Miscellaneous notes on Southeast Asian Gramineae. VIII. New species in Danthonia

3 Pages·1993·0.39 MB·English

by  Veldkamp, J F

BLUMEA 38 (1993) 217-219
Miscellaneousnoteson SoutheastAsianGramineae.VIII.
New species in Danthonia
J.F.Veldkamp
Rijksherbarium/Hortus Botanicus,Leiden,TheNetherlands
Summary
Ofthegenus Danthonia DC. (Gramineae)threenewspeciesand onenewcombination aredescribed
here.
Danthonia DC.
Danthonia bonthainica(Jansen) Veldk., comb.& stat. nov.
Danthonia pilosavar.bonthainica Jansen,Reinwardtia 2(1953)258. —Notodanthonia penicillata
subsp.bonthainica Veldk., Taxon 29 (1980) 298. —Type:Biinnemeijer11971 (BO holo;L).
Danthonia spec.: Hemsley,KewBull. (1896)36.
Culmstufted,branching mainly extra-vaginally atbase,0.12-0.5m long. Blades3—
15cm by 0.4-0.8mm, glabrous, ultimately articulating withthe sheath. Peduncle
glabrous. Inflorescence with3-13 spikelets, contracted, 2-6 cm long, axes pilose.
Spikelets chasmogamous, 4-6-flowered, 6.5-9 mm long (excl. awns). Glumes
subequal,6-9 mmlong, slightly shorterthanthe spikelet (tips ofupperlemmasand
awns protruding), glabrous. Lemmamargins with2 tuftsof hairs, hairs 1.25-1.5
mmlong, andwith arow ofa few inconspicuous few-haired tuftsabove thecallus
(obscured by the callushairs); body 3-3.5 mm long; laterallobes 2-3 mmlong,
0.5-0.92timesas long as body; setae 1.5-3.5mmlong, 0.67-1.1timesas long as
flatportion oflaterallobes; awn7-10mmlong, exceeding setae by 3-4mm(when
wet), with 1-2basal twists. Paleamargin (inside) withouttuftsofhairs. Anthers 1-
1.5 mmlong.
Distribution- Celebes (Bonthain).
Habitat- Insubalpine coppices, disturbedplaces, 2500-2890maltitude.
Note- Ms.M. Gies (FR, in litt.)reported thepresenceofaxillary cleistogamous
spikelets inBiinnemeijer 12260(L). Ihaveseenthesheet, butnot thespikelets.
Specimensseen:Biinnemeijer11971 (T), 12210,12260;vanderPijl 767-a.
D Continuedfrom Blumea 37 (1992)237.
218 BLUMEA Vol. 38,No. 1, 1993
Danthonia dendeniwaeVeldk., spec. nov.
Danthoniaspec. nov:. Stevens& Veldk.,Bot.Bull. Lae 10(1980)31.
Foliapilosa,pedunculi distaliterpilosi, glumae6-8 mm longae spiculas plusminusveaequantes,
lemmataprimasolum caespitibus(sub)marginalibusdistinctis,corpore 2.5-3mm longo,setis 1-
1.25 mm longis, loborum partibus applanatis 0.5-0.63-plo longioribus, aristis 5.25-7 mm
longis,columna brunnea spiris basalibus 0-1. Margopaleae(intus)pilorum caespitibus.—Typus:
Veldkamp & Stevens 5768 (L holo), Papua New Guinea, Northern Prov., Mt Suckling, Goe
Dendeniwa,26 June 1972,3550 m.
Culmstufted, branching extra-vaginally at base, 0.09-0.16mlong. Blades5.5-11
cm by 0.6-1 mm,pilose, articulating withthe sheath.Peduncle distallypilose. In-
florescencewith4or5(-7)spikelets, contracted, 1.5-2.5cmlong, axespilose. Spike-
letschasmogamous, 4-flowered, 7-9 mmlong (excl. awns). Glumessubequal, 6-8
mmlong, shorterthantoas longas thespikelet (then onlythe awnsprotruding), gla-
brous. Lemmawith 2tufts ofmarginal 1-1.5 mmlong hairs,with 1 tuftofshorter
hairsnext totheupperoneand withoutany abovethecallus; body2.5-3mmlong;
laterallobesfusedwith thecolumnor free, thenup to2.5 mm long,0.6-0.9 times
as long as body; setae 1-1.25 mm long, 0.5-0.63timesas long as flat portion of
laterallobes; awn 5.25-7mmlong, exceeding setae by 1.8-3 mm, with 0-1 basal
twists. Palea margins (inside) about halfway with atuftofc. 1 mm long hairs.An-
thers0.75-1 mm long.
Distribution- Papua NewGuinea, NorthernProv., MtSuckling.
Habitat- Open subalpine short-grasslands, locally dominant, 3300-3550masl.
Specimensseen:Veldkamp& Stevens 5768,5846(T).
Danthonia irianensisVeldk., spec. nov.
Danthonia vestita auct.nonPilg.
ADanthoniavestitain culmisbasi intra-vaginaliterramosis,paniculis sat densis, plusquam 20 spi-
culis 7.5-11 mm longis(aristis excl.), costa glumaepuberula adpilosa in glumasuperiore,lem-
matepilorum caespitibus densisin serietransversali aliquotspatiosub sinu,setis 1-2 mm longis,
arista8.25-9.25mm longa,paleadorsaliteraliquotpilis provisa, antheris 0.75-1.25mm longis
differt.—Typus:ANU 10952 (Hope) (Lholo;BO,CANB), New Guinea,Irian Jaya,Carstensz,
Carstensz meadow, 13Dec. 1971,3510m.
Culms tufted, branching intra-vaginally atbase, 0.16-0.3 m long. Blades 14-27
cm by 0.5-1 mm, pilose, not articulating withthe sheath. Peduncle glabrous. In-
florescence with morethan20 spikelets, contracted, dense, 6.5-7.5cm long, axes
pilose. Spikelets chasmogamous, 3-or 4-flowered, 7.5-11 mm long (excl. awns).
Glumesas longas thespikelet, midribpuberulous topilosein theupperglume.Lem-
ma with a transverse row of2.75-3.25 mm long, densely tuftedhairs some way
below thesinus, andoneabovethecallus; body 2.5-2.75mmlong; laterallobes2-
3.5 mmlong, 0.73-1.4times as long as body; setae 1-2mm long, 0.3-1 timesas
long as flatportion ofthelobes; awn 8.25-9.5mmlong, exceeding setae by 4-5
mm, geniculate, with0.5-1 basal twists. Paleadorsally with some hairs. Anthers
0.75-1.25 mmlong.
J.F.Veldkamp: Miscellaneous notes on Southeast Asian Gramineae 219
Distribution- NewGuinea, IrianJaya,Carstensz.
Habitat- Dryerareas on till,often underrockoverhangs, 3510maltitude.
Note- ANU 10976(Hope) (BO,CANB,L) from theLowerMerenvalley ofthe
Carstensz is asterilecollection, and only tentativelyincludedhere, mainly because
Danthoniairianensisis theonly tallDanthoniaknown atpresentfromtheCarstensz.
Ithas densely tufted, dullculms, quite unlikethe shiny ones ofD. irianensis, and
very distichousleaves.Ithasthehairy auricles sotypical ofDanthonia.
Specimens seen:ANU 10952(Hope)(T), 10976(Hope);Wissel21.
Danthoniajavanica Ohwi ex Veldk., spec. nov.
Danthoniapilosa auct.nonR. Br.
Pedunculi glabri.Spiculae 10.5-14 mmlongae(aristaeexcl.)aristis solum protrudentibus. Glumae
spiculas plus minusveaequantes.Lemmata primasolum caespitibus(sub)marginalibusdistinctis,
corpore 2.5-3.25mm longo,aristis 8.5-11.5 mm longis,columno brunneo spiris basalibus 3-4.
Margopaleae(intus) sine pilorum caespitibus.—Typus: Afriastini 264 (L holo;BO,K,P), Java,,
MtLawu,21 May 1981,c.3100 m.
Culms tufted, branching mainly extra-vaginally atbase, 0.13-0.6m long. Blades
5-19cm by 0.4-1 mm, glabrous, ultimately articulating withthe sheath. Peduncle
glabrous. Inflorescencewith3-11 spikelets, contracted, 2.5-9cm long, axes pilose.
Spikelets usually cleistogamous, 3-6(-8)-flowered, 10.5-14mmlong (excl. awns).
Glumesas long as the spikelet (onlythe awns protruding), glabrous. Lemmawith2
tuftsof 1-1.5mmlong hairs alongthemargin only; body2.5-3.25mmlong; lateral
lobes2-3 mmlong,0.62-1.2timesas long asbody; setae 2.25-3.5mmlong,0.9-
1.6timesas long as flatportion oflaterallobes; awn 8.5-11.5 mmlong, exceeding
setaeby 3.75-5.5mm(when wet),with 3-4basaltwists. Paleamargin(inside) with-
outtuftsofhairs. Anthers0.6-1.15mmlong.
Distribution Java: Priangan (Papandayan), Madiun(Lawu), Malang (Arjuno),
-
Besuki(Argopuro).
Habitat- Montaneandsubalpine grassfields,open,sunny, sandy torocky places,
inCasuarinaforest, locally verycommon, 2350-3100maltitude.
Note- Thisspecies was identifiedbutnot published by OhwiasDanthoniajava-
nicaOhwi.
Specimens seen:Afriastini 264 (T); Backer s.n. (7/6/1929,8/6/1929), 13317,Buwalda
8148;Koorders 43544;Posthumus 334;vanSlooten 2544; vanSteenis 4104, 4308,4820,7067,
7068, 10924,11679,11864.