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Maritime Hunter-Gatherers in the Chonos Archipelago (43°50'–46°50' S) PDF

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This article was downloaded by: [omar reyes] On: 06 March 2015, At: 09:56 Publisher: Routledge Informa Ltd Registered in England and Wales Registered Number: 1072954 Registered office: Mortimer House, 37-41 Mortimer Street, London W1T 3JH, UK The Journal of Island and Coastal Archaeology Publication details, including instructions for authors and subscription information: http://www.tandfonline.com/loi/uica20 Maritime Hunter-Gatherers in the Chonos Archipelago (43°50’–46°50’ S), Western Patagonian Channels Omar Reyes Báeza, Mauricio Moragabc, César Méndezb & Alexander Cherkinskyd a Instituto de la Patagonia, Centro de Estudios del Hombre Austral, Universidad de Magallanes, Punta Arenas, Chile b Departamento de Antropología, Universidad de Chile, Santiago, Chile Click for updates c Instituto de Ciencias Biomédicas, Universidad de Chile, Santiago, Chile d Center for Applied Isotope Studies, University of Georgia, Athens, Georgia, USA Published online: 03 Mar 2015. To cite this article: Omar Reyes Báez, Mauricio Moraga, César Méndez & Alexander Cherkinsky (2015): Maritime Hunter-Gatherers in the Chonos Archipelago (43°50’–46°50’ S), Western Patagonian Channels, The Journal of Island and Coastal Archaeology, DOI: 10.1080/15564894.2014.1001920 To link to this article: http://dx.doi.org/10.1080/15564894.2014.1001920 PLEASE SCROLL DOWN FOR ARTICLE Taylor & Francis makes every effort to ensure the accuracy of all the information (the “Content”) contained in the publications on our platform. However, Taylor & Francis, our agents, and our licensors make no representations or warranties whatsoever as to the accuracy, completeness, or suitability for any purpose of the Content. Any opinions and views expressed in this publication are the opinions and views of the authors, and are not the views of or endorsed by Taylor & Francis. The accuracy of the Content should not be relied upon and should be independently verified with primary sources of information. 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Terms & Conditions of access and use can be found at http://www.tandfonline.com/page/terms- and-conditions 5 1 0 2 h c r a M 6 0 6 5 9: 0 at ] s e y e r r a m o [ y b d e d a o nl w o D JournalofIsland&CoastalArchaeology,00:1–25,2015 Copyright©2015Taylor&FrancisGroup,LLC ISSN:1556-4894print /1556-1828online DOI:10.1080/15564894.2014.1001920 Maritime Hunter-Gatherers in the ◦ ◦ Chonos Archipelago (43 50’–46 50’ S), Western Patagonian Channels Omar Reyes Ba´ez,1 Mauricio Moraga,2,3 Ce´sar Me´ndez,2 and Alexander Cherkinsky4 1InstitutodelaPatagonia,CentrodeEstudiosdelHombreAustral,Universidad deMagallanes,PuntaArenas,Chile 2DepartamentodeAntropolog´ıa,UniversidaddeChile,Santiago,Chile 3InstitutodeCienciasBiom´edicas,UniversidaddeChile,Santiago,Chile 5 4CenterforAppliedIsotopeStudies,UniversityofGeorgia,Athens,Georgia,USA 1 0 2 h c r a M ABSTRACT 6 0 6 Thisarticlepresentsthecurrentadvancesinarcheologicalresearchon 9:5 theWesternPatagonianChannels,specificallytheChonosArchipelago 0 (43◦50’–46◦50’S).Basedonalargespatialscale,weaimtocontribute ] at tothediscussionofthedispersionandcharacteristicsoftheoccupation es ofthePacificcoastofsouthernmostSouthAmerica.Resultsshowthat y e allofthecontextsrecordeddonotexceed3600calBP.Siteformation r ar processesthatmayhaveactedinthepreservationofthearcheological m record of this area led us to question if this region was among the o y [ last to be settled (as suggested by current chronological data), or if d b thedynamictectonicactivitythatpermanentlytransformsthiscoastis e playingamajorroleinconcealingearlierevidence,therebyintroducing d oa asignificantresearchbias. nl w o Keywords archipelagic settlements, Late Holocene, maritime hunter-gatherers, site D formationprocesses,WesternPatagonianChannels INTRODUCTION Pleistocene and throughout the Holocene (Aniya1999;Fairbanks1989;Isla1989;Long The permanent degradation and transfor- 2001;NakadaandLambeck1988)hasbeen mation of coastlines caused by the fluctu- problematic for the archaeological identi- ations of sea levels since the end of the fication and interpretation of sites on the Received13January2014;accepted15July2014. AddresscorrespondencetoOmarReyesB´aez,InstitutodelaPatagonia,CentrodeEstudiosdelHom- breAustral,UniversidaddeMagallanes,Av.Bulnes01890,Casilla113-D,PuntaArenas,Chile.E-mail: [email protected] Color versions of one or more of the figures in the article can be found online at http://www.tandfonline.com/uica. 1 Omar Reyes Ba´ez et al. American coasts (e.g., Bailey 2004; Erland- of the evidence and the temporality of the son2001;GusickandFaught2011;Richard- settlementonthecontinent’sPacificmargin son1981).Thesetransformationsinfluenced (e.g.,Dillehay2009;Erlandson1993). the dependency relationship of these com- Taking into account sea-level changes, munities with their maritime or coastal theactivelocalseismicdynamics,andtheef- environment (Bailey and Parkington 1988; fectsoftheseoncoastalarcheologicalsites, Erlandson and Fitzpatrick 2006). Under- we have surveyed a previously understud- standing changes in accessibility of these iedzoneofthenorthernarchipelagosofthe spaces and the local redistribution of ex- Western Patagonian Channels. In this arti- ploitable resources on timescales of hun- cle, we present the first archaeological as- dreds or thousands of years requires pale- sessment of the human settlement in the ogeographical and paleoenvironmental re- ChonosArchipelago(43◦50’—46◦50’S;see constructions in order to guide the search Figure1),whichincludes:1)thedescription for archaeological evidence. The degree of oftheresearchmethod;2)theidentification preservationofthesespacesisalsostrongly andsystematicrecordingofthearchaeologi- associated with the changes in the land- calsitesassociatedwiththesecoastlines;3) 5 scape, which were frequently catastrophic the description and assessment of the geo- 01 (Lomnitz 1970). Only with this informa- morphologicalchangesthataffectthelatter; 2 h tion can the regional cultural development and4)thestudyofthearchaeologicalcollec- rc ofcoastalsettlementsbeinterpreted(Dille- tions.Thisassessmentenablesustoadvance a M hay et al. 2008; Fedje and Christensen an initial panorama of the settlement, both 6 1999; Mackie et al. 2011; Thompson and spatiallyandchronologically,inthecontext 0 6 Worth2011). ofmarineadaptationonthesoutherntipof 5 9: On the margin of the American Pacific theAmericancontinent. at 0 therearevastareas,someofwhichhavenot ] yet been investigated, with coastlines sub- s e jecttopermanentchangesanderosionalpro- y re cesses (Punke and Davis 2006; Rick et al. CULTURALCONTEXT ar 2006). The materials that define the occu- m o pation of these coastal spaces, the techno- On the southern tip of the Chilean Pacific [ y logicalassemblages,andtheevidenceofthe coast, there is a long and complex net of b d consumed resources, are frequently recov- channels, fjords, and thousands of islands de eredfromintertidalzonesordegradedcoasts knownasthe“AustralArchipelagos”orthe a o (ErlandsonandMoss1999).Therefore,itis “WesternPatagonianChannels”(Bird1988; wnl necessarytodevelopstrategiesfortheloca- Emperaire1963;Steffen1910).Thisareaex- o tion,recording,dating,andinterpretationof tends more than 1,600 km north-to-south D the material evidence of these settlements and reaches from the Reloncav´ı Sound to tounderstandtheculturaldynamicsthatde- CapeHorn(41◦30’–55◦60’S).Thecoastline finecoastaladaptation(e.g.,Bailey2004;Er- totals more 19,000 km with a surface area landson2001;Yesner1980).Suchstrategies of 240,000 km2, which should be added to willenableustomeasuretheunequaldistri- ˜240 km of coastline north of the Beagle bution of the archaeological records in the Channel and south of Tierra del Fuego, in region.Theimplicationsaretwofold.Atthe Argentina.Inthisvastarchipelagicarea,the regional level and for Patagonia, it is possi- distinguishable maritime adaptations com- bletodocumentadiversityofspacesoccu- menceapproximatelyat6400BP(7300cal pied by maritime hunter-gatherers in areas BP)towardthesouthernmosttipoftheconti- notpreviouslyexamined.Thisfactisimpor- nentaccordingtodataobtainedintheBeagle tant because the archipelagic geography of Channel (Orquera et al. 2011; Orquera and thestudyareaissingularforitsextension,re- Piana2009). strictionofaccess,andweatherconditions. In this regional context, several top- Additionally,atamegascale,ourresultscon- ics have been discussed. Among them, the tribute to the discussion of the variability geographical origin of the maritime adap- 2 VOLUME00•ISSUE00• 2015 Western Patagonian Channels 5 1 0 2 h c r a M 6 0 6 5 9: 0 at ] s e y e r r a m Figure1. (a) Western Patagonianchannels, (b) Chonos archipelago and sites mentionedin this [o study:1.Puqueldo´n,2.Repollal,3.Puquit´ın01,4.SenoGala1,5.IslaHarris,6.IslaSin by Nombre,7.Canalad1,8.IslaBenajam´ın01,9.Ipu´n1,10.IslaElena1,11.IslaVictoria d 2,12.CanalCuche1,13.IslaCaniglia,14.CanalDarwin(#1,2),15.Nahuelqu´ın(#1,2, e d 3),16.PosaLasConchillas,17.IslaAcuao1,and18.IslaGon˜i1. a o nl w o tation stands out, whether it occurred in accessbecauseoftheirproximitytopopula- D the southern area (the Strait of Magel- tion centers. Our research, which focused lan/Beagle Channel) or the northern area on undocumented sectors of the Western (Chiloe´/Reloncav´ı Sound) (Legoupil 2003; Patagonian Channels, aims to include vast LegoupilandFontugne1997;OrqueraandPi- coastalandinsularunexaminedspacesinthe ana1988;Prietoetal.2013;Rivasetal.1999). regional archaeological discussion (Erland- Whatwasthedispersionspeedofthisadapta- son 1993). The earliest archaeological site tionphenomenon(Bird1938;Orqueraetal. recorded in the Chonos Archipelago corre- 2011)?Wastherecontinuityordiscontinuity spondstoGUA10,yieldinga5020 ±90BP ofculturaltraditionsinthisvastarchipelagic 14Cage(5730calBP;Porter1993),whichcur- territory(Legoupil1997,2003;Morelloetal. rentlystandsasanisolatedlocalitywithouta 2002;OcampoandRivas2004;Orqueraetal. regionalcontext.Additionally,ourresearch 2011; Piana and Orquera 2007)? The study representsanimportant steptowards char- of these topics has been based on archaeo- acterizingthevariabilityofmaritimehunter- logical research conducted in the two geo- gatherersettlementsandcalibratingthefor- graphical extremes of this area, which also mation processes that inhibit or enable the correspond to the sectors with the easiest JOURNALOFISLAND&COASTALARCHAEOLOGY 3 Omar Reyes Ba´ez et al. identificationofthearchaeologicalrecordin definedastemperateDonatiaandOreobo- achangingcoastlinelandscape. lus coastal peatland (Luebert and Pliscoff 2006).Thevegetationiscontrolledlatitudi- nallyandaltitudinallybystrongtemperature gradients and precipitation (Abarzu´a et al. STUDYAREA 2004;Sepu´lvedaetal.2009). The rich fauna in the study area is rep- TheChonosArchipelagoconsistsofagroup resented by a diversity of sea mammals: of more than 150 islands that form a dense 18 species of cetaceans, two pinnipeds, netofchannelsandfjordswithnarrowand twoseals,andtwomustelids(Aguayoetal. abruptcoastlines.Thearchipelagoexpands 2006;Zamoranoetal.2010),inadditionto betweenthesouthoftheChiloe´Archipelago inthe GulfofCorcovado (43◦50’S)andthe 22 species of mollusks (bivalves and gas- TaitaoPeninsula(46◦50’S).Itcoversanarea tropods),crustaceansandechinoderms(Os- of ∼54,000 km2 that is nearly 360 km long orio and Reid 2004), 29 species of fish (NavarroandPequen˜o1979),and46species north-to-southand150kmeast-to-west.This ofbirds(Vuilleumier1985).Incontrast,the territoryislocatedtothewestoftheAndes 5 MountainsandwasintenselyshapedbyQua- terrestrial fauna on the islands are scarce 01 ternary glacier action (Bennett et al. 2000; (smallrodents).Onthecontinentalcoastline, 2 h Glasseretal.2004;Heusser2002;McCulloch the pudu´ (Pudu pudu), the huemul (Hip- arc etal.2000).Theabrupttopographyincludes pocamelus bisulcus) and, occasionally, the M puma (Felis concolor) are found (Gonz´alez slopes of hundreds of meters, with depths 6 etal.2009). 0 of200and700mbslinthefjordsandchan- 6 nels.Thisregionalsopresentsanactivesub- 5 9: duction zone dominated by tectonic plates 0 at knownastheChilean Triple Junction(the MATERIALSANDMETHODS ] Nazca,SouthAmericanandAntarcticplates) s e y and large faults that converge in the Taitao Acoastalsurveywasconductedtothewest r re Peninsula(46◦–47◦S;D´ıaz-NaveasandFrutos of Traiguen Island, the Darwin and Cuche ma 2010; Ramos 2005). The faults are respon- Channels, and the Smith Islands (Quetros, o sible for major earthquakes and tsunamis, Renaico,Chaculay,andElena),thelatterlo- [ y whichcontinuallymodifythecoast(Lomnitz catedattheAise´nfjord.Itwasperformedby b d 1970;PlafkerandSavage1970).IntheAndes, seabecausenavigationistheonlymeansof e d there is a chain of large volcanoes that re- transport that can access the channels and a nlo flectthistectonicactivity(NaranjoandStern coastsinthisterritory(Figure2).Thissurvey w 2004) and which are responsible for a sub- included 250 km of coastline in an area of o D stantialportionofthesedimentofthenearby 60kmwest-to-eastand50kmnorth-to-south. valleys. Coastalsitesareprimarilycomposedoflithic The weather in the area is typically remains, bones, and mollusk shells or their oceanicwithastronginfluenceofthewest- periostracae, which represent the archaeo- ern winds, which cause significant precipi- logical evidence of camp sites of maritime tation (>3000 mm per year) to the west of hunter-gatherers. Many of these sites have theAndes(Garreaud2009).Theannualaver- beensubmergedand“washedaway”bythe agetemperaturesareapproximately∼10◦C tides,thustheirremainsaremainlyrecorded (DGA 2003). The dense vegetation dom- intheintertidalarea.Inthesecases,system- inated by Nothofagus, Weinmannia, and aticcollectionwasperformedduringthelow coniferscharacterizestheinteriorareaofthe tide.Primarily,lithicmaterialshowedsignifi- fjords in the form of temperate rainforest, cantsignsoferosiononthesurfaces(Schiffer whichistypicalofcoldandhumidregimes 1996). (Villagr´an1988).Between43◦30’and45◦S, Inordertodetectsubsurfaceculturalde- the vegetation is defined as temperate Pil- positsinlow-orzero-visibilityconditionsas gerodendronandAsteliaconiferouscoastal aresultofdensevegetation,boreholeexca- forest, whereas south of 45◦ to 51◦ S, it is vationswereperformedusinganAMS,Inc., 4 VOLUME00•ISSUE00• 2015 Western Patagonian Channels 5 1 0 2 h c r a M 6 0 6 5 9: 0 at ] s e y e r r a m Figure2. ImageofthemaritimesurveysalongChonosArchipelago,Galasound. o [ y b d kitwith60mmdiameterFrenchandDutch dens.Withoutexception,theseremainshave e ad augers.Sedimentcoresfromdepositsmore been looted, fragmented, or removed, and o nl than 5 m deep were extracted. The bore- their integrity is low. In exceptional cases, w hole excavations detected highly obtrusive it was possible to conduct basic bioanthro- o D deposits(Lightfoot1986),suchasshellheaps pological analyses (to determine sex, age, andpaleosols,andprovedtobeanefficient andminimum numberofindividuals (MNI) tooltocalculatethevolumeoftheanthropic (Buiskstra and Ubelaker 1994). Therefore, deposits (stratigraphic expression) and to our efforts focused on determining radio- recover deep organic samples for dating. carbonchronologydirectlyfromthehuman Thus,itispossibletoestablishsedimentation remains and performing isotopic analyses ratesandtoassesstheaccumulationofwaste (δ13Ccollagenandδ15Ncollagen)tocharac- deposits. terizethediet.All14Candstableisotopesam- Thehumanboneremainsanalyzedcor- ples were carefully extracted to avoid con- respondtoasample(n=10)oftheChonos tamination.Theseanalyseswereperformed Collection of the Universidad de Chile, by the Center for Applied Isotope Studies, whichwasrecoveredinthe1980sand1990s University of Georgia, using the method of (OcampoandAspillaga1984;S´aez2008),but Cherkinskyetal.(2010).Crushedbonesam- also includes new findings (n = 10; Reyes ples were diluted in 1N acetic acid to re- et al. 2013). All of the remains were re- movecarbonates(surfaceabsorbedandsec- covered in caves/rockshelters or shell mid- ondary). Periodic evacuation insured that JOURNALOFISLAND&COASTALARCHAEOLOGY 5 Omar Reyes Ba´ez et al. evolved carbon dioxide was removed from the signs of abrasion on the surfaces of the the interior of the sample, and that fresh artifacts because these occurred frequently acid was allowed to reach even the inte- asaresultoftheirintertidalexposure. riormicro-surfaces.Chemicallycleanedsam- pleswerereactedundervacuumwith100% H3PO4inordertodissolvethebonemineral RESULTS and release carbon dioxide from bioapatite for carbon isotope ratio analysis. Residues The archeological survey on the Chonos werefilteredandrinsedwithdeionizedwa- Archipelagoallowedrecording10shellmid- terandunderslightlyacidcondition(pH= dens, two emerged intertidal management 3),heatedat80◦Cfor6hourstodissolvecol- features (locally known as fishing pens), lagenandleavehumicsubstancesinthepre- andthreerockshelterswithscatteredhuman cipitate.Thecollagensolutionwasfilteredto boneassemblagesintheirinteriors(Table1). isolatepurecollagesanddriedout.Thedried Basedontheseresults,andthoseofother collagen was combusted at 575◦C in evac- researchers(Porter1993;Reyesetal.2007; uated/sealed Pyrex ampoule in the present SternandCurry1995;SternandPorter1991), 5 CuO.Thecarbondioxideandnitrogenwere it could be established that the location of 01 cryogenicallyseparated.Theisotopicratios thesesiteswaslimitedtotheintertidalzone h 2 of carbon (13C/12C) and nitrogen (15N/14N) or a few meters from it (basal deposits be- arc were measured separately, with a Finnigan tween0and3masl)becausetheruggedge- M MAT 252 Isotope Ratio Mass Spectrometer ography and dense vegetation constrained 6 followinganoff-linesamplepreparation.The 0 settlement possibilities. Such sites are fre- 6 dual-inlet rationing system alternately mea- quentlyobservedinassociationwithcreeks. 5 9: suresunknownsampleandknownreference There is a high probability that numer- at 0 gas to achieve a high-precision stable iso- ous sites remain undiscovered along our ] tope measurement. Calibration is made us- route.Thefirstfactorinfluencingsitedetec- s e ing NIST standard hydrocarbon oil, NBS-22 y tion is related to the rugged topography of mar re (inNpoearktehsoeutsaal.n2d0(0(cid:2)6)).uTshinegratthioesδanroeteaxtiporne,sasnedd tcheesscibliiflfistya.loTnogththisefcaoctaostrs,wwehmicuhstlimadidtstahce- o aremeasuredwithrespecttothelimestone weatherconditions,whichfrequentlymake [ y Vienna Pee Dee Belemnite (V-PDB), in the navigationdifficult.Athirdfactoristhedense d b caseofδ13Ccollagen,andtoatmosphericairni- vegetation that covers the coastline, which de trogen(AIR),inthecaseofδ15N(Schwarcz preventsaccessandaffectsvisibility.Tocom- a wnlo a0n.1d(cid:2)Scfohroeδn13iCngaenrd2l0e1ss1)t.haEnrr0o.r2(cid:2)is lfeosrsδt1h5aNn. pweanssaanteefffiocriethnetswe,aythteosbaomrephleolientseecahrcnhiqfuoer Do Allofthedateswerecalibratedat2σ range anthropogenicsedimentsandremains.How- with OxCal 4.2 (Bronk Ramsey 2009) with ever, this technique is limited to in situ de- theShCal13curve(Hoggetal.2013)andare positsandwithintherangeoftheavailable expressedascalibratedyearsbeforepresent extensions(6m).Coringwasunevenlyper- (calBP). formed along the coast, favoring areas suit- Theanalysesoflithicmaterialincludeda ablefordisembarkationandwheretopogra- sampleoftheartifactsrecoveredatdifferent phyandvegetationcoverallowed(Figure2). sites of the intertidal zone, both within the Baysandbeachesprotectedfromthewind, surveyedareaandfromtheCurryCollection locations near fresh water sources, and ar- oftheInstitutodelaPatagonia,Universidad easwithplaintopographywerethemostfre- deMagallanes.Giventheselectivenatureof quentplacesforsites.Inlandincursionsdid thesample,theanalysesconsistedprimarily not surpass 40 to 50 meters from the line ofatechno-typologicalclassification,which, of maximum tide. Once a site was discov- however,includedtheobservationoftech- ered,coringwasusedindefiningitsexten- nical attributes to understand the manufac- sionanddepthofdepositswithanaverage turing method of some of the tools (Inizan of five borehole samples per site. A fourth et al. 1995). Special attention was given to factor is related to the degradation of the 6 VOLUME00•ISSUE00• 2015 Sitetype Shellmiddenand intertidallithic scatterShellmidden Shellmidden Shellmiddenand intertidallithic scatter Shellmidden Shellmiddenand intertidallithic scatterCavewith dispersesurface humanremainsCavewith dispersesurface humanremainsShellmiddenand intertidallithic scatter uedonnextpage) n Preservation Tidalerosion Minoralterations Minoralterations Tidalerosion Minoralterations Minoralterations andtidalerosion Anthropogenic disturbance Anthropogenic disturbance Minoralterations (Conti s 6 March 2015 Discoveryandrecoverytechniques Stratigraphic characterization, andboreholetestBoreholetests Boreholetests Stratigraphic characterization, intertidal collections,and boreholetestsBoreholetests Testpits,intertidal collections,and boreholetestsChonoscollection Chonoscollection Stratigraphic characterization, excavation, intertidal collections,and boreholetests 0 6 5 y eyes] at 09: Chronolog(incal∗yrBP) 3400–1270 1910 530 3600 Modern 1330–1270 1470–770 2350–1760 1760–500 r r a m m o oe wnloaded by [ inthisstudy.Distancefrshorelin(inm) 0 5 5 0 3 0 UK 15 5 o dt D sdiscusseasedeposialtitude(inmasl) –1 2 2 1 2 –1 UK UK 2 xtB e t n o d yofsitec atitudeanlongitude◦4539’S–◦46’W73 ◦22’S–45◦56’W73◦4526’S–◦56’W73◦4527’S–◦56’W73 ◦4526’S–◦06’W74◦4409’S–◦06’W73 ◦35’S–44◦43’W74 ◦4514’S–◦55’W73 ◦4527’S–◦21’W73 r L a m m 1 n n 2 1 Table1.Su Site PosaLas Conchillas CanalCuche CanalDarwi 1CanalDarwi 2 IslaCaniglia SenoGala1 ´IslaIpun1 IslaVictoria ´Nahuelquın 7 Sitetype Emerged intertidal management featureEmerged intertidal management featureShellmiddenand intertidalbone scatterCavewith dispersesurface humanremainsCavewith dispersesurface humanremains andashell middenStratigraphic depositand intertidallithic scatterCavewith dispersesurface humanremains n n servation alterations alterations tidalerosio opogenic urbance opogenic urbance tidalerosio opogenic urbance Pre Minor Minor Major Anthr dist Anthr dist Major Anthr dist 06 March 2015 Discoveryandrecoverytechniques Surfacerecord Surfacerecord Intertidal collections Surfacerecordand collections Chonoscollection Curry’scollection Chonoscollection 6 5 y eyes] at 09: nued)Chronolog(incal∗yrBP) Undated Undated 1530 1830–1660 700–660 Undated 680 r r nti ma Com o (oe wnloaded by [ inthisstudy.Distancefrshorelin(inm) 15 10 0 0 UK 0 UK o dt D sdiscusseasedeposialtitude(inmasl) 2 2 –1 1 UK –1 UK xtB e t n o d yofsitec atitudeanlongitude◦4527’S–◦42’W73 ◦28’S–45◦41’W73 ◦39’S–45◦48’W73 ◦20’S–45◦24’W73 ◦4443’S–◦13’W74 ◦56’S–45◦58’W73 ◦32’S–44◦15’W73 r L a m m Table1.Su Site ´Nahuelquın2 ´Nahuelquın3 IslaAcuao1 IslaElena1 ´IslaBenjamın 01 ˜IslaGoni1 SenoCanalad 8

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