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3 Vol. 12, No.2 The Quarterly Newsletter ApriTJune 2002 Why is DwarfPitch Pine Dwarf? WeiFang Theseexperiments are still continuing. To Jessica Gurevitch date,theoverwhelminginfluencesonthesizeofpitch pineseedlingsandsaplingshavebeenduetostrictly environmentalfactors. Seedlingsgrewfaster,survived LongIslanddwarfPitchPine(Pinusrigida) better, and began to bear cones earlier in normal areaformofPitchPinesthatgrowwithintheDwarf staturedsitescomparedtothoseplantedindwarfsites, PinePlains(onceanareaof2,500acres) inthecen- for both plants from dwarf and normal-statured tralPineBarrensofLongIsland,NewYork. Instead parents. Theenvironmentalfactorsresponsiblefor ofgrowinganuprightsingledominanttrunk, as do these differences are probably soil nutrient and normal-staturedPitchPines,theDwarfPines send moistureretentiondifferences.However,thereisalso multipletrunks fromtherootcrown. Themultiple apossibilitythattherearegeneticdifferencesbetween trunksgrowupwards,outwardsoralongtheground, dwarf and normal-statured pines, because the andarestuntedandgnarled. TheseDwarfPinesare seedlingsfromdwarfparentsgrewbetterthanthose usuallynomorethan2meterstall,whilethenormal- fromnormal-staturedparentsonthedwarfsites,while staturePitchPinegrowingcanreach20meterstall, thosefromnormal-staturedparentsgrewbetterthan althoughtheyareusuallyshorteronLongIsland.Dwarf thosefromdwarfparentsatthenormalstaturedsites. PitchPineshaveonlybeenfoundinthreeplaces in (ContinuedonPage 16) New the world: in the Jersey Pine Barrens, in the ShawangunkMountainsinupstateNewYorkandon LongIsland. MostoftheresearchonDwarfPitchPines wasconductedontheNewJerseyPineBarrenspopu- lation. Itisnotyetdefinitivelyestablishedwhether geneticallydistinctgenotypesproducedwarfandnor- mal-staturedgrowthforms, orwhetherthesediffer- encesareduetoplasticityinresponsetoenvironmental factors. Ifitisthelater,itisnotknowntowhichenvi- ronmentalfactorsthePitchPinesmayberesponding. Startingin 1997, weaddressedtheseques- tionsinLongIslandPinePlainsandBarrenswithtwo reciprocaltransplantingexperiments. Bytransplant- ingtheseedlingsstartedinthegreenhousefromdwarf or normal-statured trees, into dwarfand normal- staturedsites inthe field,wehopetoteaseapartthe effectsofgeneticandenvironmentalfactorsonseed- 111.: Audrey Watson-Wigley lingperformance. Pitch Pine {Pinus rigida) Page 1 Long Island Botanical Society Society News Founded: 1986 Incorporated: 1989 Courting Grandifolia: InFebruary of2000, afternumerous town The Long Island Botanical Society boardmeetingsrepletewithpleas ofpreservationfromenvironmental is dedicated to the promotion of groups from across Long Island, the Town of Riverhead granted field botany and a greater under- TalmageandBakstpermissiontorunbulldozersthroughtheGrandifolia standing of the plants that grow Sandhills inBaitingHollow. Thesepartners ofthe“Traditional Links wild on Long Island, New York. GolfCourse”promptlybegancarvingup the rarenativehabitat. Now, two years later, the Appellate Division ofthe Supreme Court Executive Board ofthe State ofNew Yorkhas foundthat “the Town ofRiverhead has President brokenthelawandimproperlysegmentedthereviewprocessmandated Eric Lamont (631) 722-5542 bytheStateEnvironmentalQualityReviewAct(SEQRA)”. Thedecision VicePresident Skip Blanchard (631)421-5619 washandeddown,unanimously,bythejudicialpanel. Theincomplete Treasurer golfcourse andthe scarred sandhills await furtheraction. Carol Johnston (516) 676-6648 HauppaugeHerald: The efforttopreservethe Hauppauge Springs Recording Secretary continues. SuffolkCountyhasmadeits offertothethree owners. The Barbara Conolly (516) 922-5935 Corresponding Secretary owners have not accepted the offers. The situation is in stalemate. John Potente (631) 361-6756 John Potente, Directorofthe Hauppauge Springs Coalitionwill give apresentationtotheHauppaugePTAattheHauppaugeMiddle School Committee Chairpersons onthe comerofTownLineRoadandLincolnBoulevardonTuesday, LocalFlora April 16 at 7:30 PM. All are welcome. Steven Clemants (718) 623-7309 Belauding Beitel: Joseph Beitel, a founding member ofthe Long FieldTrips Island Botanical Society and its first Vice-President, is remembered JennyUlsheimer (516) 625-0989 asanenthusiasticandaccomplishedbotantist. Whilegoingtocollege,he Programs Rich Kelly (516)354-6506 workedduringthesummersforSuffolkCountyParks. Heinventoriedthe Membership fernsandidentifiedmanypreviouslyunknownvarietiesoccurringonLong Lois Lindberg (516) 922-0903 Island (andNew York State) at Montauk County Park. He ledmany Conservation stimulatingfieldtripsfortheLongIslandBotanicalSocietyandpassed John Turner (516) 829-3368 away in 1991 while workng as abotanist at the New YorkBotanical Karen Blumer (631) 821-3337 Education Garden. LIBSinitiatedaprojecttoplaceamemorialplaque,inhishonor MaryLauraLamont (631)722-5542 atBigReedPond(nowcalledTheodore RooseveltPreserve). Karen Thomas Stock (631) 727-5250 Blumer, Conservation Chariperson for LIBS, has contacted Suffolk Hospitality CountyCommissionerofParksPeterScullyforpermissiontomountthe Jane Blanchard (631)421-5619 plaqueuponanappropriateboulderatthesite. BothKarenandBarbara BettyLotowycz (516) 676-2047 Zu Proly (516) 676-4023 ConollywerecontactedbytheSuffolkCountyDepartmentofParksand Editor RecreationandnotifiedthatLIBSisauthorizedtoplacethememorial. It John Potente (631)361-6756 willread: “JosephM.Beitel, 1952-1991,Talentedbotanist,Giftedteacher, Valuedfriend. Werememberhimhereinoneofhisfavoriteplaces. Long Membership IslandBotanicalSociety.” AnnualDuesof$15payableto: LongIslandBotanical Society Mailto: Plants in the News LoisLindberg,MembershipChairperson Mars Trek: TheNationalAeronautic and Space Agency (NASA) 45 Sandy Hill Road OysterBay, New York 11771-3111 is currently assessing he possibility of life on Mars. A $1 billion Article & News Submissions experiment placed on Mars in 1976 by the Viking mission is now LongIslandBotanical Society yielding results. The Viking probes placed nutrients in Mars soil P.O. Box 5001 samples and detected gas emissions related to the metabolism of Hauppauge, New York 11788 microorganisms. The recent detections ofa rhythm in the readings [email protected] are consistentwith life as we know it. Uponhearing ofthe findings, PrintedbyNativeAmerica LIBSbeganplanningforamartianfieldtrip. Page 14 5 Letters to the Editor That “Top Twenty Invasive Plants inNew York State” list (Jan.-Mar. 2002) is an excellent summary—looks likeyouroundedup alltheusual suspects”andmore! Noquarrelfromme-although I don’tmindBlackLocust...initsplace. But, I am wondering ifthere might be another potential problem species or two. I first became familiar, not long ago, with Yam-leaf Clematis (<Clematis terniflora) around my former house in Greenport (North Fork)—not a lot and not an unattractive “escape”. Last season, I saw several Yam-leaforSweetAutumnClematis(Clematisterniflora) patches of ternifolia up the Hudson, around Constitution Island. So far, no “menace”, I thought... unlike that damn Swallow-wort Cynanchum ( ). However, on a trip to southern New Jersey last September8,1was startledtoseethisplantliterally all over Cape May County. It was in flower, so thatmanyormostoftheroadsideedges (atvarying heights) seemedasiftheywereblanketedinsnow! True, most ofmy previous trips down there have been later in the month when I would not have noticeditso obviously. Perhaps, ithasbeenthere alongtime? Idon’tknowwhetherornotthisvine is spreading Map ofrecognized sites ofYam-leafClematis rapidly on Long Island (orNewYork State),but I think it can surely be considered a serious New “invasive species”,atleastinsouthern Jersey at present. I’d be interested in you reaction/ comment. Say, while I’m at it... isn’tthere another one we should keep a wary eye on—that scourge of the WesternPlains, Leafy Spurge Euphorbiaesulafl ( Perhaps not on Long Island, but upstate surely a potentialthreat, don’tyouthink? GuyTudor, NewYorkCityButterflyClub Editor’s note: I am all too familiar with Clematis terniflora and Euphorbia esula. Bothhaveinvadedandaregrowingatgreatspeed on my own property in Hauppauge. Both seem to enjoy temperate climate and are entrenching themselves on Long Island. Leafy Spurge (Euphorbia esula) Page 1 (ContinuedfromPage Effect ofLight Competition 13) Anothersuggestionthattheremaybesome upon the Dwarf Growth Form geneticdifferencesbetweenthetwogrowthformis justbeginningtooccur,asthesaplingsbegintoreach of Pitch Pine reproductivematurity(about4-5 yearsold). Plants fromdwarf populations havebeguntobearcones earlierthanthosefromnormalstaturedpopulations. Meryl del Rosario Sofar, onlypinesplantedintothenormal statured AndrewHarrison siteshavebeguntoproducecones. Herbivoryisalsoimportanttoseedlingper- formance. Pine seedlingsthathadbeenbrowsedin The dwarfgrowth form ofthe Pitch Pine (Pinus early age tendtobe shorter, fork earlier and grow rigida),whichisreadilyobservedinthedwarfpine lessuprightthanseedlingsneverbrowsed. Buther- plains of Long Island’s Central Pine Barrens, is bivoryhistoryisnotacritical drivingforce forthe consideredtobegloballyrare. Amongitsdistinctive perpetuationofpitchpinedwarfism. Firefrequency characteristicsareacontortedtrunkandlowcanopy canbe theconsequence ofdrierandpoorsoil. Itis heightwhichranges from 1 to 3 meters (Reschke, possiblethatfrequentfiresmayleadtothecreationof 1990). Incontrast,maturetreesexhibitingthemore thegnarled,multitrunkformofDwarfPitchPineby common,non-contortedgrowthformusuallyrange repeateddestructionofthesaplingleaders(orgrow- inheightfrom 15to25meters,withamaximumof30 ingtip,theapicalmeristem).Thathypothesiswillre- meters(BumsandHonkala, 1990). maintobetestedinthefuture. Inanycase,itisclear Establishingthecauseofthedwarfgrowthformhas thattheDwarfPinePlainsareauniquesiteinwhich beenthe subject ofconsiderable debate. Possible thedwarfpinesareabletogrowandpersist,whether genetic differencesbetweenthegrowthformsand ornotthesepopulationaregeneticallydistinctaswell. environmentalfactorssuchassoiltypeandchemistry, windandlightcompetitionhaveallbeenproposedas theprimaryagent. Previousresearchhasshownthat soilcharacteristicsinstandsofnon-contortedpines are similartothose indwarfstands (Black, 1997), andthusitislikelythatsoilpropertiesarenotadirect cause ofthe dwarfgrowthform. The focus ofthis studywastheeffectsoflightcompetitiononthegrowth ofPitchPine. We chose as our study area a firebreak that intersectsastandofdwarfPinesattheAirNational GuardfacilityinWesthamptonBeach. Predominantly contortedtreesinhabittheareastothenorthandsouth ofthe firebreak, while pines with straight trunks Wei Fang and Jessica Gurevitch ata study site populatethe firebreakitself Theareas outsidethe inthedwarfPinePlains ofeasternLongIsland firebreakarecharacterizedbyanunderstorythatis dominated by Scrub Oak (<Quercus ilicifolia). However,onlyaverysparsepopulationofScrubOak Editor’snote: Thisarticlewassubmittedforpubli- is present within the area of the firebreak. We cation in LIBS on March 1, 2002. examinedthisassociationofScmbOakwithdwarf, JessicaGurevitchisProfessoroftheDepartmentof but not the non-contorted. Pitch Pines and EcologyandEvolutionatSUNYatStonyBrook. hypothesizedthatthedwarfgrowthforminpitchpine WeiFangiscurrentlyworkingonherPh.D.inEcol- resultsfromcompetitionwithscmboakforavailable ogy & Evolution at SUNY Stony Brook light Page 16 Ateachofeightsites inandaroundthefirebreak Oakincreased, the C-indexalso increased. Since wecollecteddataontreeheights,trunklengthsand C-indexisameasureofthedegreeofcontortionin percentscrub oakcover. Inthesites outsidethe pines, andscrub oakpercentageprobablyreflects firebreak, wealsonotedpresenceorabsenceof theamountoflightcompetitionthatthepinesface, opencones. Animportantadvantagethatthe the logicalconclusionweinferredfromourdatais firebreakofferedwasastudysitewasthatit thatobservedcontortionisdirectlycorrelatedwith seemedtominimizethepotentialforgenetic thedegreeoflightcompetitionexperienced. Away differencesbetweenthedwarfandnon-contorted to confirmthis hypothesiswouldbetoperforma trees, sincethemostlikelyseedsourceforthepitch factorialexperiment;i.e.,growtomaturityseedlings pinesinthefirebreakwasthesurroundingdwarf fromnon-contortedparentsundershadedconditions pinecommunity. andseedlingsfromdwarfparentsinfulllight(aswell Our study sites 1 through 7 were located in this as the reciprocalpairings). The result from such dwarfpinecommunity. Theaverageageofthetrees long-termexperimentmightfinallyprovideadefini- herehadpreviouslybeendeterminedtobe33years tiveexplanationforthecurious,butcontroversial, (Black, 1997). Studysite 8, astandofyoungpitch dwarfpinegrowthform. pines,waslocatedinthefirebreak. Onlytwoscrub oaksinsite8weretallerthan3feet(mostwereshorter than .5 meter) and they were not located in close proximitytoanypines. Themajorityofthepinesin References: this site were too small to bore and thus we aged thembycountingbranchwhorls. The largerpines BLACK, J.A. 1997. Forest Composition, Sedi- wereagedbystandardmethods,inwhichcoreswere mentandChemicalCharacteristics: Westhampton obtainedandtreeringscounted. BeachAirNationalGuardSiteandAdjacentWood- The age datawe obtainedinthis study, together lands. UnpublishedReport,Dept, ofDefenseLeg- withthepreviouslyobtainedagedatafromsites 1-7, endGrant. supportourassumptionthatthe dwarfpineswere the seedsource forthenon-contortedpines inthe BURNS,R.M. andHONKALA,B.H. (Technical firebreak. The oldestpineinthefirebreakwasap- coordinators) 1990. Silvics ofNorthAmerica: 1. proximately26yearsyoungerthantheaverageage Confers. Agricultural Handbook 654. ofthesurroundingdwarfpines. Also,openpinecones U.S.Department ofAgriculture, Forest Service, werepresenton90%ofthe surveyeddwarfpines. Washington,DC. Sincetherewerenotreesinthefirebreakthatwere significantlyolderthantherest,itseemunlikelythat RESCHKE, C. 1990. EcologicalCommunitiesof one(orseveral)seedlingfromnon-contortedpines New York State. New York State Natural Heri- tookrootthereandthenbecametheseedsourcefor tageProgram. NewYorkStateDepartmentofEn- theremainingtrees. vironmentalConservation,Albany,NewYork. Priortothis studytherehadbeennoattemptsto quantifythedegree ofcontortioninthedwarfpine growthformandrelateittopossiblecommunityin- teractions. Thereforewe developedwhatwe call theC-index. TheC-indexisaratioofthetrunklength Editor’s Note: Thisarticlewassubmittedforpub- ofapinetotheheightofitscanopy. Theindexranges lication to LIBS on June 19, 2001. upwardfromaminiumvalueof1 (indicatingatree CharlesA. HarrisonwasaResearchAssociateat withoutcontortion)toatheoreticalmaximumofin- the CenterforCommunityResearchat Suffolk finity. C-indexvalues increasewiththe degree of Community College in Selden, NY. contortion. MeryldelRosariowasaffiliatedwithBellportHigh We observed a clear relationship between the School/ Suffolk Community College percentage ofscrub oakatthe siteandtheaverage SummerFieldProgram. C-indexvalueforthatsite. WhentheamountofScmb Page 17 8 Pitch Pine, Hauppauge, Long Island, New York Page 1 Potente E. John Dwarf Pitch Pine, Westhampton, Long Island, New York Page 19 Abandoned Scrub Oak Gall Refugia Inorderto determinethe diversityofthe second- arycommunityinscrub oakgalls intheDwarfPine Amy Ho Plains,theteamgatheredpost-emergencegallsfrom Hannah Kang the ScrubOakshrubs. Duringthe summersof2000 and 2001, a total of 148 samples were collected. Krista Biendenbach Preservationoftheanimalswasaccomplishedbyplac- ingthegalls inseparatebags atthe site. Theywere Scrub Oakgallsareformedwhenacynipidwasp laterstoredinafreezertopreservethegallcontents. oviposits her eggs on a selected oak leaf. The The galls ofeachshrubwere dissectedandgalldi- gallsthemselves areproducedinresponseto spe- mensionsandinnerchamber/outerexitholesizeswere cific plant growth regulating chemicals or other measured. The animals were then taken out and stimuliproducedbythe insects. Itis thoughtthat placedinsmallvialsof70%alcohol. these chemicals change the DNA composition of Artificialplasticeggs(simulatingnaturalgalls)were the plant, thereby forming a hardened structure. placedonthe scrub oakplants. (Figure 2). Seven- “Tiny flies in the family cecidomyidae, order teeneggswerecamouflagedwithgreenpaint Exit diptera, inducegalls inmembersofabout50plant holeswerepuncturedintotheartificialeggstoresemble families. The othermajorgall inducing family is exitholes ofagall. Contents ofthepseudo-havens cynipidae, order hymenoptera, a family of tiny werecheckedaftera21-dayinterval. wasps. The cynipidaehaveamorerestrictedhost Measurements ofthecapsuleholewerecompared range,beingfoundonlyonoaksandspeciesofthe to those of the gall emergence hole. The graph rosefamily. Therearemorethan700kindsofcynidpid (Capsule Hole vs. Exit Hole; Graph 1) gives a waspsthatinducegallsonoaksintheUnitedStates.” slope ofless than one. This indicates thatthe exit (F.W. Howard). The Scrub Oakgallthatisstudied holes tend to be larger with respect to the capsule isaproductofthecynipidaeAmphibolipsilicofoliae hole. Thissuggeststhattheanimalmayhaveunder- onScrubOak(Quercus ilicifolici). gone atype ofmetamorphosis afterhatching. The The ScrubOakgalliscomposedoftwowalls:the animalmayhavebeenengagedinswelling,growing, outergallshieldthatisvisibleandtheinnerportion orhardening. Numerousgallswere foundwithout thatisasmallcapsule.Theun-emergedcynipidwasp gallemergenceholes (notshownongraph 1) indi- dwellsinsidethecapsule. Afterhatching,itwillleave catingthatthewaspemergedfromthecapsule,but thecapsuleandthegall. Ahardenedstructureisleft: didnotleavethegall. Therewere alsogalls found thegallwithaninnercapsule. Twoholesremain: the with exit holes that did not have a capsule hole emergenceholeandthecapsulehole. Thisenables (not shown on Graph 1) . This is evidence ofthe otheranimalstoutilizethegallasarefuge. presence of an invader. Questions as to whether Theaimofthisstudyistoinvestigatethesecond- the invaderwaspresentbefore orafterthepupae’s arycommunitythatinhabitstheScmbOakGall. Ar- demise beginto arise. tificialgallrefugeswerealsointroducedforcompari- sonstudy. The studytookplaceintheDwarfPine PlainsofWestHamptonBeachonLongIsland. ScrubOakleafgall Graph 1 ScrubOakCapsuleHolevs.GallEmergenceHole Page 20 1 Themainincentivetothepresence ofallthe spe- After placing artificial galls on the shrubs, we cies is the nutritious consumption products. This waited for 21 days. At the end ofthat time, the includesnutritivetissuesforthecynipidwasp,mite, gall’s capacity forhousing animals was checked. beetle and larvae. Mites and thrips, along with Only 1 of 17 galls was inhabited. Fourteen adult thecynipidwasp feedonthisnutritious galltissue red ants and one larval ant seemed to have fa- to survive. These two families make their own voredthe single artificial gall. Ithadbeenplaced galls and they come into this gall formore access inanopenareanearmanyantmounds. Whenthe to gall tissues, since their own gall has its own pseudohavenswereretrieved,theantshadalready community of secondary inhabitants. died. (Ananthakrishnan, 1992;Westphal, 1992). Asshown The invention was made to resemble the actual inTable 1,miteswerethemostcommonspecies in gall. The texture ofthe artificial gall wasplastic. thegallrefiigia. Miteswereoftenfoundinsideagall This isadifference ofnature’sprovisionsvs. man- withotherspecies. This indicatesthatthemitesdid madematerials. notpreyupontheotherspeciestheywerefoundwith. The gall itselfis a very complex structure pro- Bycontrast, antsarepredators althoughtheymay duced by a living organism. The occupation of be preyed upon themselves. Spiders appeared to the abandoned scrub oak gall shows great diver- preyuponants. Antswereplentifulinthesecondary sity for such a smallbiological structure. community(Table 1). Another finding was the presence ofnumerous References: larvaeleftbehind,manytimeswithoutparents. This showsthatthegallisnotonlyahomeforthecynipid BLAND, Roger G., How to Know Insects. Third Edition. Central Michigan University. Wm. C. Brown Co. Pub. wasp,butalso arefuge forthe larvae ofotherspe- BORROR, Donald J. & TRIPLEHORN, Charles A. & cies. The larvae have abetter chance ofsurvival JOHNSON,NormanF. 1982. An IntroductiontotheStudy inamicrohabitat,ratherthanasitethatismore ex- ofInsects: SixthEd.OhioStateUniversity. posed. Oftenthe larvae areconsumedbythepara- FORMAN, Richard. 1979. Pine Barrens: Ecosystem and sitizingwasp Chalcidoidea Torymidae. Thiswasp, Landscape. Academic Press NY, San Francisco, London. ROHFRISH, O. 1992 Patterns in Gall Development, Bi- part ofthe secondary community, acts to keep in ology of Insect Inducted Galls. J.D. Shorhouse and O. checkthepopulationofcynipidsinandarea. Rohfrish.NewYork,OxfordUniversityPress:60-86. ROSOMER, Wm. & STOFFOLANO, J., 1998. The Sci- enceofEnt.: FourthEd. OhioUniv.andUniv. ofMA. ROSS, H. & ROSS, C. & ROSS J. 1982. A Textbook of CommonName ScientificName Abundance Entomology: Fourth Edition. University of Georgia and Western Washington University. John Wiley and Sons. Ant FormicidaeMyrmicinae 29 SOKAL, R. R. & BOWERS, M. 1995 Biometry, W. H. Ant-like LeafBeetle ColeopteraEuglenidae 21 FreemanandCompany.NewYork. Beetle Larvae 4 SOUCY, S. &ROOT, K. 1998. ConservationBiologywith Bristletail ThysanuraJapygidae 4 RAMASBiolab.AppliedBiomathematics.Setauket,NY. CCaatreprepitllBaeretles TCoorlteroipstiedarea Dermestidae 11 TOWNSEND,C.J.&BEGON,M.&HARPER,J.L. 1996. CobwebWeaverSpider AraneidaTheridiidae 3 Interactions. Ecology-Individuals, Populations and Com- CommonThrip ThysanopteraThripidae 13 munities:ThirdEdition, 211-482. Lady-birdBeetle ColeopteraCoccinellidae 1 VON ENDE, Carl & BROWER, James & ZAR, Jerrold. LLeaarfvaoerPlantBugs Hemiptera Miridae 402 1998. General Ecology: Fourth Ed. WCB McGraw Hill. Leaf-Hopper CicadeilidaeHomoptera Nymph 1 WENIG,J. 1976.AnInt.toEnv.Sci.TheEcologyofLI.Env. Mites Arthropoda,AcariOribatida 42 Pub.Ass.LTD.p. 10. Spider AraneidaDictynidae 5 WHEELER, A.G. 1991. Plant Bugs ofQuercus Ilicifolicr. WUnaisdepntifiedInvertebrates HymenopteraCynipidae 21| Myriads ofMirids in Pitch Pine-Scrub Oak Barrens. HemimetabolistNymph 1 Hemiptera Nymph 1 Amy Ho, Hannah Kang and Krista Biendenbach attained semifmalist statusupon submission ofthis project in an Intel High School Competition Table1. TableofSecondaryCommunityInhabitants Page 2 Additional Notes on Maritime Oak-Basswood Forests Philip Marshall In belated response to articles by Eric Lamont significant Long Island Native American archeo- (1997) and David Kunstler (1998), on Maritime logical site. Couldtherebe aconnectionbetween Oak-Basswood forests of Long Island and humanoriginforthedisjunctdistmbutionofBlack WestchesterCounty,NYIwouldliketooffersome Walnut(Juglansnigra)inupstateNewYork. Why thoughts and observations on a similar associa- notthe same for Basswood (whose phloem fibers tionfoundonthenorth side ofLongIsland Sound were valued for cordage) on the shores ofLong onChaffinchIslandinGuilford, CT. Island Sound? (I admit it is unlikely that such is Located almost directly across central Long Is- the caseonSouthDumplingIsland!) landSoundfromRoanokePoint inRiverhead,NY, Chaffinch Island is a small island (I estimate itto be no more than one hectare in area) now con- nected by a narrow causeway to the mainland at MulberryPoint. Althoughconsistingofalump of granite bedrock protruding from the level ofthe surrounding Spartina saltmarsh, thus differing markedlyfromtheLongIslandlittoral, Chaffinch Islandoffers aminiatureflorathatis immediately familiar to the Long Island botanist. The forest cover is mainly Oak (Quercus alba, Q. velutina, Q, coccinea), with a small number ofRed Cedar (Juniperus virginiana) and Pitch Pines (Pinus rigida) and an understory of Huckleberry Potente (Gaylussacia baccata Lowbush Blueberry ), E. Vacciniumpallidum BlackChokeberry Aronia ( ), ( melanocarpa Bayberry(Myricapennsylvanica), John ), andPoisonIvy Toxicodendronradicans). Inone American Basswood (Tiliaamericana) ( sunlit spot can be found Prickly Pear (Opuntia humifusa)andPastureRose (Rosa Carolina and ), atthewater’s edgeareBeachPlum (Prunusmar- itima Hackberry (Celtis occidentals), and the References: ), great curiosity, several gnarled and stunted indi- viduals ofAmericaBasswood (Tiliaamericana). Kunstler,D. S. 1998.AmericanBasswoodonHuck- The soil ofthe southern tip ofChaffinch Island leberryIslandinWesternLongIslandSound. LIBS appears tohavebeenformedfromagiantmidden Newsletter8: 3-4. of clam and oyster shells certainly of Native Americanorigin, andtheresultinganthropogenic Lamont, Eric. 1997. TheMaritimeOak-Basswood adjustment ofsoil pH (which I have inferred, not ForestonLongIsland’sNorthFork. LIBS News- actuallymeasured)maypartlyexplainthemodem letter7: 27-28. presence ofTilia americana on the site. But only partly. Still, I recall having seen T. americana in Wykoff, M.W. 1991. Black Walnut on Iroquoian asimilarmaritimepositionatthe ChandlerEstate landscapes. Northeast Indian Quarterly, Summer in Mount Sinai, NY, which is known to be a 1991: 4-17. Page 22

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