,7 a8sa s epa-­prim1a-r1in0eu,sm befrroemtd ho eu termost rastpee caindedis nd o ats stioigt tnh Cea uca(­lP) P;C - primcaorvyeR rt sr-;e ct1r-i6c,es sibainr (dSsy lcvaiuac aOsgince&av B ank­numbefrroemtd hc ee nptar(iaRrll ). ovs1k9i1,H0 e)c .o nsitdhelera etttdeob r ea subspoefcS. i ceusr rusctar,e smsuicnhSg y lcvuirar huaclai moSduesnhd1kr9ii0-n 4, smalslieozrfet hebiiarls cl o mpawrietS.dh KazakhLsetsaWsnheiirat ne throat althaea. Thnee rxetv iosfi ZotInhS mePat er(imaolr eS ush(k1i9nh0 a4ds)e sctrhisibuseb ds pecies tha7n0 0s pecimweanscs a)r roiuebtdy verbyr iefrfloyms isxp ecicmoelnlsbe yc ted Portenko (1a9v 6e0rw)yi. d ienA tdeohrpi­tmis naignna d ttsh l eo wIerrg ahnildzo wTeurr ­ pretoaftt hipeoo nl ytsyppeic_ciha eess s,i gngeaRdii veAr mso.rd ee tadiilaegdno ofts hies toS .c urr1u0sc uab spseicoxift e hse(,mSc .. subspiencciletushdd eii nffge rfreonmS.c ce.s currS.u cc.ca ,a ucaS.s ci.ac lat,h S.a ce.a ,c urrSu.cc m.ai ,n u(laSas.m inusocrSu .cl .a jaxartS.i cc.haa ,/ imoadennS.ddr c it. e lenmgiin­usacnudSl a.m) a rgelSatnoilczam ann, ticnae)s tiintnh tgee rroifMt oonrgyoan ldi a1 89a7sw, e alsla dditdiaootnnada ils tribution thfoer meUrS SaRn odn meo r(eS.c .m ar­anldi hfei stwoerrgyei vienan n otahretri cle gelapnriocbaa)b liynt hnPeea smtiirnog- (ASluasi1h.9k 0i8n),. Theo thseurb spoefcL ieesssW ehrit ethrAo raetv iasnesddu ppledmeesnctreoidfp tion recogbnyiP zoerdt e(n1k9o5w 5e)rS. e c . types peciomfSe .cn .hs a limoidspe rnedr­i minuS.lc ac.,h uancahnS.edc i m.co an ticoslean.ht eerdIe ns. p ring-psluumm(maAegprer il Asa r esoufdl ets croifnp etfowir omnbs y toJ ultyh)foi,rs md ifffreormS.s c c.u rriunc a Sush(k1i9n0 149,2 5S)n,i gir(e1w9s2k7ti,h s eh orbtiwelirln ,fog r mu(l6a> 2v>sP7.2= 192a9n)Pd o rte(n1k9o5 h5o)l,o tyoprse ysn ­P6r,a r6e>l2y>o 7r5 >2>a6s)h,- glreesys, typsee roifse istx a xoaft hseu persSp.ebcrioewscn riosawhnn nd a paenl,di gahntgder re y curr-uS.c aal thaarkeeea ip ntt hl! collreeocsfutt pi popenar rI titscs .l otsoee a stseprne ci­ ofZ ISTPh.ie n teirnte htseat x onaonmailcy mseinsos ft hneo minostyupbiscpa[elSc uisehsk in ofg eogravpahriiciaanttlh i eoWsnae r bilne­rdse sigtnhaetamesv d a ra.ffi ni(sn oSw.c . creacsoends idientr harebe lcyyee natBr es­.b lythhaiv)it]nh,sge a mwei nfogr muAlsa . caumsoeso tfo rigdiensaclr iaprbteri ioencfso mpawrieStcd.hh . a limodesnodurtih,e.rtnh e ansdo metiinmeex(safo cirtn sttahndeci e­,S. c.m inu(l=ma i nushcauasls ah)o rbtielrl mensioofbn isrw desrg ei vbeySn n igireawnmsdok rire o unde(do ftewPni2= n P g7o re ven anPdo rtewnikthoor uetg aforrs de dxi ffer7­>2>w8i)tl,hi g chrtoeawrnn rd u fobuasc S.k; enceisbt)e ,c onmeecse stsoda ersyc trhiebc e.m argelhaaansl i ocnawg ienrag ns dh arper tysppee ciimdnee ntsa il. differiencn ocleosro aft"icanoadnpb "a ck. ZOOSYRSOTS.S IVCoAl1 .0• V. ML oskToytpi:eSyn sl vciuarr uacnSada. l thaea 221 Upppearrt osft hsei nbgilriednj uvenOinloeeft hmea l(eSsu s1h9k0i68n9:,l1 o,c ality plumaargyeee lltohwaetnrh oosfae d ultnso1..) c ,o lloenc2 t7e.dI V[9i.nsV a]n.d1s8 98 The·yaa rlels iog ahntldee rbs rso awscn o m­Sarykoontp hale,o wIerrg RhiivzSe o rf,. the parteotd h bosfS.e· d ciu rraunS.cd ca v. a arf­. towInr ghiisaz b,s eAnctc.o rtdoti hneg, ar­ fini{s=S. c.b lyitnh.ij)u pvleunmiaalngede c hiovfet shO er nithDoelpoagryti mhtea nst , greayneddra rkeesrp>e coinca rlolwtynh ,a nn obte erne cebiyZv IeSdAP l.sl y nthyapvees thoosSfec m.i n(u=lm ai nuswciutlhtaoh)ue,ta no riglianbbaeylPl . S ushankdia nl abel welelx preisnSs. c e.md i nuyleal ltoiiwnn t "ZoIon!As.Nt S .S S(Ri"Cn y riclhlairca cters; lowpearor fct r oawnnod nn ape. latoencr,i taesZd I SlPa bfoeulsr)p ;e cimens Notitnhrgee alaintycd o nstoafmn ecny­ havael ast oh isrtda ndarlda b"peCrloi lnlt.e d tiotnreadSi utsshh,ka issnt retshisene tde rPm.eS­uschwkiitcnhh" a racteristic rounded left diaptoes iotfS.i coh.na limobdeetnwS.dreie mna rgaintnde wxrti tbytS euns hiknGi enr man. cv.aarffi.n (i=Ss .c b .l·y atnhSdic. )m. i n(u=laT hlea tltaebireJs la ckoinnlgoy n seip ne cimen minuscula). (n3o).w, h ihcahbs e egni vbeySn u shtkoin Ina·d di,tttiohom eno rphologicaMl. AMc.eh nazrabanichdeta rebs re ser ne cebiyZv IeSdP mentiS.o ·nceh:da ,l imriod dieffneidrnts h ew ihtihcs o llection. breeadriaennaghd· a biBtiaratdr.sfoe u nidn Ont hfaec eo ft hoer iglianbsaeelldx ,a, t e NEAr·a iS eaar,e iant, h orbnuys hHeasl,i (­OlSdt yanldel )o caalrgieit vybe unnt,o t th e modendrboine berZseiret(io=anv riagi e ntceoulml encatm(oeir S.'uess. h kIinns)c.ro inp tions Bie1b8.0n,8o L;n. 1,7 7a4n)Ed l aeagnsuosn ­thbea cokft hlea baermlea dlea taeser v,i ­ gar(iBceare nxSh c.h leScchhtl.(e)�ch hotr.­d encbeydd i fferdeanrtcko,el roo fuI rn dian _teBnesriensxhS .c hlegcrhotwo.in)mn ogi sitn tkh;ei yn cludeo:f t hnseup mebceiirmn e n sansdoyio lnls o wIerrg ahnildzo wTeurr gatihs ea mpolf.e ts hpee ciesi ntc hotilrsli epc,t ed Riveirnsc,l uhdiilnslgay n Tduss uImn. leng(tmhmo )f w in(gU a)n tdh aotfb. i ll Sushkoipni'ntsih oen ,M ugaondtdzh hea( rCyu lmme)a sutrose kdua lnlwd,i nfogr mula aroefaKN a zakphisfontreae nas rtiesn habistheodw. itnphgoe s iotfti ho2enP .rteilpa ttoi ve byn ortfohremr[snS.c c.u rrauncS.d ac v.a r.o thPe( rfo erx am6p.l2e.,T7 h.se)a .md ea ta argei voenln a boeflSsu shckoilnl'ebscu tti on, affini(s= S. c.b lytThhie)hr]ae.vb ee enno herdea toefcs o lleacritenti hnNege wS tyle, daotnat hseo utbhoeurnnd oafdr iisetsr ibution alfolu srp eciamrieedn esn ats"i Sfiyeldcv uira­ ofS .c h.a limoTdehnfiedrn iod.f a b iwridt h ruchaal imoSdeunsdrhika intndh" we,o r"dt y­ charaocfS.t ecr.hs a limoadennS.ddr ic . pusi"ws r itbtySe uns hoknti hnle a boefol n e minublrNa . MP.r zewianlt shUkeri u ngu oft hmea lTehsi.rs e cotrodg,ew tihtiehnr d i­ Rivvearl·il nAe pyr ailll oSwuesdh( k1i9n0 8: vidduiaffle rbeentcweese n wshyinictnhy­ pes, 691�t6os9 u2g)g ave asdstit s troifS.b cu.tion cluidned ivoifdd iuffaelarsge ceno tl laetc ted halimodetnhedear sit..i n diffedraetnpetes r,mt iodt ess igtnhmateee n ­ LatSeriri,g ir(e1w9s2ck9oi)n firtmheids tioned asslp eecctiomteynp e. suggeAsctcioortndoh.i i nS.mg ,c h .a limoden­ Lecto(dty epsei ghneart"ed'e),[d. s1 ] S,a ry­ drnie sfrtosmt hNeEs hoorfte hC ea spSieaan kop[aN,K azakhssatnoadnnsl , o.wI errg hiz tfoo otohfWi AllltsZa aii,Ds eapnr easnsdi on ° Riv4e8r2,2 '6N°1, 3 82'8E.,I V[lOU. V].1898. TarbaMgtasti:ant,y hs eo uutpth ot hleo wer =6 3C,u l=m 1 2.62.;2n .o72.," P ,.S ushkin Amudacre'nyptaar,ora fttl h Key zylkDuem­ (ZIS1P2,0 9W8i)n.6g 2 .t6a,5i 5l. t6a,r sus seartn Tda shkMeingtrb.ai nrtwd esrc eo l­ 19.b7ifr,lo lmfe ath8e.rb3si,fr l olmn ar6e.s8 , lecitnte hdSeA ltSaeim,i reFcehrygea,n a,the widotfbh i 2ll. d7e,p otfbh i 2l.lW9 i.n fogr ­ PamiArmsu,d aErT 'ryaan,s caarsaepnaEid a n mul6a> 2>t7io;pfP 2s itunateeadtr ote irop f Iraann,di ,na utuamlnis,noD .z hunagnardi a P6;o utero ntv hateno oepf P 6w itshh allow KashgeaatrsHoito a t wainn;t ienIr nidniga . notCcohl.o roaftR i6to ynp iocfya ela rling ThKea zakshusbtsapnwe acasic ecseb pyt ebdi rwdh:i t(inspohut r weh ifiteel)od c cupies mosttax onombiuisttst scs o,wp ae.s s ometsiommeesw mhoartteh aonn e-otthiinrvndae nre , underdsitffoeorVdeo nltclhya(.n1 e9ti5sn4k­)ni a rroawlionsnghg a tftofe athbearso en;l y cluidnS.e d c h.a limoadelnmsdrooir s eo uthr­emnoafans tmsaw lhli tsippsohrt e setnhteo n erpno puldaetsicor_niassbs ee pdas ruabtsep weo­rtnio pf R5. ciSe csj. ax arStniicgai r1e9w2as9nkS. di c ,. Paralec(t1o)ty.[p'le'sssd']a.,,n a dmts o uth snigiSrteawcshk1ai9ni2o b9vu,t,t h oipsi nioofnI rgRhiivze2 r8,. V[9.VUI ]=6. 31,8­98. hansor te ceainvsyeu dp port. Cul=m1 2.65 2,=. n7o,3. "P ,.S ushfrkoimn , Fivoeft hseis xy ntoyfS.p ceh.sa limodecno­lleocfMt .iM oenn zb(iZeIrS1 P0,0 729). drairk ee pitnZ ISTPh.ew ye rceo llbeyc tWeidn 6g2 .t7a,5i 5l. .8t,a 1r9s.ub7si, frl olm PP.. S.u shiknsi pnr ainnsdgu mmeri.nt1 h8e9fe 8a th8e.rb6si,fr l olmn ar6e.sw7 i,d otfbh i ll aroefta h:Te u rdgeapir e(sNKs aizoank hs2t.a7dn,e)p .ot fhb i2l.l(8 2.") d' ,[ lss?a]n,d s 222 VML oskTypoetsi: Sn y lvciuar raunScda a l th•Za OeOaS YRSOTS.S IVCoA1l 0. Tusu[ma bo1u0t0k mE oft owInr ghizT,h eb rideefs crbiypS tuisohinkn icnl udes 20.VI[2.UV=I6I4CJ,u. l1=m8 1 9286,.. 2.t7h,foel lowdiant"gaA..b opvaelg er,e ywiisthh, n.o4. 'P \.S ush(kZiIn1S 2P0,0 W9i9n)6g.4 .4w,eldle velbolpueids ohn-t ghprei elyae nudm ta5i4l.t 4a,r1 s9u:bs7i fr,lo lmfe ath8e.br7is, l lpure grey lciotvtetlrtaesi ,lu -pcpoevre rwtisn g fromna6r.ew8si, d otfhb 2i.ld7le, p otfbh i lalnrde ct_rDiicmeesn.ls airbogineals,t l r ifle 2.(93.") cl [,l ss]a,nT duss u2m0,. VI[2s.hVoItrIht]iae.ncnr u rrbuulctoa n tgheiarnhn a li­ 189U8 .6= 4 C,u l=m1 26,. 2n.o75.," P ,. modenndotrthi iw,ni ;ncl g 6 3.5�-6 659-,6 6, Sush(kZiIn1S 2P0,1 W0i0n)6g.3 .t8a5,i7 l. 9c,ulm1e2n- 1w3i.nfo5gr; m umloas 6t>l2y> 7. tar1s9u.sb1 i,frl olm fe at�h8 e.rb6i,frH o m 18s peciSmEeA nlsta.an iNd W Mongolia. nar6e.sw4 i,d otfbh i 2l.ld7 e,p otfbh i 2l.l8 .T ypcle asd:cl. j ,.1 ,5 .VIIClh.u1si9ta1e 2p,­ (4")?s e0x,u vs.a]n,Td uss u2m0,. VI[2.VII]. pec;o Sluls.h kin". 189U8 .6= 2 C,u l=m1 1.65.,2n .o76.," P ,. Oft hmee ntisoenre1id6es sp,e ciamreen s Sush(kZiIn1S 2P0,1 01/4W6i6n6-g294 ,6, 0). keipntt h ceo lleocftZiIToShnPde .a tatw.oo n ta5i6l. 6,1 9t.ba6irfr,lso lumfe s a th6e.br7is,l l birndosrt e cebiyvZ eIdSa Pr gei vuennd er fromn ar5.e 4sw, i dotfbh i 2l.l(4 g roowfbt ihl l "Mateirnti hamelo "n og.ornaA plhtb aiir ds nocto mpleted). (Sush1k9i3n81,:8 2n)u:m be6rbsi( scl , DiscuTshsceio olno.r oafAt li3Po,Cna nd 12:VI.191R4i,v CeEhrlu,say ntage apSspEhe , R5-a6nt dh seh aopfte i opfPs C a nrde ctrices Altaanild)O b(io"s3, . VIll.olw�eIr�B,o ku­ shotwh tahtle e ctaont3ydp p aer alectotypes MorRiinv Kehra,r aN-WGM oobnig,o Elaicah) . aryee arilnifi nrgsssut m mpelru mPalguem.­ ofsy nthyatpshe rlsea eba esl sd,e fosrcS rc.i. bed agoef t wmoa lceosl loenc2 tV.eI idIsv ery halimoOdennt dhbreai co.kft hoer iglianbaell mucwho rens,p ectihcaeel nlptyar oiafrrl e c­ (wittehwx rti tltaetinenv r i oilnektt)h o,er igi­ triic1ne2 s0 099. naild entimfiacragteilioassnnu ibcsatb iyt uted AsS ushhkanison tPe2id n,m al1e0 07i2s9 telenagntidht neiu cmabo eftr hs ep ecoifm en equianlle ntgoPt6 ha, n idtn h ree osfpta ralec­ thsipse ccioelsli enAc ltteiadgsii v(etnhi esr e totyipnecsl,u dPi2ni gssjo umve.ws,hh aotr ter, ana bbrev"iAalbtteu.for"tre he ne u mbIenr ). buittt sii pss i tunaetaterotde iropf P 6t hatno addittihoweno ,r" dt ypiussw "r itbtye n tiopfP 7a,si nt hlee ctoTthtyerp aeoc.fte hse Sushoknti hnse a me ofsti hdee o lfaa dab nedl notocnth h teoo pft hoeu teri Pnv6 aa rvneei s­ lwm alceosl loenc1 t5e.dV IIiln". C1h9u1y2a ibiljneu va.nm da l1e0 07i2nt9 hr;ee maining stepapne"d"c otyopnlu asb"oe fol tsh tewro twmoa l(eiwsno rpnl umatghceih)sa, r iasc ter noetx pressed. bircdosl ltehceitrntee hds e a mdea Tyh.e isne­ Isth obuesl tdr etshsSaeutsd h kicno'ns­ sncortiepa trrieeo pnesaa ltosenosd t anldaabredl s cerntihineng t ermedioafSt .ce .h aploiso­ifSt uisohnk in'sT hcpeor lelseoecftnt wcioeo n. modenbdertiwm eeesno phniolrotSuh.sce .rns peciwmietanhus t hdoers'isg "ntaytpiuosn" currauncS.da c ·.b lyatnhsdio utdheesrenra tnc do nsiddeirffaebrilepnen l cuemsca ogned i­ formcsl otsoSec. m .i niutslr amu aei ansrl ey­ tiboent wbeierfrndo smt hsee roifse ysn types gardtihdneig m enosfbi iornIdsnts h.pe l·u m­gigvreo ufonrdd ess igonfta htleie ocnt otype. agceo lorS.a ct.hi aolni,m oidnehnadbriit Liencgt o(tydpees ihgenra"eto")e[,.da dv]i,c in­ semidoefsKNea rztask ihcssl toatsnose oru thi­toyfK osh-AgacShE A [lRtCuahsius,yi ­a, erfonr mIsna. l pll umaigtuesps p,pe arra trse s kasytae p4p°95e6,'8 N°84,2 '1E7,0m 0a. s . noticleiagbahlntyhde a rv dei styienlclto wIi.s2]h[, 1 5).VIII.n1o49.,t1 y2p,u Ps.A" l,t .[ai] tienstp,e coinma alnlatynlo deu tveafno efrs e ­ Sush(kZiIn1S 2P0,1 11/4W6i6n-g9 6607)..3 , migaenstd h egirrec aotv eIrnst usm.mp elru m­ ta6i0l.t 5a,r1 s9u.ts4o o,pfb iblrlo kweind,t h agSe.c,h . a limomdeanbydr ei aelassdioi­l sy ofb2i.ld5le, p otfhb 2i.lP8l2. -s 4th ialvnleo t tingufriosmhn eodr tfohremr•dsn u teo t he complgertoewidtnt h hp er ocoefps oss tbreed­ purgerre( yl ebsrso wcno)lo of"u cra mpu"c h inmgo ualnttd h ree wailnfo gr muclaan bneo t morseh arply cwointtthhr·gea r setyiinsgh deter(mPi2nt)eho;deu tvearno ent hteo opf bromwann talsie sn,o utfohremrTsnh. we h ite P6w itshh alnlootwc h. Pcuorleoo nu wrh ite pattoenRrn 5 a neds pecoinRa 6li lsay l so mordee velTohpede idff.e rfreonmcS e.cs . R6i sv erwye ldle velaonpodec dc utphiee s telenagrpiert eiscean ted beloww.h ooluet vearn em oarntedh a3/no4 ft hien ­ nevra nael ;a rwgeed,g e-swhhaisptpeeiod st, Sylvciuar rtueclae nSguisthi1kc9ia2n -5, presoentn hatep ipcaaroltf sRt 5h;we h istpeo t TelenLgeistsiearn Whitethrooantt h teo opfR 4i ss maolnlt ;ht eo opfR 3, theirosen aln ya rwrhoiwt eidsgshet ripe. Paralectotyp"etypsu slaa"bn "edcl oltye­d Thseu bspiensca imeaesftd eT re lenagni tes, etihcgn rooufAp l taiinahnasbt ihvteai lnlgpe uys b"yP .S ush(kwiibntr hin eofto ends i men­ oft hKea tRuinva enrid tt sr ibutariessi.o (n1s"))cl ., v[ilcw]io,nfK i otsyh -Agach, ZOOSYRSOTS.S IVCoAl .V. M1L 0o s•kTyp oetsi: Sn y lcvuirara unS.cda a l thaea 223 2[15].VAIlIntIo.3.. ,ty1 p9u1sP2".,S, u shkmionu nletfew:a thaerprers e soennn patem ,a ntle (ZI1S2P0,1 10/4W6i6n6-g59 .t64a05,i)8 l.. 5a,n udn deortfsh ibedo ed y. tar2s0u.sb4 i,frl olm fe ath8e.rb3si, frl olm S.c.t eleniglsia tritgcheaaor nt hseurb spe­ nar6e.sw4 i,d otf.h b 2i.ld5le, p otfbh i 2l.l8 .c ieesx,cS. e cp.tm argel(a=Sn .ci .cc hau an­ (2")se?x,[ w1] v,i cionfKi otsyh -Agcahcehi,icnma a)l:oe fts h tey psee r(in=e 1 s2 ), 2[15].VIAIlItn..o1 2.9, C1 o2t,y pPu.s"w,i n6g7 .8(±615..84 t-a76i00l.. 86)(±,51 8..72 - Sush(kZiIn1S 2P0,1 09/4W6i6n6-g89 .6206,)4.. t1a)r,2s 0u.s4 (±104..74 -b2ifr1lo.lm3 ), ta6i4l.t 1a,r2 s0u.bs6i fr,lo lmfe ath8e.br3is,l l feath(en=r 1s1 8). 7±(08..33 -b9ifr.lo1lm) , fromn ar6e.sw7 i,d otfbh i 2l.ld5 e,p otfbh i ll nar(en=s 1 17). 1±(06..44 w-i7d.ot7fbh) i ,l l 2.T8h.ce o nsidleernoagftbt hlhwee i nsgh ows (n=1 22). 5±(02..14 d-e2p.ot7fh)b ,i 112.9±0.l thtahtse p eciimsae pnp araem natll(ey3. ) (2.7-I3nt. h1rfe)em.ea lweisn,6g 4 .644,. 7, "d'[[?w1]] ,v, i cionfKi otsyh -A2g[a1c5h),. 66.t4a;5i 6l. 670,,6 50,.t 7a;r 1s9u.s26 0,. 1, VIIL1A9l1nt2o.,1. ,C otypPu.Ss u"s,h kin 20.b4ifr;lo lmfe ath7e.r86s.,8 1 .,b2 i;frl olm (ZI1S2P0,1 08/4W6i6n(-gl9 oe6nft06e )4).. 4, nar6e.s61 .,66 .,w9 i;d otfbh i 2l.l25 .,26 .,6 ; ta6i0l.t 7a,r2 s0u.bs4i ,l lfe afrtoh7me. br6is,l l depotfbh i ll2 .327..,07 ., fromn ar6e.sw1 i,d otfbh i 2l.ld5 e,p otfbh i ll Inm osstp eci(mn=e 1 n0st) h,we i nfogr ­ 2.I7n.t hoer iglianboaeftll h sipse ciimsen, rriulia6s > 2>i7nh;a lbfiorftd iospfR, 2 b eing writtthieaittnas 9 ,b utths ee ixcs h antgoed situnaetaetrdoet rht ei opfP 7i;nj uvenile d' int hlea boefSl u shkcionl'lseT chteion. mal1e2 01P12=9 P ,7 b,u itnt hriened ividuals lenogftt hhre i gwhitni g6s 8 .b6ui,tt c sa rpal colleocn2t 6e.dV l.n1e9aK1ro5 s h-Agach, joiinsdt a magjeudd;gfr ionmtg·h r ee latively P2= P 6T.h oeu tvearno efP 6w itdhi stinct smalleln ogftt hhne od ta malgeewftdi ntgh,e notocnth h teoi pn1 1b irwdist,hn ooutitc3n h specmiambyeea n9 . bir(d1s2 01121001,11 260,1 i2no2 n)me;o re Othpearr alec(tnoo4tys-p.1ew 5si) td,ha tab irtdhv,ia sni vese mruyc who rn. ont heliarb [eRluss:Ss EAi lat,Ca hiu,y skaya DiscuSs.cs t.ie olne.n igmsio tscitlc otaso e step1p7e0,0 -m1 9a0.s0 .I .n]e,aK ro sh­ S.cb.l yitnth hsiei aznepd l umcaogleo ration, Agac(h4d',),[ adA]l,nt o.1. 4( 20(55)d'),, buittu sp ppearra trlsei ghCtoemrb.i noaftion [lsA]ln,to .1. 5( 20(66)9),,[ lsA]ln,to .1. 6 ash-cgarspeh,ya rcpolnyt rwaistbthri onwgn (20174)[,2 7].VV.Pl e.r1e9v1(e5Zr,Iz SiPem,va ntnlooetn liynt hseu mmpelru mbaugte 120112200,1 12210,1 22/4(67l6)o- w9e6ur0s )u;a alllsiyon t hfree swhi ntpelru mage, Chagan-RBiuvrd'eg,r[ia ,uz vyA. lJnt,o. 1. 3 ,b rowngirseuhyp pteari l-acnogdvr eerytiss h 18[31).VPI.SI u.s1h9(k11i42n,0 119b/r4o6w(6nn- obtr owcne)n tpraaiolrf rectrices 960E)l;a nRgiavs[ehbr e twKeoesnh -Ag(aRcldh)i stinwgeultilhsm ihoseu sn tsauibns pe­ anCdh agan-RUizvue(nr8 d']),[, l sA]ln,to .. c iefrso mn ortS.h ecrn.c urranudcS a.c . 5(,9 d'),[ aAdl]n,to .6. 3, 0 .V[l129.1PV4.l, .b ]l.y Itnah did.i itnidoinv,wi idtaunh ao ltoscn h Sushkin (ZI1S2P0,1 1132/0416t61h-2oe9,u 6 tv0ea)rn;o e fP 6p redomiinS.n ac.te [NW MongoKlhiaar,a -lGoowbBeiar]k :u ­telenagnidt•iwbciiattr hhdt esio pfP 2s ituated MorRiinv (e1rd'0,,} [ adA]ln,to .7. ,( 119), neatrote hrte io pfP 7t hatnoP 6o.c cmuorr e [adA)ln,to .8. ,1 4[27).(V1ld'2.,)[1 a9d1]4fr,,e quently. Altn.o 1.0 (,1 d'3 ) [,l sA]l,tn .o 1.1 , Asc ompawrietS.dh c .h alimotdehnedr i, 20.VI[3.(V1I4d'l),)[ .l1sA9]l1.nt4 o.1,. 2 ,TelenWghiittieaitnlsh a rro(gaiwetnir n tga,i l, 21.VI[4.PV.SI uls)h.k1i9n11 42(,0Z 1I1Sa4Pn,td,a rsbuudsti) ff,e rientn hcseei soz feb ill 120111250,1 11260,1 11270,1 18/46n6o-t9be6ydS0 u)s;hakrineno cto nfirImnea dl.l (15r)eendesaA rc hNiutu Lra k[ew esteprlnumoafgS. cet.se lenugpiptteiarci al,-c overts shorjeu]vA,.l ,tn .o9 ., 1 7[30].PV.l.a1nR9dl1 a 4rg,er eyneodrta, rb kr owinns; u mmer Sush(k1i2n0 628/466-960).plumtahgmeea ,n itsslo em e\Vhatmdoarrek er, Thlee ctaonttdyh prepe aer aleccotlo­tgyrpeeywsii sthhy,oe ultl otwiiwnshthi,d c ihs tin­ lecitnme idd -Abuegaauvr se trfr ye swhi ntegru isihnedsi voifS.d cu.ha allsi moadelnsdroi plummaogset; oft(o8dpa on2td9 y )hp aevse i wni nptleurm age. as ummeprl umiabtge eic;no gn siwdoerrinan b lFyr otmh ree osfts outshuebrsnp S.e cc.ies, fivbei rcdosl lielnca tJteuedan nd ee ayrJ lulIyn. t elendgiifftieintrc hsdae a rukpeprse irad ned onbei irjndu vepnliulm(ea1 g2e0 r6e2m8i)g,ae lsse ox,c efoprS.t c .m argelianln aircgae,r anrde cthraivnceoes tst ill. compslieztee.d growth '(wlienng5g 4tm.hm6 t, alieln 3g8trn.hm7 ) b,u t Thvea li.dotifht miyos r pholroagtihcearl ly tarshusar s� acheds i(dz2ee0fim .nm6i)t t;hje eu ­wells eparsautbesdpw eac·nsi e evsde oru bt­ venmial(lee1 20i1fis1n 9i)st hhpieon sgt juveedn.i le 224 V.M.L oskTypoetsi: Sn y lcvuirara unS.cda a l th•Za OeOaS YRSOTS.S IVCoAl1 .0 Sylvciuar ruscnai girSetwasckhiain tohwoi,as s (iSsn igi1r9e2wa8sn)kcd io ,ui lnd­ 192-T9u rkmLeensisaenr Whitetchlrutodhaelit os c ailnti hrteay n ogfte h seu bspe­ cioenslb ya soentd h foel lowsipnegc iomfe ns S.cs.n igihraebswe segknii vaiesa nn erwe ­thZeI ScPo lleecxtaimoibnnyh e idim n1 926: placenmaemfonerSt ylcvuirart uucrak men(i1"c)'i'a[, l s3].,/ 16l.T,Ir Va.n1s9k1Sa tsap­ien, Snigir1e9w2ps7rk,ei o,c bcyuSy plivmeiydas ­tiDoonr t-KuMjeur vH[].nL, eo aurdn oon., tacteuar cmZeanriuc&dBa ni yl ke1v9i1c8h.1, 2 01(5Z"I 1S0P0,8 47/5W5i9n6-g29 .161,). Snigir(e1w9sh2ka7dis)e scrtihsbiuesbd ­ ta5i6l.t 3a,r1 s9u.bs5i fr,lo lmfe ath7e.br8is,l l speciinde estfr aoim1l 5 s pecicmoelnlse cfrtoemnd a r6e.sw9 i,d otfbh i 2l.ld6 e,p otfbh i ll int heea stpeaorrfntt h Kea raDkeusmae nrdt 2 .W7i.n fogr mu6l>a2 >t7io;pfP 2s ituated desigtnhhaeot leod(t p3y.7p )e. neatrtoe hrte io pfP 7o;u tvearno ent hteoo pf Holoty"p'i'ae d2..6 .VIInIe.Ra1er9p 2e5t.Pe 6kw itthr aocfen so tocnhl (y2.") 'i' [,l s], [TurkmeLneibsatpas(nk=fo,ay r am eCrh ar5­./18l.,IT Vr.a1n9s1k aSstpaitDeionor,nt ­ dzhousPkrayo3av°83),4 '6N°31,3 'nEo]2.,6 2 Kuju [neHa.rL oMuednroovn1.],2 , 0 79" inm y[ S.SIn.i girCeowlslkei(c]Zt IiSoPn(,"Z ISP, 100W8i4n86g/0 5.t52a95,i-3 l9. 131,). 100849/2I4na8 d-d9it2tot6i h)oe.dns a eto an, t ar1s9u.sb6 i,frl olm fe ath7e.rb7si, frl olm thoer iglianboaeftll h h eo lotthyepirease;l son ar6e.sw8 i,d otfbh i 2l.ld6 e,p otfbh i 2l.l7 . ani nscr"iSypltviciouanr rtuucrak menWiicnag fo7r m2=u;o l uatv earno ent hteo opf Snimga.d"be y S nigirienbw lsaIkcnikd iaPn6w iats hh alnlootwc h. intkh,le o cailsgi itvayes "n R epeTtreakn,s ­Thien scr"icplttoiokaso rena kumSeInSs"i s caspainatd"h n,ea moeft hceo lliegsci tvoer[n S =IS S.SIn.i girmeawdbseypk ein]oc nti hle ("CSo.lSIln..i girTehwoesr kiign"iu)nm.ab leb ra cokfo riglianbiaensllo sst u ffitcosi teantte ofS nigirewskiisl' asc kciionnlsglt,eet cahtdtai htoe tnsw efeo m albeesl toont ght ey psee ­ theirasle o canluimtyb( e"rLN o6 e7."W )i.n gr ieNsoo .t hseirm ilar wshpiecccohiu mledn s 62.t7a5,i3 l.t 6a,r2 s0u.bs4i ,frl olmfe athebresw itcho nfidence asdp easriagtyonpfae tse d 8.b4i,frl olmn ar6e.ws8i ,d otfbh i 2l.ld6 e,p tS.hc .s nigiwreerwfoesu kniidnti h ZeI ScPo l­ ofb i3l.lW0 i.n fogr mu7l>a2 t>8io,pfP 2s itul­ection. atneedta irop fP 7 t;ho eu tvearno ent hteoo pf DiscusVsaiuroine. (1h9a5cs4a ,s t1 959) P6w idtihs tniontPccuthrw .eh ictoel oonRu 6r douobntt h vea liodfti htsiyus b spVeocli­es, I wedlelv elaonpodec dc unpeiaetrshlw eyh olec han(e1t9sp5ku4iit) tn sa mient hsey nonymy outvearna enm do rteh uapnp hearol ftf h ien ­witS.h c .h alimoadnePdno drrit,(e 1n9k6o0 ) nevra nweh;i stpeo atrspe r esoenan pti cialn cltuhdweeh doT luer kmientn hiear oafn ge parotfsRa 5n Rd4 . S.cj.a xarvtailcioadf;S.i ct.ys nigirewskii ParatyApcecso.rt doti heneng t irtny hr ee g­waasc ceopntlbeyydfe wt axon(oKmoirset­s istrbaotooikfto hnOe r nithDoelpoagryt mleon1vt9,,7 2). onltyh eh olothyapsbe e egni vebny Howevtehrfoi,rs m d iffdeirsst iinnic ttsl y SnigitroZe IwSs(Pko1 in7 .V.H1i9os2t 6h)e.mr o rphoalnlodig hfeyi stIotprsly u.m age col­ collemcatdiieonK n asr aiknus mu mmaenrd oratinisoo tni cleiagbthlhtyiae nntr h ree osft autu1m9n2w 5e rree cebiytv heZedo ologmiocranelo rtfohremrasnsl, i gahsi tS.nc m.i nula, Museoufmt hMeo scUonwi ve(rRsuisttyba u­ttht ey poifS.c amlci .n ouclhatr ioenn tt h e mov1,9 5A4m)o.n tgh etmh,ea rrteew oot heurp ppearor ftt h beo daynfl da nikassl maobs­t femaloefLs e ssWehri tetchorlolaebtcy t seedne ts,p eciinta hsleul mym pelru mS.a gc.e . Snigiroen1w 8as nk2di3 A ugustn e1a9rs2 n5i gidrieffwweserklsil i o frtohsmeu rb species Repe(tceokl lneucmtbieRor-ns3 57a2n8d ofS. curriuntc hpaeu rleisggthr tce oyl oofu r .5 732 S5u)d.i lo(1v 9s7ke2ar)yr ao nleiosutsmeladyn tilnae l pll umaIgnea sd.d iS.ti co .n, thsee coonftd h ebsiera dsas y ptea,nK da ­snigirhearwses lkatiisivh eoalrnytrd o unded lya(k2i0n0 r1e)p etahtmieisds thaakvei,n wgi nagsi, Sn c. m. i nula. cittehd·is sp eciamste hnhe o lotByopteh. A ccortdomi yno gb servSa.ct .si noingsi,­ speciimnZe MnMsUc olleacritenfai coptna ­rewsiksait iy pidcwaello lfse arn ddeys erts ratyOpfte hs1e.4 p aratsyibpxie rsfrd,os m usuailnlhya bsittaibnidglu inwseiestdHa h ­ Karawkeurrmee cebiySv neidg iornel wosankl io xyalpohny lHl puemr,sa incodut mhp esra m­ fromt hper icvoaltlee ocfMt airWoi.neS tromm ophitlroeauensssd h ruIbtns e.sj tusis nt (Snigi1r9e23w75s:)k .i , suchha biatlasitonsv icionfiR teyp etek Ina l atpeurb licSantiigoinr(,e1 w9s2k9(i:S op1y9ev6t,7h ;ae u thoobrs'esr viatni ons 260n)ottehdS.a ct.t urkmiedsni isctar i1b9u5tT8eh)dbe.i radrsrt iotv hKeea raaklurme ady in" EasptaeorrfTtn u rkmetnoMi esrtovaa n­,i nb etw1e0etanhn 1 d2 tMha racnhbd e gtion siisnt hwee sHto"w.e vheehr a,ns o vti sitberdeeeadr cloym:p lfreetscehl utwcehrees ZOOSYRSOTS.S ICA1 0•V V. oMl L.os kTypoetsin: S ylcvuirar auncSdaa. l thaea 225 founidnt hfier tsetn -pdearoyifA o pdr (iRlu sP-7o;u tvearno ent hteo opfP 6w itadh i stinct tamo1v9,5 S4o;p y1e9v6,I7h )a.vm eefl te dg-notTchhwe.h iatreoe naR 6i wse dlelv eloped linagnosdb secrovpeudl oafat diuowlnht isc,ha nodc cumpoirteehs ha anol ftf h ien nvearn e; begtahn_e s ecnoensdtc iynciglnHa e l,o xy'l·ao s nm awleld ge-wshhiatpsiepsdoih tsp resent des·ew roto dlnaenadDrsz heb(e3°3l94 'No,n a pipcaaroltf sR 5. 5°41'lE)o n2 2.V.T1h9e7ds9ae.ta ana,dl as o In addittoti hoheno lotyIhp aev,foe u dn finodf an ewsitt fohu erg gisnC hil'maomtehd1e-5br i rfrdosmt htyep es erdiuetesot, h e kumo n1 2.IV.(1R9u4s6t a1m9o5ev4v,)i -inscr"iSpytlcivuoirnar ucaJSaxnaidrge.t.t i ca dentcheSa .cts . n igiirnehwassbkaiindtides y- SerS.n igir1e9w2so6kn"ti h bea cokft he seratloslv etrhs eo uotfTh u rkmewneisstto arni,go iron laldla boeftl hsZe I ScPo.l lection. upt toh Cea spsihaonr e. Apparoenntl1ly2by i,ra dmso ntgh erme ally Thmee ntidoinffeedr ientn hcmeeo sr phoble-lotnotg hp eo pulianthiaob'nit lthoiewn egr ogayn ldi hfei stoofTr uyr kmebniiraradens SyrdavraylalT ehyer. e maitnhirsnekgei ns imporatniadnn dti tchdaeit set ionfct thniesws ist hqouuetsb teilotonomn igg raitnidnigv idu- formfr omS. c.m inucloan,t troWa orlyt earlsos fo thseurb sp(escbeieeel so w). (198a0n)ed,s pecfrioamS.l c.lax yaJ rtaisc a,Pa rtypaesb elontgoiS. n cg.J axartica. oppotsoSe tde pa(n1y9a17n98 9,W0 a)t,s oKna zakhstanR,i vS[eybrre dtawKrez'eyyn-a (198a6n)Rd,o se(l1a9a9rR2 e)c.o gnoift Jioornad naKd a zalinscko]lE,l. E. v1 e8r5s3-, thseu bspS.e cc.si neisg iriesw wesljklui sitm ia-n(n1,s) e ?x[, alda,st uem m(e2rs)]e ,?x , tied. [adl,a stuem m(eZr]I S1P0,0 808, 100809); colNl..AS .e ver(t3scf)o,[ v a,d1 ]7,[ 29].VII. Sylcvuirar juacxaa rStniicgai r1e9w2-s9k 1i8,5 (84,cf) , [ ad1]8,[ 30].V(IZII.S1P8, 58 SyrdLaryeas Wsheirt ethroat 100810010,8 (054cf)), ,[ ls9][,2 1].Vl.1858 (ZI1S0P0,7 (9699)),[ , l w1]4,[ 26].VII.1858 Thseu bspiensca imeaesftd eJ ra xtahrtae,n ­(ZIS1P0,0 7(976cf)), [, l w2]3,. VII[4.VIIIJ. chineanmoteft hSey rdRaryiav er. 185(8Z I1S0P0,8 (085cf)),[ , l w1]7,[ 29].VIII. Snigirheabwsrs ikedifle ys crtihsbiuesbd ­185(8Z I1S0P0,8 (093d')],[j , u v1.3J[,2 5].VI. specfrioem1s 7s pecimZeInScsPo lolfe c1t8i5o(8nZ ,I 1S0P0,8 (0120cf)),,[j uv(.1]1cf,), mainfrloymt hleo wSeyrr �Rairyvave arl le[jyu v1.6J.,V I.[12885(.8ZV IIS.1P]0, 0 797, [betwPeeerno( v=sK kz yloarnddKa a)z a­10080(01)2cf,),[i uv./1l4w[]2,6 ].VII.1858 linasnkdd]e ,s igtnhhaeot leodt ype. (ZI1S0P0,8 06). Holot"ycfap de2..4 .VII[.O1lS8dt5 y8l e!F]i,vp ea raty(p1e00s8 01-1100008r80e06­4), Ssyr-DCaorljSlas.,e wet[zSoewv ertstoavNi].n,AS .e verotrsiogvli'ansboa enoll l p da ­ Origina1l0n2u(6ms7bnu eobrm m .i nublea­)p;e rin;t hree osftt heimti, sr eplbayca e d findliimZc ohoM !u.sA .c add.W. i sseinns csht.a nodlaZdrI dSl Pa bweiltr ho unldeeaftdn d Peters(bZuIrS1gP0",0 79T8h)eo. r igirniagelhdtg es. , Severltasboievsll' a sc kbiunttgh ,ea rranee S.cJ.ax aritasic coam parlatairvfogerelm y: oladn adn elwa boefZl I ScPo lleOcntt hieoi nnm. a lfreosm t htey psee rwiiens(g,n= 8 ) facoef t hfier lsatb ienal d,d ittoti hoaenb ov6e 4.9±1.7 t(a6(i2nl=. 111-5)68 7..43±)2,. 0 dattah,ei rasen . insc"rSiyplmtviiinoaun s c(u5l4a.,4 -t6a1r(.sn3u= )s 1, 1 1)9 .8(±108..57 - Humea"n odn,t hbea ctkho,er igniunmable 2r0 .b4i)l,l fefartohm(e n=r1 s08 ). 1±(07-..46 ofS everctoslolve'"cnsto 8i.7co 2.Sn sa.n"d 8.8b)ifr,llo mn ar(en=s1 06). 8±0;1 (6.6-6.9). inscr"iSpytlcivuoirnarJ uaxcaar Stniicdgae. t .W infogr mu6l>a2:(> n=71 )P,2= P 7( n=4 ), SeSrn.i gi1r29e6wm"sa kdbieyS nigiirne w7s>k2i(> n=82 )a nPd2= P S( n=1 )o;u tvearn e blaIcnkd iinalkn;a ttehrwe,o r"dT ypiuss "o nt hteoo pfP 6w iatd hi stniont(ccn=th8 ). addedb yatd hieffreehr aennIdntt. h e lnae­wP aratynpoebtse lontgoS.i cnJ.gax artica. betlh,de a otfce o llectthbiein irggsd i ovifen n O neb iridd entbiySfi neidg iraesS.w cs.k i thNee wS tyl(e5 .VIII). Jaxartwiictlaha, b" eSly rdRairv9ye,a[r l ,w ], Thutsho,er igniunmable r g(i1vb0ey2n 6 270). V[III.I1I.8X 5.N8].A,. .S ever(tZzIoSvP", Snigiirtseh woasfttk h ioe l ZdI ScPo llec1t0i0o7nb0,e8 l)ot,npo ga ss sS.i cnb.gl yTthhiiiss. ratthhe·aronS f e verctoslolve'Jcsut digoinen.vg i debnytc hetedy piccoallo roafut pirpoen fromt hpel umtahgaeeg o,ef t hbei hradbs e enp arrtusfo:bu rso nwanpo eft hsea mceo laosur deterimniancecdu iritas nt oe"tla myda: l e"m,a nt(ltelh aet htaemsra rkemdolriyen tense buaty earilnaiv negrfr ye swhi nptleurm agceo lortahtaiinnto hnoe t hsepre ciomfte hnes [lwW]i.n 6g7 .t3a,5i 8l. t6a,r 1s9u.sb6 i,l tly speer aineudsp )pt earci olv ertssl iognhltyl y fromfe ath8e.rb6si,fr l olmn ar6e.sw9 i,d otfh d artkheamrna ntIlnae d.d iPt2i= oP n6a, n d bi2l.ld5 e,p otfbh i 3l.lW0 i.n fogr muPl2=a thneo tocnth h oeu tvearno efP 6i asb seIntt. VML oskTypoetsi: Sn y lcvuirara unScda.a / th•Za OeOa SYRSOTS.S VIoC1l.A0 226 shobueln do ttehdta htwe h oulpep seirad ned Amudavraylaal nemdye ntiboySn neidg irew­ flanokfts h biishr adva eni nternussbetr yo wns k(i1 922597a:)s c lotsoS.e c h.a limodendri, tint,iwushn iucsfohurt a hlse p ec(iperso baabrlaeyc ,t ucalloilstnyeh s ei tzSoec. j. ax artIi ca. iti sa ni ndivwiidtauhba elr crharnotm,i csotuinlcodfi t n adn dyi fferientn hcpeel su mage coloration). coloroafclt ,ai do1,n7 .V(IZIII1S.0P 0,8 25), Furtthwebori riddesn tbiyfiS endi girewwhsiikcfisihn isphoisntgb rmeoeudfrliotnmS.,g asS. cj.ax artwiictlaha, b" eKlysz yDlek­ucmj. axaritnti hsceaa mp el umOatghese.er v en seSrta,r bai[-vbiucloiyfTns aihmt dyy b(u=l maakl aensod n feem alienfr esfihr .swti nter Tamdayc)c,o rtdoai rncgh .diatvoaen t hep lumdaigffefre ormfo urs peciomfS.e cn.s Glazunroovui'tnse1 8922] s,e ?x, [ ls]j,ax artiintc hase a mpel umoangleiy nt he 19.IV[l.DV..K]G..l 1a8z9u2n(,oZ vI"S Ps,omewlheaostcs h raucpepopeuarsr T thsu.s , 100752, b1e0l0ot7no5pg 3a )s sS.i ncg. thdee lotfta hA em udaRriyviaeps rr obianb­ly minuTlhaii.sse videbnytc hevede rtyy pichaalbibtyS.e cdj. ax arotrip coap ulianttieorn­s oft hsiusb spceoclioerosaft t huiepo pnse.ir d emeditaoS.t ce.s nigirAedwdsiktdiiaiot.na al ofb odfiyr,as ntfod r emtohsseth acropn troans tdtihset rnoi fLb eustrWsi heoiteittnhh reoat betwleiegbnhl tug irse"hyc..a a pn"yd e llowlioswher Avmauldalarenreyey cae stsoda ertye r­ ligbhrto mwann tglree,yb irsohuw pnp teari mline pmroerceit shbeeol uyn dbaertyw een covearnatdls ss,mo a sliloz fbe i r(dwsi5 n8g. 6thessueb species. an6d0 .I2it)ps .o sstihbtalhtdee e scroifption thpel umcaogleo riantht eoi roing diinaagln Soy­lcvuirar cuhcuaa ncPhoertiec1na9k 5o-0, siwsas i nfluebnyic necdl oufst ihoebnsi er dAslashLaensiWsahenir t ethroat int htey speer "iDeiFseii.: r biuswntigb e e S.i cm.in.u".l( .aS nigi1r9e22w95s:9k )i., Whedne scrtihbsiiusnb gs pecies, Portenko DiscuTshsbeii ornfrd.os mt hleo wSeyrr ­(1955055:h) ae sx ami1n6se pde cicmoeln­s darvyaal dlieffyce orn sidfreormS.a bcl. yl ecbtyeN d. MP.r zewalsakniMd,. EG.. E. minunlooatn liynt hlera gseirz beui,tnt heG rum-GrzahniPdm. aKKi.ol zoli,onC v h ina plumcaogleo raaswt eilTolhnt.e y speer iine­s( upcpoeuror fts heHe u aHneg( YelRliovwe)r clud(e1fos)u: ar d ublitrfi dnsi sphoisntgb reed­ antdh Ael asMhtasan.n )hd a dse sigtnhaet ed inmgo u(l7t- s9ttahg( e2fos)u) yr,e ar(liinn­gs holoatnyodpn peea ratype. cludtihhneog l otiynv peerfr)ye sfihr wsitn ter Holoty"pcl,e[ .a d], U[pCphHeiurna an]g, plum(a3go)enm ,ea liern a twhoernrfi rssutm ­ Hen,e laorc aGloimMtiay,y1 88(0O lSdt yle), meprl um(a4 g)te h,rm eael ejsu viepnnl iulme­ N.MP.r zew(aZlIs1Sk6Pi3,"7 65/4O2n5-974). agaen,(d 5 o)n mea l(eri 1o0.0 8i0np6 o)s tju­ thbea cokft hoer iglianbaell i,sa nti hne­re venmiolue(l mto trheha anol fsf m afelalt hers scrimpatdibeoyP n o rteinnrk eoid n "kS:y lvia arnee w). currcuhcuaa ncPhoeritc1e9an5 kT3oy. p us!". Thuep ppearro tfys e arliinfrn egsswh i nter Win6g8 .t1a,6i 0l. t4a,r 2s0u.sb6 i,frl olm plumaargteeh m eo srtu foaunshd a vaed is­ feath8e.rb2si,fr l olmn ar6e.sw8 i,d otfbh i ll tioncchtr aceTohutisisi n asttb i snietjnn.u t v e­ 2.d6e,p otfbh i 3l.lW0 i.n g formutliap 6>2>7; nilaenasdd ublitri dnfrs e swhi nptleurm age; ofP 2s itunaetaetrdoet rht ei opfP 6o;u ter theuiprpp earra trssei mprluyfob urso winn, vanoent hteo opfP 6w itdhi stniontTcchthe . adublitra dlsss oom ewdhaartkT ehrue.p per whiatreoe naR 6i wse ldle velaonpodec dc u­ parotfts hm ea liewn o rfinrs sutm mpelru mage piemso rteh ahna loffi nnvearn ael ;a rge arlei gbhrto wngirsewhyi ,t hyoeultlo orw wedge-wshhiatspipesiodpsht r esoenan pti cal ochratcienotu.s parotfRs 5 o;n lawy h itsitsrihips pr ee soennt ThuS.s c,j. ax artdiicffaefr romsn orthern thteio pfP 4. subspienccileS.us cd,h.i a nlgi moadennSdd. ri Paraty"p<;?[e,l. sH ]u,aH neg[, l ocGaulyi­ty c.te/enignti htleii cgauh,pt peperar rti sna ll Duy1]4,. VIIG..a1 n8Md9. 0G ,r um-Grzhi­ plumaangdea,ss a r ulient, hm eo rreo unded mail(oZ"I 1S0P0,6 T2h9oe)r .i glianbiaesll winagnsfrd,o m.soSuc.ts. hn eirgni arnedwskii lackointn hbgea, cokft hoelZ dI SP tlhaebreel S.c.m inuilnta h,le a rsgiezSren .i girewski iasn i nscrbiyPp otritoienrnn e kidon " kS:y lvia (1922599s:)u ggetshS.ta cetj.d ax armtuiscta currcuhcuaa ncPhoeritce1an9 k5Po3a .r aty­ difffreormS. c.m argelaanswi eclaalsfr, o m pusW!i"n.6g 5 .t4a,5i 6l. t9a,r 1s9u.sb6 i,l l· S.cm.i niunlta hl eo nbgieTlrhl pe.l ausibility fromfe ath8e.rb1si,fr l olmn ar6e.sw6 i,d otfh oft hedsieff erweinblcelee xsa miinnme odr e bi2l.ld5 e,p otfbh i 2l.lW9 i.n fogr muPi6=a ·detianti hfoler thcormeivniogsfs i uobns pecies ofSc .u rruca. P2o;u tvearno ent hteo opfP 6w itthr aocfe s Ninbei rcdosl lbeycN t.Se edv eritnts hoevn otch Whoinatlreyeo .an R 6o ccupies not sechoanoldfAf u gunseNtau rk iunts h e lomwoertreh oanne -otfih ninrvedar n e. ZOOSYRSOTS.S VIoClA. S. 227 JO•V. ML oskTypoetsi: Sn y lcvuirara uncdaa / thaea Ina ddiIth iaovne,. ex1a2tm oipnoety7dp esI:wt o ublevd e irym porttoka nnowtwh eirse <:!,3 a�n d 2juovf.t hMeomcs,otwl elirenec ttehndee stairneogafb irddess crfriobmte wdo AprainlMd a yo,n loyn ea n;bdQ'!o tjuhv enmiilger ating assS pyelcmviaimraeg nesl anica biridnJs u,lT yh.eb sierar dets h e-lianSr .g Setsotl z1m8a9n(7n2d' , ,U zbekiFsetragna,n a, curriunmc aal:(e isn cltuhhdeoi lnognt yp=ne e,a Mra rge3l.aXn.,1a 8n95d3. IV.T1.8 94, 8)w,i n6g8 .5(±615.;66 -t7a60i0.l.9 4)±,1B .a9r eUyn)fo.r tuntahlteoe clayot,fit ohne se (58.5-t6a3r2.s06u.)s7, (± 02.04. 1-b2i1l.ls4 y)n,tr yepmeasi ns.Tuhnewkyen rkoeew ipnnt. fromfe ath7e.r7s±( 07..34 b-i8frl.ol2mn ) a,r eBsr aniMtusskeiui'mnsW arszTahwoau.g h SushakniSdnn igirheawvnseoke tix amined 6.6±(06..32 -w7i.do2tfb)h i, 2l .l5 ±(02..13 - thtyep sep eciomfS.e cn.ms a rgeltahneiyc a, 2.6d)e,pot fbh i 2l.l9 ±(02..17 W-i3n.fog0r )-. consitdhefoirrsemt d ob ew iddeilsyt riinb uted mul6a> 2>(7n= 6)p redomiinnfia vtee s; NorCthhi Snnai.g irreegwasirkatdisn e e ds ting birtdiospfP, 2 b eisnigt unaetaetrdotei rpo f fromL obnLoark(= e L oNpu ri)nt hwee stto P6i;no nmea lPe2=, P 6i;no nmeo rmea le thAel asMhoaunn tRaainangn eOd r dionts h e P2= P 7o;u tvearno ent hteo opfP 6w idthi s- easatnfr,do m4 3°Ni nt hneo rttoTh s aidamin tinnocttic nfih v mea lweisst,hh a lnlootiwcn h thseo uStuhs.h iknicnl iunid tensde stairnega twmoa lweist,h nooutitcno h n mea lIenfo. u r altshoKe u nlMuonu ntwaeistntots hP ea mirs. femal(eisn clpuadriantwgyi pn6eg2) .6,15 ,. 4, � andd' collebcyPt rezde wianlt shkei 66.515,. 55;6 .5t66a,.5i 98l,.5 39,.t la;r1 s9u.s6 , Russ(k= iKya xtSahxa Mno)u ntRaainng e 20.220,.2 60,.b 7ifr;Uo mn ar6e.3s3. ,66 .,9 . (ZI1S0P0,5 12070,5 i2nM8 ay) a nadm oult- Thrmeael ceosl l.eicAntp eradin lMd a yi n in<:!g, a dc ollbecyMt .eVPd.e vtisnto hve EastTesrani( d=Qa uma idbayPm .)KK .o zlov don odti fffreorm ttyhpee osf cs..c huanK-halaRsitvavenar l luepypT,ei rz nHaeip n Augu1s8t8( 9Z IS1P0,0 5e3v1i)d etnhcaet cheiintc hape l umage'acnoadlr ooern altyi on nestoiftn hlgia srfo gremi nn ortfohoetrhni lls somewshmaatli lnte hrae v erdaigmee nsions ofKunilpsuo ns sMiybe lxea.m inoaftt hieosne (wi6n6g.6 16,. 626,.t 6a5;i7 l.5 48,.5 39,. 2), busti gnifilcarantgthels.arycn m. i nula. specismheotnwhssta htde oyn odti ffseirg nifi- canfrtolmSy c. c. h uancheica. Int hed ifferednitaigan(loP soirst enko, Inc onttrota hsoetp inoifSo uns hankdi n 1955S).c, .c huancwhaesdi icsat inguished Snigirews(k1i9,s5 u5Pg)og rettsheS.tan etkd o fromS .c m,i naunlS.da c m.a rgelSatnoilcza- c.margeliasnn eisctoain nligynt hEea stern man1n8,9 7. DiscusTshiedo ens.c rbiybP eodr ten(kCoh inTeiseen)-T Shhicason n.c lcuosubileodn (19d5i5ff)e rientn hcpeel su mcaogleo rabtaisooennad s infegmlaecl oel lbeycP trezde wal- anddi menbseitownS.es e cn.c huanchsekiiicnt a hC eh aid(u= -KGaoiHlde uv) a lilney (bifrrodmts h seo uth-peaaorsfttt heserp ne -May( OlSdty l1e8)7( 7Z I1S0P0,8 T5h8ee)x .- cireasn agneS.d) c m.i nluolqoauk i ttraeun ed acdta otfec olleocftt hibinisigr usd n known, werwee lkln owlno nagg( os Senei girebwusttkh iee,x peddiitaiironynd ictahtateths e 192295:9 -R2e6a0lb)li.yr i,dd se natsS.i cfi.e dl owreera cohfCe hsa idwue-rGceor lo sisned chuancahrmeeui clchaa ragneidrns prainndg t hvee brey ginonfMi any(g O lSdt yfolret )h;i s summpelru mhaagvdeea rukpeprpe arrtw si thr-easioctno ,ub lead m igrianndti vTihdiusal. outty poifcS. a cl.m inuyleal lotwiiTnshthe. femalwea sa lwaaysss itgons. e cd.m ar- differienns ciezasere es pecnioatlilcye gaebllaein itims co ass;ti miintlh aperl umcaogle- witcho nsidfoerrt ahtseie ooxfn b ir(dsse eo r·a tsiioaz·ndne ,w · m gfo rmu(lPa=2 P 6t)o abovIeitp)s o. s stihbWatle es,tT esrani idsa m thtey pSe.cs .c huancihnce oircrae sponding inhabbyi tbeoidfir ndtse rmpehdeinaottey pes. plumage. Buitst h obueln do taeldts hota htae v erage s. Altlh ree maibniirfrndogsmt hZeI ScPo l- winlge nignt chc. h itanictshh seeai macesa lectiona sSi cdm.ea nrgteilfia(en9d<:fi a cnad iS.nc m.a rgel(aPnoirct1ae9 n55k50o:t5, a, b . 5� )a nedx amibnyPe odr taernaeku ot ummin- 11A)p.p artehndetoyln yo.d,t i ffaelris nto h e gratnwtoosft : h ewme rceo llienSc etpetde m- plumcaogleo rsaitn"icboern o,w gnrietsyih n t ofb acakns dh oulmdeenrtsib"oyP n oertde nkhoeo rn,oe n,l. XaIn,td h ree isnOtc ,t onbienre; forS. cc.h uancahsde iisctai nigfrtuo ims hsipnegc iwmeercneos l lieFnce tregdLa anraig.ne - s.c .m argeliasan siscoacw iiatsthee ads odniavli pdrueavlbasui ttlh p,el umcaogleo ration varioaftp ilounm argaetsth,he anwr i tghe o-variaenstd,h rbeier sdhso ublead s signed, grapvhairci aAtlbilio rnod.fS.s c:c huanw-ithqouuets ttoiv oanr,io otuhsseu rb species. cheiecxaa mibnyPe odrt enakrioen w orn Partiactutleasnrht oiubolengd i vteoan d' , sprainnsdgu mmfeera twhheir1ls4eo, f 1 5 [lwC]h,i nnao,r thern ofofNo atnhsilhlasn biradsss ibgynh eidtm o S ·.c .m argelaniMctasS .y,d uBne,i s2h1a.n1I,8X 9.G0 .a, n Md. (ZIcSoPl leacritefrni e oswnhi) n ptleurm agGer.u m-Grz(hZiIm1Sa0Pi0,l8 Oo6n 5t )hb.ea ck 228 V.ML oskTypoetsi: Syn l vciuar raunScda a lt ha•e aZ OORSOYSSCSTAVI. o Il .0 oft hoel ZdI SlPa btehlei,rsa e ni nscribpit2li.lod9 ne, p otfbh i 2l.lW2 i.n fogr muPl2=a "marge(lSaInmSia)cd"bae y S .SIn.i girePw7so;ku itv earno ent hteo opfP 6w itdhi stinct inp enLcairldg.ie m enosfti hobeni s(r wdi nngo tTchhwe.h itairsoehnaR 6o ccuapbioeust 68.t4a,6i 3l. w7i)n,fog r mu(lPa=2 P 6a)n d haolffi nnvearna ew ;h itsipsoihtsp resent upppearrm tusc lhi gthhteatrnh oisnSe .c . alosnoa pipcaartolsf Ra5v ;e nrayr wrhoiwt ­ blytihnid itchaatitetb elotnogS. s c. issht roinplpeyr esoentn heted goeft hien ner chuancPhoeritcheaan.ssk e oet nhb iisbr udt v anoeft huep ptehrio rfPd 4T .h pea raitasy pe dindom te ntiitIoi.tnps o sstihbhaleate l ass­o year.lbiiinrngvd e rfrye swhi ntpelrum age sigintteSo d. c m .a rgeltahnoituchgmaiha,s l es ooanft tehrpe o stjuvenile moult. wacso llmeucctfaher dt thote hree a frsotmt he Among otfrhoemrw hibtcihhrse du sb spe­ East(eCrnh inTeiseen)-n Sehatarhnn e,o rthc­iewsa sd escrtihbfoeel dl,o wing specimen westbeorunn doafrt iheresa nogfeS. c. shoublend o t"e9da,:d T ,a jik[iBsatdaank,h ­ chuancohuetilbciyhan i emd. shanv]i,c ionfKi htoyr [o=Kg hho ru3g0h.]V,I . Thaeb odvaest hao twhV aatu (r1i9ew5 a9s) 193A6.,II v.a (nZoIv1S 0P0,6 68/1O9n5 -938)". corwrheechtnep latcheneda mceh uanchtehibeca aco kft hoer iglianboaefltl h biisr d, Porte1n9ko5i,5sn y nonwyihmmt ya rgelaniPcoar taednd"kemod o ntPiocroitlnb.al " iu netk o Stolz1m8a9n7n., thien scr"iSpytlaivloitnah b ayAe .aII"v. a nov inb laIcnkd iinakn. Sylvailat hmaoenat iPcoorltae1 n9k5-o5, DiscuTshsceio omnp.a orfbi isroidnsnsm u ­ MidAdslieMa onu nWthaiitne throat meprl umfraogmte h tyep see roifS.e a sm. o n­ ticwoiltafoh u rb reesdpiencg imens from Thsiusb spweacdsie essc arsiS byelcdvu ira­N orth-WIensd(ti3ecla r a ndfr omK ashmir rucmoan tifrcoom5l 0sa p ecicmoelnlsie nc taendcld l sfr omB alticsotnafitnrh)mde si ff­er thKeo petPdaamgihAr,l saa,in Tdi eSnh ane ncneostb eyPd o rteunpkppoea:rrt sm'uacrhe Mtsa.lfr,lo mt hZeI ScPo lletchhteoi loont;yd paeri knIe nrd biiarndd asrb,kr orwan,tt hhearn anpda rawtyeprede e siginntat heoedr igibnraolw ngiryse.Th h see paroaftt hiIeon nd ian description. Mountain WSh.ia tletthhareoaa ta,l thaea Holotyp"ecl[.,a d], [GTiasjMsitaksri. sH,tu amne1],8, 7 a8n dM iddAlsei Maonu ntain locaKlviaty3k 0,k mN· ofS tali[n=a bWahdi tetS.h ar.mo oantt,iP coorltae1 n9k5io5s, Dusha1n9b0em0] a ,s. I. .1 ,1J un1e9 4A8.,I .q uijtues tIitfis ehdo.ubl eod n lnoyt etdh at Ivanov"4 4{6Z9I/S2P4,A4 .-II9.v4 a8n)o.Pv o rt(e1n9k55o05 e6:)r ronienoculsBulady­e d wrotthese p ecniaem"seS ylavlitah iane al"o ch(iisntsaotnfBe aaldt iisnat dadnit)to,i on blaIcnkd iinaoknnt hbea cokft hoer iglian­aKals hmiintr h,re a nogfet hneo minotypical beble;l otwh,ei rsae n i nscrmiapdtbeiy o snu bspecies. Portienrne kidon "kS:y lviam ocnutrircuoclaa Porte1n9k5To3y,.p usW!i"n6.g8 .t2a5,i7 l. 9A,c knowledgements tar2s0u.sb8 i,frl olmfe ath1e0r.bs8i ,frl olm nar8e.sw7 i,d otfbh i 2l.ld9 e,p otfbh i 3l.l2 . Ia mg ratteJofu. KlMe .r zfohrnc eorn sulatnadt ions reatdhimena gn usTchrcJio pltlo.etf ch tZeio oonl ogical Winfgo rm6u>l2a>t 7io;pfP 2s itunaetaerd InstRiutsusAticeaa,nd oefSm cyi esnucpepsbo,yr ted -t-iopfP 6o;u rtv eanoent hteo opfP 6w ith thReu ssFieadne rMaitniioosfSnt c riyea nnTcdee c h­ notTchhpe.u wrhei atreoe nRa 6 i wse dlelv el­ nolgorgaynn.o2t 0 00-w0a3us-s 1e6d . opeadno dc cumpoiretesh aonn e-otfht ihred innvearna es ;m awlhli tsipsiohspt r esoenn t References apipcaarlot fRs 5 a;n arrow swthriiistp ies h onlpyr esoentn iotpf P 4T.h heo loitsay np e AlSi.,& RiplSe.yD1,.9 7W3a.r btloRe erdss tarts. adumlatli ewn o rsnu mmpelru mage. Handboooftk hb ei rodsfI ndainaPd a ki8s2.t7a 7n, Paraty"p9je,u. [v w1.] T,a jikliebsftat nakn ,p B.ombeatyc,. ofK hingou RiverT,a v'di-olDw an·rBs aatk reKer.1a, 9 m9Wa 7ro.b flo efEr usr oApseia,an No dr th (=T abi-[D=Ta avrial)' 3d°84ar4a',7N °03,0 'E]Afr,i c4a0.p 0L. o ndon. 14.VIIAI..II1v.9a 3n(5oZ,vI "1S 0P0,6 74/C2h2e8nT-g.,H1 .9 8A7 s.y noopfst ihasev ifaoufn a 936O)n. btahcoekft hoer iglianbiaenall d ,d i­Chiin-ax:1 v-i1,2B 2e3i.j ing. titooin d enti"fiScyalatvliitoahn( a ibenla a"c EktchecRo.pD&a. Hr i,l Fe.1, 9 8L3e.Os i sedaeu x ChidneeMo ,n goelidte eC orPiaises.e 7re0pa7.ux . Indiinabkny)A .IIv.a ntohvei,ras en i nscrip­ Paris. timoand beyP ortienrn ekidon "kS:y lcvuira­H affeJr.1, 9 9S1y.lc vuirar Guecoag.r aVpahriis­che rucmao ntiPcoorlat e1n9k5Po3a,.r atypuast!iI"onG.n:l. uv toBznl otzUh.eN&iB. ma ,uK e.rM,. Win6g9 .t4a6,i0 l.t 2a,r2 s0u.cs6u ,l mfreonm (BeaHarnbd.b)ude.cr hV ogMietlt rleu1r2o/1p1a,s , feath9e.rc6su, l mfreonmn ar8e:sw1 i,d otfh Passer(i3Tfo.er i7m!9e)5s:- W 7i9e6s.b aden.

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