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NEW LYONOTHAMNEAE, A THE ROSACEAE TRIBE IN (ROSALES) Luc Brouillet notably the Potter (2007) paper, have changes e, et al. led to in the Molecular phylogenetic analyses angiosperms more of the and, of Rosidae Tahkht. (summarized specifically, in Angiosperm Group Phylogeny and Morgan 2003) Rosaceae 1994) have changed Juss. substantially (e.g., et al. our perception of the relationships of Rosaceae, and have changes resulted in slight to circumscription. its & now Traditional Rosales Bercht. Presl Cronquist 1981) included families that are dispersed (e.g., in J. many and now orders subclasses (Angiosperm Phylogeny Group 2003). Rosales has circumscription a that & members includes former Rhamnales Linnk and of the ex Urticales Bercht. (Angiosperm Juss. Presl J. Phylogeny Group new 2003). Similarly, phylogenetic analyses have led to a classification of the Rosaceae with (Potter et al. 2007), a reorganization of subfamilies and, to a lesser extent, tribes. Hypotheses of rela- among and tionships tribes subfamilies were considerably modified One may also as a result of this analysis. compare, Kalkman for instance, the traditional classification used by (2004) to that of Potter (2007): et al. the classic subfamilies Spiraeoideae Arm, Rosoideae Arm, Prunoideae Horan., and Maloideae Weber, rec- ognized were in the former, reduced three in the Rosoideae, Dryadoideae and to Sweet, latter, Spiraeoideae. now The subfamily last incorporates genera formerly included in three subfamilies, including Prunoideae and Maloideae. In the phylogenetic analysis of Potter et (2007), Lyonothamnus A. Gray appeared the al. at members base of Spiraeoideae, sister to all other of the clade. In their classification, Lyonothamnus was not assigned Lyonothamnus has been to a tribe. Historically, considered part of tribe Quillajeae Endl., along with now Don Quillaja Molina. Quillaja is treated as the only genus in Quillajaceae D. of the Fabales Bromhead (Angiosperm Group Phylogeny The and 2003). subfamilial tribal classification of Potter (2007) be- et al. is used ing as the basis for treating Rosaceae in the Flora of North America project (volume Rosales in part). 9, name no am Currently, tribal exists for Lyonothamnus (Pankhurst 2007; Reveal 2006). Therefore, proposing I the creation Lyonothamneae, accomodate of a tribe, to Lyonothamnus. Lyonothamneae nOV Tribus Brouillet, tribus Lyonothamnus Amer. Type: A. Gray, Proc. Acad. Arts 20:291. 1885. mple compound; or caducous, stipules free; \ Journal of the Botanical Research Institute of Texas 2(1) 386 hypanthium campanulate; torus none; carpels distinct, 2(-3), perigynous; epicalyx of (l-)2-3 bractlets; 2-4 more, Fruits aggregated (follicetum), styles ovules or clustered. follicles terminal, distinct; styles free, x = tardily falling, 9. )hn Strother for their helpful c< Angiosperm Phylogeny Group the orders An update of the classification for 2003. Angiosperm Phylogeny Group. APG Soc :399-436. and 141 families of flowering plants: Bot. Linn. J. II. New Columbia An system of of flowering plants. University Press, AJ. 1981. integrated classification Cronquist, York. Flowering Dicotyledons: Celastrales, Oxalidales, Rosales, Kalkman, 2004. Rosaceae. Kubitzki, ed. plants. C. In: K. The and genera of vascular plants. Berlin. Pp. Comales, families Kubitzki, series ed., Ericales. Vol. VI in K. 343-386. sequence Morgan and 1994. Systematic and evolutionary implications of rbci Robertson. D.R., D.E. Soltis, K.R. Amer. :890-903. variation Rosaceae. Bot. 81 in J. Orga- Lyonothamnus. Rosaceae database. Provisional global plant checklist. International 2007. Pankhurst, In: R. [http://bgbm3.bgbm.fu-berlin.de/iopi/gpc/PTaxonDetail.asp7NameCa nization for Plant Information (DPI). che=Lyonothamnus&PTRefFk=1 8 (accessed Oct. 2007)]. 2, TA M. Morgan, M. Robertson, Arsenault, Dick- Potter, D.,T Eriksson, R.C. Evans, S.-H. Oh, J.E.E. Smedmark, D.R. Kerr, K.R. 266:5-43. and Campbell 2007. Phylogeny and classification of Rosaceae. PL Syst Evol. inson, C.S. nominum supragenericorum plantarum vascularium. [http://www.pl a nt- 2007. Rosaceae. Indices Reveal, J.L In: systematics.org/reveal/pbio/fam/famRS.html (accessed Dec. 2007)]. 1 1,