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MISCELLANEOUSCHROMOSOMENUMBERREPORTS FORPOA(POACEAE)INNORTHAMERICA RobertJ.Soreng DepartmentofBotany NationalMuseumofNaturalHistory SmitlisonianInstitution Washington,DC20013-7012,U.S.A. [email protected] ABSTRACT Thefollowing66chromosomenumbersandvoucherslor24speciesofPoaL,arereportedfrom Canada,Mexico,andtheU.S.A.:P.ahhrayiaiasubsp.pattfrsonii,2n=42,2n=42,2n=42;V.aipina,2n =28+11,2n=32+1(2x),2n=40+1,2n=42,2n=56;P.arctnasubsp.aperta,2n=98+1;subsp.arctica.In =56;subsp.arctica(longipilaform),2n=56-59,2n=80,In=88;P.atropurpurea,In=28;P.Ingelovii, In=28+1;Pcusickiisubsp.pallida,In=56+11;P.fcndkrianasubsp.albescens,In=28+11,In=56; subsp.fendleriana.In=56(4x),2n=59,In=58-60,21-1=58-64;subsp.longihgula,In=56,2/ica.56; P.glaucasubsp.giflucti,In=56,In=56-58;subsp.rupicola,In=48,2n=48-50,2n=54-56,2nca.100; P.interior.In=42;Plaxasubsp.banjjiana.In=84;P.leptocomaIn=42(2x);Plettermanii.In=14; Pnapensis,In=42;P.nervosa,2n=28;P.iKcidentalis,In=14(2x),2n=28;P.paucispicula,In=42;P. pipe?'!.In=28;Ppratensissubsp.alpi^^cna.In=56+111;Prejlexa,In=28(4x);Psecundasubsp. junci/olia,2n=63;subsp.secunda,2n=84-88+11;Psierrae,2nca.58;Ps(rictiramec],2n=28-29,2n= 29+11;Psupinac.v.SUPERNOVA,In=14;PIracyi,In=28(5x),In=28+1;Pumlaierahssubsp. pachypholis,In=42;andP.unilattralissubsp.unilateralis.In=84. RESUMEN Secitan66numeroscromosomaticosytcstigosde24especiesdePoaL.deCanada,Mexico,yE.E.U.U.: Pabbreviatasubsp.patlersonii.In=42;/'alpma.2n=28+11,2ii=32+I(2x),2n=40+1,In=42.2ti= 56;Ptjreticasubsp.aperta.In=98+1;subsp.arctica.In=56;subsp.oreticfl(formalongipila),2n=56- 59,2n=80,2n=88;Patropurpurea,In=28;P.bigclovii,2n=28+1;Reusicljiisubsp.pallida.In=56+11; P.Jendlerianasubsp.albescens,2n=28+11,2n=56;subsp./endleriaiia,2n=56(4x),2n=59,In=58- 60,2n=58-64;subsp.longiligula,In=56,Inca.56;Pglaiicasubsp,glauca,2)i=56,2n=56-58;subsp. rupicola.In=48,2n=48-50,2n=54-56,2nca,100;Pinterior,2n=42;PJaxasubsp.banjfiana,ln= 84;RleptocomaIn=42(2x);Pletter/nanii.2n=14;Pnapensis,2n=42;Pnervosa.In=28;P occidentalis,In=14ilx);P.paucispicula.In=42;Ppiperi,2n=28;Ppratensissubsp,alpigcna,In= 56+111;Prejlexa.In=28(4x);Psecundasubsp.junci/oiia,2n=63;subsp.secunda.In=84-88+11;P. sierrae.Inca.58;Pstnctiramea.In=28-29,2n=29+11;Psupinac.v.SUPERNOVA,2n=14;Ptracyi, In=28(5x),2n=28+1;Punilateralissubsp.paehyp/ioli.s,2n=42;yPunilateralissubsp.unilateralis, 2n=84, TheFloraofNorthAmericaNorthofMexico(Morinetal.1993)editorialpohcy requiresthatchromosomenumbersbeindependentlypublishedpriortobeing reportedinthetreatments.Therefore,Iamreportingaseriesofchromosome countshereforPoathatareunreportedoronlymentionedwithpartialvoucher SIDA21(4):2195-2203.2005 Table1.VoucherinformationforchromosomecountsinthegenusPoathatarenewormentionedwithnooronlypartialvoucherinformationinSoreng (1985,1990,1991a,1991b,1993,1998)andSorengandHatch(1983).RJS=RJ.Soreng,RWS=RW.Spellenberg. Taxon County Specificlocation,date,collectionno. Chromosomeno.(2n), &State &herbarium andnotes Poaabbreviatasubsp. U.S.A. ClearCr.Co.:RockyMts.,Mt.Evenstop,S5Eof 42(Soreng1991b,withpartial pafferson/7(Vasey) Colorado: Georgetownca.13km,3Aug1984,RJS,R.Bayer, voucher)information A.Love,D.Love& M.Dunford&G.L.Stebbins2555(US) B.M.Kapoor Colorado: Summit/ParkCo.boundaryRockyMts.,TenmileRange, 42(Soreng1991b,with NorthStarMt.,HoosierRidgeWofHoosierPass, partialvoucherinformation) 2Aug1984,RJS,RBayer,M.Dunford&G.LStebbins2548(US) Montana: DeerLodgeCo.:Anaconda-PintlarWilderness,Mt.Tiny, n=21,frompollendivision aboveStormLk.,6Aug]980,RJS&RWS1165-2{US) (Soreng1991b,withpartial voucherinformation) PoaalplnaL. CANADA. BanffN.P,ca,100kmNofBanffonhwy93,Eslopesof 42(new) Alberta: Mt.Peyto,SofPeytoLk.,NofBowLk.,28Jul1980, RJS&RWSWT8{\JS) Alberta: PlateauMt.,betweenMt.LivingstonandMt.Burke,ca.67 56(new) kmdueNofColman,2Aug]980.RJS&RWST105{US) U.S.A. SagaucheCo.:SanLuisMts.,NslopeofBaldyChato,offBig 28+11(new) Colorado: MeadowRd.PR790,17Aug1980,RJS&RWS1406-a(US) Colorado: SagaucheCo.:SanLuisMrs.,NslopeofBaldyChato,offBig 32+1(new) MeadowRd.PR790,17Aug1980,RJS&RWS1406-b(US) Wyoming: ParkCo.:BeartoothPass,Esummit,8Aug]980,RJS& 40+1(new) RWS1213-5{US) Wyoming: SublettCo.:LittleSheepMt.,NWofNendofGreenLakes 32+1(new) ca.6km,10Aug]980.RJS&RWS1290{US) Poaarcttcasubsp.aperta U.S.A. SagaucheCo.:SanLuisMts.,NslopeofBaldyChato,offBig 99(Soreng1985,without (Scribn.&Merr.)Soreng Colorado: MeadowRd.,FR790,17Aug1980,RJS&RWS1412-a(US) voucher) Table1.(continued) Taxon County Specificlocation,date,collectionno. Chromosomeno.(2/?), &State &herbarium andnotes O o O PoaarcticaR.Br. CANADA. KananaskisProv.R,atKananaskisSummit(HighwoodPass), 56(new) subsp.orcf/co Alberta: 4n0e,aronMtW.Asrideet,h3u1saJucla.]69780k,mRJSS&ofRSWeSeb1e09a4nd{USH)wy1,onHwy U.S.A. PitkinCo.(WofLakeCo.line?),RockyMts.,SawatchRange, 88(new) Colorado: IndependencePass,15Aug]980,RJS&RWS1391(US) Montana: DeerLodgeCo.:Anaconda-PintlarWilderness,Mt.Tiny, 80(new) aboveStormLk.,6Aug]98Q,RJS&RWS1180(US) Montana: GlacierCo.:GlacierN.P.,PiganPass,4Aug]98Q,RJS& 56-59(new) RWSl142(\JS) PoaatropurpureaSaibn. U.S.A. SanBernardinoCo.:BaldwinLake,1985,RJS2632(US) 28(Soreng1993,without California: voucher) PoabigeloviiyaseySi U.S.A.New LincolnCo.:WhiteMts„NEofSierraBlanca,below 28+1(Soreng1985,without Scribn. Mexico: MonjeauL.O.,16Jun1981,ff75/5S4f(US) voucher) Poacusickiisubsp.pallida U.S.A. ParkCo.:NEofGardner10km,EofJardine,Jun081984, 56+11(Soreng1991a,with Soreng Montana: RJ52453-a{US) voucher,butlocation incompleteandnumber erroneouslyreportedasRJS 2456) Poafendlerianasubsp. MEXICO. SierraMadreOccidental,Creel,nearairstrip,15Apr1984, 56(new) albescens(Hitchc.)Soreng Chihuahua RJS&RWS2309(DS) Sonera: 5kmNWofCananeaonmicroondasroadNfromroadto 28+11,withinversionbridge Sonora,19Mar1982,ftJS&RlV51780-5{US) &fragments(Soreng1985, withoutvoucher) Poafendleriana(Steud.) U.S.A.New CatronCo.:ShendanGulch,21May]983,RJS&D. 56,withmultivalents(new) Vaseysubsp.fendleriana Mexico: Ward2125{US) Mexico: DonaAnaCo.:OrganMts.,Wside,belowandEofBaylor 56(new) Pk„1 Feb1984,RJS&R.Neilson2/90-b(US) Table1.(continued) Taxon County Specificlocation,date,collectionno. Chromosomeno.(2/?), &State &herbarium andnotes Mexico: GrantCo.:BlackRange,19l<mNWofMimbres,D.Ward 58-64(new) 81-04(NMQ Mexico: LincolnCo.:WliiteMts.,MontgomeryBiologicalResearch 56(new) Mexico: SStaantdioonv,al8Ckom.:NSaonfdiRauiMdtoss.,o,W18baAspe,rJ1u9a8n1T,RaJbSo1P5i8c0ni(cUSA)rea, 58-60(new) NEofAlbuquerque,6Jun]983,RJS&RWS2172{US) Mexico: SocorroCo.:SanMateoMts.,21Mar]984,RJS2303 56(new) [novoucher] MEXICO. SierraMadreOccidental,7kmEofTomachic,14Apr1984, n=28+1,mitotic,pistillate Chiliual-iua RJS2306(US) plant(new) Poafendlenanasubsp. U.S.A. ApacheCo.:ChuskaMts.,6.7kmNEofLukachukai,on 56(new) longiligula{Scr\bn.& Arizona: BlA-13,9Jun]983,RJS&RWS2177{\JS) T.A.Williams)Soreng Wyoming: ParkCo.:MammothHotSprings,08Jun1984,/?JS2454(US) nca.28,mitotic(new) PoaglaucaVahlsubsp. CANADA. PlateauMt.,betweenMt.LivingstonandMt.Burke,67km 56-58,meiosisirregular glauca Alberta: dueNofColman,1Aug1980.RJS&RWS1098-3(US) (new) U.S.A.New TaosCo.:WheelerPeak,ndge3.3kmNofpeak,0.4kmSof 56,multivalentsandlaggers Mexico: FrazerMt.,19Aug]980,RJS&RWS1454-1(US) common(new) Poaglaucasubip.rupicola U.S.A. PitkinCo.:RockyMts.,SawatchRange,IndependencePass, Ca.TOO(new) (Nasli)W.A.Weber Colorado: 15Aug^980.RJS&RWS1372-18{US) Wyoming: ParkCo.:ClayButteLook-Out.,ca.2kmWofBeartooth 54-56,multivalents(new) Lk.,8Aug]980,RJS&RWS1221-2{US) WWyyoommiinngg:: dSfutbol,eRtJtSC&o.R:WToSp1o2f21Li-t5tl{eUSS)heepMt.,NWofNendofGreen 4488-(5n0e,w)multivalents(new) o3D Lakesca.6.25km.1 Aug1980,RJS&RWS1299-6(US) PoainteriorRydb. U.S.A. SagaucheCo.:SanLuisMts.,NslopeofBaldyChato,offBig 42(new) Colorado: MeadowRd.FR790,17Aug1980,RJS&RWSM22-a-3(US) 2 Table1.(continued) Taxon County Specificlocation,date,collectionno. Chromosomeno.(2n), &State &herbarium andnotes o o Poa/oxoHaenkesubsp. U.S.A. GlacierCo.:GlacierN.R,PiganPass,4Aug1980,RJS& 84(Soreng1991b,with banffianaSoreng Montana: RWS1137{[]S) partialvoucherinformation) Poa/epfocomaTrin. U.S.A. GlacierCo.:GlacierN.P.,PiganPass,4Aug1980,RJS& 42(Soreng&Hatch1983) MUtoanht:ana: RSuWmSmi1t148C-o.4:(MUtS.)MurdockEofBaldMt.Pass,Hwy150,1 42(new) Aug]980,RJS&RWS1347-2{US) PoalettermaniiVasey CANADA 14(A.Love,pers.com.,letter orU.S.A.: ca.1982,reportedbySoreng 1991a,voucherunknown) PoanapensisBeetle U.S.A. NapaCo.:Calistoga,SendoflandingstripWofLincoln 42(Soreng1991a,with California: Ave.,27May1986,RJS2926(US) partialvoucherandlocation) Poanervosa(Hook.)Vasey U.S.A. MarionCo.:SilverCr.FallsS.P.,WinterFalls,6Jun1986, 28(new) s.str,(excludingPoa Oregon: WS2960(US) wheeleriMAsey) PoaoccidentalisVasey U.S.A.New RioAribaCo.:SWofCoyote,PuercoC.G.,ca.33kmNW 14(Soreng&Hatch1983) Mexico: ofLosAlamos,15Aug1978,RJS&SI.Hatch48(US) NewMexico: OteroCo.:SacramentoMts.,ca.8.3kmENEofCloudcroft 28(Soreng&Hatch1983) SofNM-244onCR-7,Decearly1978,RJS123b(US) NewMexico: OteroCo.:Cloudcroft,SI.Hatch-2222(TAES) 14(Soreng&Hatch1983, countbyS.LHatch) PoapaucispicutaSaibn. CANADA. BanffN.R,ca.100kmNofBanffonhwy93,Eslopes 42(Soreng&Hatch1983, &.Merr Alberta: ofMt.Peyto,SofPeytoLk„NofBowLk.,28Jul1980, reportedasPoaleptocoma) RJS&RWS1016{US) PoapiperlHitchc. U.S.A. DelNorteCo.:Offhwy1990.6kmonPatrickCr.Rdabove 28(Soreng1990,1993, California: theMiddleForkoftheSmithRv.,2Jun1986,RJS2950 withoutvoucher) PoapratensfsL.subsp. U.S.A. (US)Nome,Jul]983,G.L.StebbinsA-3107!{\JS) 56+111,originaldet.asPoa alpigena(Lindm.) Alaska: arctica(new) Hiitonen , Table1.(continued) Taxon C&oSutnattye &Spehceirfbicarlioucamtion,date,collectionno. aCnhdronmootessomeno.(2f?), PoareflexoVasey&Scribn. U.S.A.New TaosCo.:WheelerPk.LaCaiBasin,ca.1.7kmNNW 28(Soreng&Hatch1983) Mexico: ofpeak,19Aug]980,RJS&RWS1478-4(US) Utah: SummitCo.:Mt.MurdockEofBaldMt.Pass,Hwy150, 28(Soreng&Hatch1983) Wyoming: P1a1rAkuCgo.:]C9l8a0y,RBuJtSte&RLWooSk1O3u3t6.,{\cJaS.)2kmWofBeartooth 28(new) Lk.,8Aug]980,RJS&RWS1227{[JS) Wyoming: SublettCo.:SsideofLittleSheepMt.,NWofGreenLakes 28(Soreng&Hatch1983) Poajunsceicfoulnidaa{Sscuubbsnp..)Soreng NU.eSv.aAd.a: HcLaaw.ny5dek5rm0,,C1o1.J:uTlAou]ig9y8a01b,9e8R0RJ,aSnR8gJ2eS1,&(ERUoSWf)SAu1s2t6i0n-3ca.(U1S3)kmon 6pa3rt(iaSlorveonugch1e9r91ibn,fwoirmtahtion) PoasecundaiPresI U.S.A. GlacierCo.:GlacierN.P.SiyehPassTrail,4Aug1980, 84-88+11(Soreng1991b, subsp.secunda Montana: RJS&RWS1135{US) withpartialvoucher information) Poas/erraeT.Howell U.S.A. EldoradoCo.:DeepCanyon,N.ForkofAmericanRiver, ca.58(new) California: EofColfaxoffhwy80ca.2m,ca.0.8kmNEofriver crossingofIowaHill-ColfaxRd.,30May1986,RJS&G.L Stebbins2931(US) PoastnctirameaHitchc. CMhEiXhIuCaOh.ua: SDiiearbrlaoMPaadssr,e2OkccmidWentoanl,roWadoftoSaMnadJeorsae,B1a3biAcporra,19C8.4E,l n19=911a4,+w1i,tmhitvoosuicsh(eSroraenndg RJS&RWS2304-a{[JS) partiallocation) PoaSUsPupE/RnNoOSVcAhrad.cv. CMU.ahSri.yhAl.uaanhdu:a: CCdluallyoto,iuRveJatStte&ed5Rf9Wr5oS0m-2bc30o(U4mS-m)ber{cUiS)alseed,2000,/?J5<SJ. nnun==re71p,4o-mri1tte5od+s)1i1s,(Jm.iCtaosyiosue(tnteew,) Poafracy/Vasey U.S.A.New BernalilloCo.:SandiaCrest,rimNofTram,16Jul1981 28(Soreng&Hatch1983) MNeexwicMoe:xico: RRCoJJlSSf&&aKxS.ICG.oa.H:dazWticNahW/664o4f2(URS(a)UtSo)n,RatonCityPark,16Aug1978, 28(Soreng&Hatch1983) Table1.(continued) Taxon County Specificlocation,date,collectionno. Chromosomeno.(2/j), &State &herbarium andnotes o o NewMexico: ColfaxCo.:NWofRaton,RatonCityPark,31May1979, 28(Soreng&Hatch1983) fiJS266(US) NewMexico; ColfaxCo.:NofRaton,JohnMayer'sRanch,downcanyon 28(Soreng&Hatch1983) fromRatonPassonsideofBartletMesa,EsideofU5-25,31 o May1979,/?JS26/(US) O NewMexico: ColfaxCo.:RatonRanch,NWofRatonca.7.5km,3 28(Soreng&Hatch1983) NewMexico: CJoulnfa1x97C9o,.:RRJSat2o7n2R(aUnS)ch,NWofRatonca.12km,3Jun 28(Soreng&Hatch1983) 1979,ftJS2/4(US) NewMexico: LincolnCo.SierraBlanca.circbelowthePeak,10Jul1982, 28+1(new) RJS&RWS2007iUS) Poaunilateralissubsp. U.S.A. PacificCo.:Ilwaco,RWS&D.Southerland1522A(NMC) 42,R.W.Spellenbergcount pachypholls(Piper) Washington: (Soreng1998,withpartial Soreng voucherinformation) PoaunilateralisScnbn. U.S.A. CurryCo.:3.3kmSofGoldBeachonserpentineroadcut, 84,asPoaunilateralis,]. subsp.unilateralis Oregon: BuenaVistaWaysides,300ftabovetheocean,22Jun1949, Clausenunpublished J.Clausen2151(CAS) (Soreng1991a,withvoucher butnolocation) 2202 BRIT.ORG/SIDA21(4) informationinSoreng(1985,1990,1991a,1991b,1993,1998)andSorengandHatch (1983).Table1includesiullspecimencitationsandherbaria(acronymsfollow- ingHolmgrenetal.1990)wherethevouchersaredepositedforallofmyprevi- ousandnewreports.Myowncountsweredonebetween1978and1988.Meth- odsforthechromosomepreparationsweregiveninSorengandHatch(1983). Inaddition,vouchersand/ornotesofafewcountsdonebyotherbotanists/ authorsthathavenotbeenreportedpreviouslyareincluded.JacquesCayouette providedhisnewchromosomecountofP.supinafromtherecentlyintroduced (inNorthAmerica)cultivarSUPERNOVA.ThecountreportedinSoreng(1991a) forP.lcttcrmai}uwasmentionedtomeinaletterbyA.Love,ca.1982.Thisre- portisinterestingasitraisestothreethenumberofdiploidspeciesintheNew World(theothersarePoccidentalisandP.pseudoahbreviata).Verificationof thecountforP.lettermanii isneededsince1onlyhavethecorrespondence record.1foundanunpublishedreportforPunilateralissubsp.unilateralisof 2n=84onaherbariumspecimenatCAS,thecounthkelydonebythecollector ofthespecimen,geneticistJensClausen.Myers(1947)reportedacount by StebbinsofIn=42,presumablyforthetypicalsubspecies(asPunilalcraUs stuhebscpo.asptacohfyNphWohsOriesgroanthaerndloacdalj,acmeonrteWarsecheinntgltyonp)u.blRiishcehda,rdanWdrSepsetlrlicetnebdertgo madeacountof2n=42forPunilaleralissuhsp.pachyphohs. Althoughemphasishasswitchedawayfromcytogeneticcomparisonsof speciestoDNAanalysesinPoa(Gillespie&Soreng2005;Soreng1990),itis importanttohaveanunderstandingofthecytogenetichistoryoftaxainorder tointerpretresultsofotheranalyses,andtobeabletolocatevouchersandknow wheretheywerecollected.Ofthe66countslistedinTable1,34areunreported elsewhere.ThebasechromosomenumberinthegenusPoaisx=7,andthe countsreportedheregenerallycorrespondtomultiplesofseven,butunbal- ancedsetsol chromosomeswerefrequentlyencountered.Romannumerals givenafternumbers(i.e.;2n=2S+11)representunpairedchromosomesinthe metaphaseoranaphaseofmeiosis,orunbalancednumbersinmitosisorlater stagesofmeiosis.AlthoughsupernumaryorBchromosomeshavebeenreported frequentlyinPoa,noattemptwasmadetodistinguishunbalancedchromotin ot thistypeIroinfragmentsresultingfromirregularmeiosis,etc.Mostofthe countsreportedhereconformtonumbersreportedbyotherauthorsforthesame taxa.Poasierrac{2n=28)istheonlytaxonreportedhereforthefirsttime.In taxawithpreviouslyreportedcounts,otlicrthanmyown,anddisregardingthe extrachromotin,theonlythepreviouslyunrecordednumbersinanytaxonre- portedhereare;In=56inPfcndlcrianasubsp.albescens,andIn=48,48-50, 54-56,andca.100inPghiucasubsp.rupicola.Thisworkcontinuestoshowthe patterninPoaoffewdiploidtaxa,numeroustaxav/ithlow,fairlystabletetra- andhexaploidnumbers,othertaxawithhighereupolyploidseries,andtaxa . . 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