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Long Island Botanical Society Vol. 20 No. 1 The QuarterlyNewsletter Winter2010 Checklist of the Bryophytes of Long Island EricC. Morgan1 Jon A. Borysiewicz1 and Jon A. Sperling2 , BartlettArboretum Herbarium, 151 Brookdale R<±, Stamford, CT 06903 2Queens College Biology Dept., 65-30 Kissena Blvd, Flushing, NY 11367 Thebryophytes,commonlyknownashornworts,liverworts name.Withonlyahandfulofhornwortspeciesintheregion,an andmosses,arenon-floweringplantsthataredispersedbyspores amateurbryologistcaneasilylearntorecognizetheseplants. as opposed to seeds, and can be found in a tremendous array Liverworts are alargegroupwhich can be separated from of habitats, often with impressive diversity and abundance. mosses by their absence of leaves, orif leaves are present, by Whilemoderncladisticshas separatedthebryophytesintothree the arrangement of those leaves. Leafy liverworts have leaves groups,theAnthoceratophytaorhornworts, theHepaticophyta arrangedintwo orthreerows, usuallyon creepingstems,while orliverworts, and the Bryophytaormosses, forthe textof this mosses have theirleaves spirally arrangedwith few exceptions. paperthe termbryophytewillbeused Amongleafyliverworts, the leaves are inthebroadsense. never found on the upper side of the With over 15,000 species stem, which usuallygives the plants a worldwide, the bryophytes make up a flattenedappearance.Theleavescanbe sizeable portion of the approximately entire,lobed,ciliateorotherwise,justas 300,000 land plants estimated to be invascularplants.Manyleafyliverworts living today. There are about 1,500 canberecognized bywhatare known native vascularplants on LongIsland, as complicate bilobed leaves inwhich and the bryophytes may potentially thelobes are foldedontooneanother. haveover400species,bringingtheland Non-leafy or thalloid liverworts have plant total to nearly2,000. In spite of a flattened rosette or strip-like plant theirrelativelysmallsize,arequirement bodywhich is not divided into stems due to their lack of a developed andleaves. vascular system, the bryophytes playa Mosses are also divided into majorrolein ourplanet’s ecology. For subcategories. They are regarded many years the bryophytes, like the as either pleurocarpous, with stems lichens, have been used as indicators creeping or trailing and branches of environmental quality such as air numerous,oracrocarpouswithupright quality (Pott & Turpin 1996; Davis et stems (sometimesinconspicuouslyso), al. 2001). Evenmoreimpressiveis the andfewtonobranches.Forthecorrect CALLIERGONELLA factthatthereismorecarbonstoredin identificationofmossesandliverworts, asinglegenusofmoss.Sphagnum than CourtesyoftheFloraofNorthAmericaAssociation, aproperunderstandingofthesegroups , inanyothergenusofplantsworldwide illustratorPatricia Eckel isessential. (Clymo&Hayward1982). For over a century, the more familiar and conspicuous The three main groups of bryophytes are relatively easy vascular flora of Long Island has been well studied and to distinguish from each other based upon characters visible documented through both local floristic studies and larger with the naked eye or a hand lens. Hornworts are the least projects documenting entire families and their distributions encounteredof thesegroups as theytendtohave quite specific acrosstheisland.ThestudyofLongIsland’sbryophyteflorahas habitat requirements. Our local hornworts tend to have a beenmuchless exhaustive.While severallate 19thandearly20th distinctly flattened thallus, often in irregular-shaped rosettes, century studies such asJelliffe (1899) and Cain and Penfound usuallywith sporophytes present. These sporophytes, the spore (1938)includedthedocumentationofbryophytespecies,studies bearingstructures,resembleananimalhorn,hencetheircommon focused solely on bryophytes are nearly non-existent for the (Continuedonpg3) ) Page 2 Long Island Botanical Society Vol. 20 No. 1 Long Island News Society Botanical Society Founded: 1986 The LongIsland Botanical Societywas foundedin 1986, and to celebrate Incorporated: 1989 our25thAnniversarywearebeginningtoplana10-daybotanicalfieldtripto FloridaintheSpringof2011.LIBSmemberAnnJohnsonhasvolunteered ; TheLongIslandBotanicalSocietyisdedicated to serve as tripleader.Weplanto startintheFloridapanhandle andmake ; to the promotion of field botany and a ourwayintopeninsularFlorida. Staytuned formoreinformation. greater understanding of the plants that ; growwildonLongIsland,NewYork. LIBS has joined a coahtion of 33 other organizations that formed to ; expressoppositiontothe saleanddevelopmentofaparceloflandowned Visitthe Society’s Web site bySuffolkCounty.The250-acreparcel,whichiswithintheCarmansRiver www.libotanical.ora watershed, would be Hsted as surplus by the County which would then enterintoacontractfortheintensivedevelopmentof thistractformixed- Executive Board usedevelopmentknownas “LegacyVillage.” I • • President At LIBS’ annual “Members Night” on December 8, 7 members gave Eric Lamont 631-722-5542 presentations.MarionHubbardofferedadisplayoninvasiveplantspecies; \ • AndyGrellershared shdes of beeorchidsinGreece;PeterWarnyshowed • Vice President Andrew Greller 516-364-9377 highhghtsofhistripsinsoutheasternU.S. toCanada;RayWelchproduced \ “The Waning of the Season” with music; Marilyn Jordan showed the • Treasurer Florida panhandle spider lily (Hymenocallis choctamnsis) which grows only : CarolJohnston 516-676-6648 in rockymarshes, and thewaterprimrosewhich has invaded the Peconic RecordingSecretary RiversysteminForgePondbutwhichhasbeensuccessfullyweededoutby Barbara Conolly 516-922-5935 volunteers; CarolJohnstonshowedafewshdes ofgrass-of-Parnassus and fringedgentianinsouthernVermont,andsharedherdiscoveryof therare CorrespondingSecretary (SI) stiff gentian (Gentianaquinquefolia inasmallquarryinNorthPownal, JohnPotente 631-361-6756 Vermont; and Rich Kellyshowed some eye-catchingmothsinEdgewood Preserve andelsewhere. Committee Chairpersons : LongIslandFloraCommittee—Update Flora j GerryMoore 718-623-7332 Afterthe untimelydeath of committee chairSteve Clemantsinlate 2008, j I FieldTrips the Flora Committee needed to regroup and push the Long Island Flora Atlas projecttowards completion. James Mickley 717-278-9126 j • Programs To this end, the committee met on December 1st, 2009 at the Brooklyn • RichKelly 516-354-6506 Botanic Garden (BBG). The attendeeswere GerryMoore, (thenewchair \• of the FloraCommittee), Barbara Conolly,AndyGreller,RichKelly,Eric Membership • Lamont, and A1 Lindberg. The feasibihty of continuing with the old • Lois Lindberg516-922-0903 process of manuallyplacing specimen and sightrecord data on maps, by Conservation township/county, was discussed. Since BBG’s New York MetropoHtan ; • BillTitus 516-671-7351 Flora (NYMF) database apparently includes Long Island flora data that • Education wasinputbyStevebefore his death, GerrysuggestedthatLIBS coulduse I MaryLauraLamont 631-722-5542 theNYMF datafromLongIslandforits atlas. This dataisby5-kilometer I mapblocks,sotherewouldbemuchmoreinformationonanyonespecies • l Hospitahty map. The numerous data points would almost approach a range map Kathleen Gaffney 516-742-4019 on Long Island. The NYMF database includes both specimen and sight j • ZuProly 516-676-4023 recorddata. Itwasalsodiscussedthataweb-basedatlaswouldbethebest • DorothyTitus 516-671-7351 approach. • • NewsletterEditor SteveGlennofBBGwillworkwithLIBStogettheprojectofftheground. ; Margaret Conover 631-357-3065 Agreatamountofworkwillbeneededtoproduceandarrangethespecies ; witShkisppec&ialJathnaenkBslatonchard wmhapisc.hwIitlwlabsescuogngdeusctteeddthaatttLheIBBSBhGiruesainggraadssotfutdweanrtetsoitpeeHrcfeonrsmetohwisnweodrbk,y : BBG. TheLIBSexecutiveboardapprovedtheexpenditureof $3000,plus Webmaster :• travelexpenses, forthiswork. DonaldHouse : [email protected] • ) Long Island Botanical Society Vol. 20 No. 1 Page 3 (Cont.frompage 1 1936a), and eastern Suffolk County (Biechele 1993), or a island, the one significantworkhavingincluded the entireNew focus upon vascular plants, with less emphasis upon the YorkGtyregion(Grout1916). bryophytes (Jelliffe 1899). Other studies such as Grout The last few decades have seen occasional papers on the (1916) and Ketchledge (1980), included Long Island as a subject such as Biechele’s (1993) work on the liverworts of part of a larger geographic area. Grout’s work includes the eastern Suffolk Countyand a statewide checklist of mosses by bryophytes of much of New Jersey, Westchester County, Ketchledge (1980)which,due to agrid systemformappingthe and even the Delaware Water Gap, regions likely to have state, lumped species foundin Staten Islandwith species from westernmostLongIsland,andlumpedcentralLongIslandwith portions of Westchester and Bronx Counties. Studies such as Table1.Numbersofbryophytetaxa. these canprovide abase fromwhichtobegin future studies of ReportsincludeallorpartsofLongIsland. thebryophytesofLongIsland,howeverwiththenarrowhabitat requirements of manyspecies, smaller-scale studies and species Source Hornworts Liverworts Mosses listsofnaturalareasarecurrentlyneededforaproperassessment Biechele (1993) 3 70 NA of bryophytes in our area. In the past fewyears severalpapers Cain(1936) NA 11 23 havebeguntoaddressthistopicthroughlocalfloras (Morgan& Cain&Penfound(1938) NA 28 48 Sperling2005),individualspecies distributions (Trigoboff2005) NA NA & Grout(1916) 370 andecologicalassessments (Morgan Sperling2007). FloristicworksonLongIslandbryophyteswillbecomemore Jelliffe (1899) 2 25 109 NA NA important as ecological changes occur throughout the region. Ketchledge (1980) 289 Withtheprospectofglobalclimatechange,bryophytesaswellas Currentstudy(2009) 4 78 226 vascularplantscanprovideindicationsastotherealityandextent ofthesechanges.Slack(1977)pointsoutthatsomecoastalplain speciesrange onlyas farnorthas LongIsland and are notseen different floras than those of Long Island and likely to be anywhereelseintheNortheast,apatternquite familiartothose majorfactorsinhis highnumberof taxaforthe area.Within studying vascular plant distributions. Species such as Telaranea Ketchledge’swork, thebryophytes thatmayhavebeen found nematodes,whosenorthernmostoccurrenceisintheLongIsland insouthwesternmostLongIslandareplacedinwiththoseof region, can provide valuable data towards an understanding of Staten Island, a separate entitywith avery differentvascular climatechange shouldthe abundanceof species suchas thisbe flora,andpossiblybryophyte floraaswell. alteredoverthecomingyears.Itwillbeimportanttomonitorthe In this contribution from the Long Island Botanical distributionofmorenorthernspecieswhichoccursparinglyon Society,theauthorsbegintoprovidedataregardingthenumber LongIsland. For these reasons, documentation of these plants of bryophytes found on Long Island, including both extant in individual parks and preserves on Long Island will become and historic species. This datais compiled from avariety of increasinglyimportantoverthenextfewyears. sourcesincludingpublishedliterature,herbariumrecords and Aside from using particular species as ecological field surveys. Severallocal herbariaincludingbut notlimited indicators, floristic change can be looked at on alarger scale to the NewYork Botanical Garden (NY), Brooklyn Botanic in bryophytes to determine larger trends in a given area. In Garden (BKL), Bartlett Arboretum (BART), and Queens a study of Alley Pond and Cunningham Park, in Queens College Biology Department (no official acronym) were County,LongIsland, thespeciesrichness,ortotalnumberof searched both online and in person. Each of these herbaria species present, dropped 36 percent for mosses, 58 percent holds significant collections of Long Island bryophytes for liverworts, and 50 percent for hornworts from the mid ranging from hundreds to thousands of specimens collected 1980s to the early part of this century (Morgan & Sperling onLongIsland overthe past 100-plusyears. Fieldvisits have 2007). These numbers provide strong evidence of a larger been made from 2002 to the present and have resulted in ecological shiftinthe area, although notnecessarilynegative. approximatelytwothousandcollectionsofbryophytesbythe Could this shift be due to forest maturation, increased park authorsandcoworkersonrelatedprojects.Allspecimenshave traffic,lowerrainfalls, highertemperatures ornone of these? beendepositedattheBartlettArboretumHerbarium(BART) These are questions thatwill need to be examined furtheras orthe Queens CollegeBiologyDepartment. more reliable species and distribution data are recorded for Nomenclature for this checklist follows that of the LongIsland. USDA/NRCS,Plants NationalDatabase (2009), ortheFlora In the past, reports of bryophyte numbers for Long ofNorthAmerica(2007)whenfamilytreatmentsareavailable. Island have been varied and incomplete (Table 1). This is Field andlaboratoryidentificationswere done usingavariety due to several factors includinga focus on particular smaller of guides including the Flora of North America (2007), regions of Long Island such as Cold Spring Harbor (Cain Andrus (1980),Crum(1983) andSchuster(1949; 1953). This (Cont.onpage4) . Page 4 Long Island Botanical Society Vol. 20 No. 1 checklistincludes a total of 68 families and 308 taxa, which Miller,N. G.andTrigoboff,N.2001.AEuropeanfeathermoss, can be broken down as follows; hornworts, 2 families and Pseudosckropodiumpurum,naturalizedwidelyinNewYork 4 species; liverworts, 26 families and 78 species; mosses, 40 Stateincemeteries.Bryologist104:98-103. families and 226 taxa. A full list of these taxais included as MorganE. C.andJ.Sperling.2003.Asurveyofthebryophytes Appendix 1. ofShuSwampPreserve,MillNeck,NewYork.Long IslandBot.Soc.Newsl. 13:25,28. LiteratureCitedandAdditionalReferences: 2005.ThebryophytefloraoftheCranberryBog . Andrus,R.E. 1980.Sphagnaceae (PeatMossFamily) ofNew Preserve,SuffolkCounty,NY,withnotesontheir YorkState.ContributionstoaFloraofNewYorkState. distribution.Evansia22:78-84. R.S.Mitchell,Ed.Bull.NewYorkStateMus.Sd.Serv No.442. .2007. Changesinthebryophytefloraof Cunningham Biechele,L.T. 1993.AchecklistoftheHepaticaeand ParkandAlleyPondPark,QueensCounty,LongIsland, AnthocerotaeofeasternSuffolkCounty,LongIsland, NewYorkCity.Evansia23:56-60. NewYork.Evansia10:99-103. Pott,U.andD.H.Turpin. 1996. Changesinatmospherictrace Britton,E. G. 1902.NoteonaLongIslandmoss.Bryologist elementdepositionintheFraserValley,B.C.,Canadafrom 5:66-67. 1960to 1993measuredbymossmonitoringwith Burnham,S.H.andR.A.Latham. 1914.Thefloraofthetown Isotheciumstoloniferum.Canad.J.Bot. 74:1345-1353. ofSouthhold,LongIslandandGardiner’sIsland.Torreya Ross,J.H. 1931.SomebryophyteassociationsofColdSpring 14:201-225. Harbor,NewYork,andvicinity.Bryologist34:88-91. Cain,S.A. 1936a.Thecompositionandstructureofanoak Schuster,R.M. 1949.TheEcologyanddistributionof woods.ColdSpringHarbor,LongIsland,withspecial HepaticaeincentralandwesternNewYork.Amer.Midi. attentiontosamplingmethods.Amer.Midi.Naturalist Naturalist42:513-712. 17:725-740. 1953.BorealHepaticae.Amanualoftheliverwortsof . . 1936b.SyrrhopodontexanusSull.inLongIsland. Minnesotaandadjacentregions.Amer.Midi.Naturalist.49: Bryologist39:118. 257-684. W and T.Penfound. 1938.Aceretumrubh:Thered Slack,N. G. 1977.SpeciesDiversityandCommunityStructure , mapleswampforestofcentralLongIsland.Amer.Midi. inBryophytes:NewYork Statestudies.Bull.NewYork Naturalist19:390-416. StateMus.Sci.Serv.No.428. Clymo,R.S.andP.M.Hayward. 1982.Theecologyof Sperling,J.A. 1958.Keytothebryophytesof Cunningham Sphagnum.In:BryophyteEcology,A. E.Smith,ed. Park.Nucleus2:9-13. J. Chapman&Hall,London,pp229-289. . (Dateunknown).BryophytesofLeedsPond, Crum,H. 1983.MossesoftheGreatLakesForest,ed.3. inVegetationalAnalysisofLeedsPondPreserveand UniversityofMichigan.AnnArbor,Michigan.417pp. CommentaryonHabitatRestoration,30pp. Davis,D.D., R.McClenahen,andR. Hutnik.2001. andE. C.Morgan.2003.Thebryophytefloraof J. J. , Useofanepiphyticmosstobiomonitorpollutantlevels CunninghamParkandAlleyPondPark,QueensCounty, insouthwesternPennsylvania.Northeast.Naturalist NewYork.Anewflorafromoldspecimens. 8:379-392. Evansia20:11-14. FloraofNorthAmericaEditorialCommittee.2007.Floraof Trigoboff,N.2005. Tortulapapilhsaand Tortulapagorum NorthAmerica,vol.27,Bryophyta,part1.Oxford (Pottiaceae)inNewYorkState.Evansia22: 85-89. UniversityPress,NewYork, (additionalmaterialpublished USDA,NRCS.2009.ThePLANTSDatabase onlineatfna.org) (http://plants.usda.gov.October2009).NationalPlant Grout,A. 1906.AdditionstothebryophytefloraofLong DataCenter,BatonRouge,LA70874-4490USA. J. Island.Bryologist9:26-28. Wickes,M.L. 1937.AnotherSyrrhopodononLongIsland, 1916.ThemossfloraofNewYorkCityandvicinity. NewYork.Bryologist40:71 . Authorpublished.NewDorp,NewYork. 121pp. Jelliffe,S.E. 1899.TheFloraofLongIsland.Privatelyprinted. Join LIBS today! 163pp. AnnualMembership is $20payable to: Ketchledge,E.H. 1980.Revisedchecklistofthemossesof LongIslandBotanicalSociety NewYorkState. ContributionstoaFloraofNewYork State,R.S.Mitchell,Editor.Bull.NewYorkStateMus.Sci. Mailyourdues to: CarolJohnston Serv.No.440. Latham,R. 1917.HabitatofCephaloziafranciscionLong LIBS Treasurer 347DuckPondRoad Island.Bryologist20:63-64. Mapes,H.M. 1962.SphagnumportoricenseandDiscelium Locust Valley NY 11560 nuduminNewYork.Bryologist65:68. NOTE: MembershiprenewalsaredueinJanuary Long Island Botanical Society Vol. 20 No. 1 Page 5 Appendix 1. Checklist of the bryophytes of Long Island, NewYork ANTHOCERATOPHYTA FOSSOMBRONIACEAE Telaraneanematodes ANTHOCEROTACEAE FossombroniacristulaAustin (GottscheexAustin)M.Howe AnthocerosmacouniiM.Howe FossombroniafoveolataLindb. AnthocerospunctatusL. Fossombroniawondracyekii(Corda)Dumort. LUNULARIACEAE Phaeoceroslaeiis(L.) Prosk. Funulariacrudata(L.)Dumort. GEOCALYCACEAE NOTOTHYLADACEAE Chiloscyphuspallescens MA11C1lANtilACEAE Notothylasorbicularis(Schwein.) Sull. (Ehrh.exHoffm.)Dumort. MarchantiapolymorphaL. Chiloscyphuspolyanthos Preissiaquadrata(Scop.)Nees HEPATICOPHYTA (L.) Cordavar.riiularis(Schrad.)Nees ANEURACEAE Geocalyxgraveolens(Schrad.)Nees PALLAVICINIACEAE Aneurapinguis(L.)Dumort. Harpanthusdrummondii(Taylor) Grolle Pallavidnialyellii(Hook.) Carruth. Riccardialatifrons(Lindb.) Lindb. Fophocoleaheterophylla(Schrad.)Dumort. Rdccardiamultiflda(L.)A. Gray FophocoleaminorNees PELLIACEAE Riccardiapalmata(Hedw) Carruth. Pelliaepiphylla(L.) Corda GYMNOMITRIACEAE PelliamegasporaRM.Schust. AYTONIACEAE Marsupellaemarginata(Ehrh.)Dumort. Pellianeesiana(Gottsche)Limpr. Asterellatenella(L.)P.Beauv. Marsupellasphacelata(Gieseke)Dumort. PLAGIOCHILACEAE BLASIACEAE JUBULACEAE Plagiochilaporelloides BlasiapusillaL. FrullaniaasagrayanaMont. (Torr.exNees)Lindenb. FrullaniabolanderiAustin CALYPOGEIACEAE FrullaniaeboracensisGottsche PORELLACEAE Calypogeiafissa(L.)Raddi Jubulapensylvanica(Steph.)A.Evans PorellapinnataL. Calypogeiamuelleriana(Schiffn.)Mull.Frib. Porellaplatyphylla(L.)Pfeiff. CalypogeiasullivantiiAustin JUNGERMANNIACEAE PSEUDOLEPICOLEACEAE Barbilophoyiabarbata CEPHALOZIACEAE (SchmidelexSchreb.)Loeske Bkpharostomatrichophyllum(L.)Dumort. Cephaloyiabicuspidata(L.)Dumort. Gymnocoleainflata(Huds.)Dumort. Cephaloyiacatenulata(Huebener)Lindb. Jamesoniellaautumnalis(DC.) Steph. PiiLlDlACEAE Cephaloyiaconnivens(Dicks.)Lindb. Ptilidiumpulcherrimum(Weber)Vain. Cephaloyialunulifolia(Dumort.)Dumort. RADULACEAE CephaloyiamacrostachyaKaal. Cephaloyiapleniceps(Austin)Lindb. Fophoyiabicrenata Radulacomplanata(L.)Dumort. Cladopodiellafluitans(Nees)H.Buch (SchmidelexHoffm.)Dumort. Cladopodiellafrancisci Fophoyiacapitata(Hook.)Macoun RICCIACEAE (Hook.)H.BuchexJorg. Fophoyiaindsa(Schrad.)Dumort. RicciafluitansL. Odontoschismadenudatum(Mart.)Dumort. Myliaanomala(Hook.)A. Gray RicciahuebenerianaLindenb. ssp.sullivantii Odontoschismaprostratum(Sw.)Trevis. Nardiainsectalindb. (Austin) R.M.Schust. Odontoschismasphagni(Dicks.)Dumort. Rdcdocarposnatans(L.) Corda LEJEUNEACEAE CEPHALOZIELLACEAE Fejeuneacaiifolia SCAPANIACEAE Cephaloyielladivaricata(Sm.) Schiffn. (Ehrh.)Lindb.emend.H.Buch Diplophyllumapiculatum(A.Evans) Steph. Cephaloyiellahampeana(Nees) Schiffn. Scapanianemorea(L.) Grolle Cephaloyiellarubella(Nees)Warnst. LEPIDOZJACEAE Scapaniaundulata(L.)Dumort. Nomlliacurvifolia(Dicks.)Mitt. Bayyaniatrilobata(L.)A. Gray TRICHOCOLEACEAE Kuryiapauciphra(Dicks.) Grolle CONOCEPHALACEAE Kuryiasetacea(Weber) Grolle Trichocokatomentella(Ehrh.)Dumort. Conocephalumconicum(L.)Dumort. Kuryiasylvatica(A.Evans) Grolle Fepidoyiareptans(L.)Dumort. (Cont.onpage6) Page 6 Long Island Botanical Society Vol. 20 No. 1 BRYOPHYTA BrachythedumrimlareSchimp. DicranumfulvumHook. AMBLYSTEGIACEAE Brachythedumrutabulum(Hedw.)B.S.G. DicranummontanumHedw. Amblystegiumserpens(Hedw) Schimp. Brachythedumsalebrosum DicranumontariensePeters. Amblystegiumserpens\T2£.juratykanum (F.Weber&D.Mohr) Schimp. DicranumpolysetumSw. (Schimp.)Rau&Herv. Brachythedumvelutinum(Hedw) Schimp. DicranumscopariumHedw Amblystegiumvarium(Hedw)Lindb. Bryhnianovae-angliae(Sull.&Lesq.) Grout DicranumspuriumHedw Calliergoncordifolium(Hedw)Kindb. Bryoandersoniaillecebra(Hedw.)H.Rob. DicranumundulatumBrid. Calliergongiganteum(Schimp.) Kindb. Hurlynchiumhians(Hedw.) SandeLac. Rhabdomisiacrispata(With.)lindb. Calliergonstramineum(Brid.) Kindb. Calliergonellacuspidata(Hedw) Loeske Homalothedellasubcapillata(Hedw.) Broth. DITRICHACEAE Campyliumhispidulum(Brid.)Mitt. Platyhypnidiumriparioides(Hedw.)Dix. Ceratodonpurpureus(Hedw)Brid. Campyliumradicale(P.Beauv.) Grout Pseudoscleropodiumpurum(Hedw) Fleisch. Ditrichumpallidum(Hedw.)Hampe Campyliumstedatum(Hedw) GE.O.Jensen Ditrichumpusillum(Hedw.)Hampe Drepanocladusaduncus(Hedw.)Warnst. (Hedw)Jaeg.&Sauerb. Hygroamblystegiumflmiatile(Hedw.)Loeske (Bruch&Schimp.) Bruch&Schimp. Ilygroamblystegiumtenax (Hedw.)Jenn. BRYACEAE Pleuridium subulatum(Hedw)Rabenh. Hygrohypnumeugyrium(Schimp.) Loeske Hygrohypnumochraceum BryumcaespitidumHedw. ENTODONTACEAE (TurnerexWilson) Loeske BryumcapillareHedw. lintodoncladorrhiyans(Hedw.)Mull.Hal. Feptodictyumhurnile(P.Beauv.) Ochyra BryumflacddumBrid. lintodonseductrix(Hedw)Mull.Hal. Feptodictyumriparium(Hedw)Warnst. Platylomellakscurii(Sull.)Andrews (Hedw) G.Gaertn.,B.Mey.&Scherb. EPHEMERACEAE Scorpidiumscorpioides(Hedw.) Iimpr. Feptobryumpyriforme(Hedw.)Wilson Ephemerumcrassinermm(Schwagr.)Hampe Warnstorfiaexannulata(Schimp.) Loeske Pohliaannotina(Hedw)Lindb. Ephemerumserratum(Hedw)Hampe Warnstorfiafluitans(Hedw)Loeske Pohlialescuriana(Sull.) Grout EphemerumspinulosumBruch&Schimp. Pohlianutans (Hedw)Lindb. ANDREAEACEAE FABRONIACEAE AndreaearothiiF.Weber&D.Mohr (F.Weber&D.Mohr)Andrews Anacamptodonsplachnoides RJjodobryumroseum(Hedw)Limpr. (Froel.exBrid.)Brid. ANOMODONTACEAE Anomodonattenuatus(Hedw)Huebener BRUCHIACEAE FISSIDENTACEAE Anomodonrostratus(Hedw.) Schimp. Bruchiaflexuosa(Sw.exSchwagr.)Mull.Hal. FissidensadianthoidesHedw Anomodonrugelii(Mull.Hal.)Keissl. Trematodonambiguus(Hedw)Hornsch. FissidensbryoidesHedw. Haplohymeniumtriste(Ces.) Kindb. Fissidensbushii BUXBAUMIACEAE (Cardot&Ther.) Cardot&Ther. AULACOMNIACEAE BuxbaumiaaphyllaHedw. Fissidensfontanus(B.Pyl.) Steud. Aulacomniumheterostichum Diphysdumfoliosum(Hedw.)D.Mohr FissidensosmundioidesHedw. (Hedw)Bruch&Schimp. FissidenstaxifoliusHedw. Aulacomniumpalustre(Hedw) Schwagr. CALYMPERACEAE SyrrhopodonincompletusSchwagr. FONTINALACEAE BARTRAMIACEAE SyrrhopodontexanusSull. Dichelymacapillaceum(With.)Myrin BartramiapomiformisHedw. Dichelymapallescens Schimp. Philonotisfontana(Hedw.)Brid. CLIMACIACEAE FontinalisantipyreticaHedw. Plagiopusoederiana ClimadumamericanumBrid. FontinalisdalecarlicaSchimp. (Sw)H.A.Crum&L.E.Anderson Climadumdendroides Fontinalisnovae-angliaeSull. (Hedw)F.Weber&D.Mohr Fontinalissullivantiilindb. BRACHYTHECIACEAE Brachythedumacuminatum(Hedw)Austin (Renauld&Cardot) Grout FUNARIACEAE Brachythedumacutum(Mitt.) Sull. FunariahygrometricaHedw.var. hygrometrica BrachythedumcalcareumKindb. DICRANACEAE Physcomitriumpyriforme(Hedw.)Hampe Brachythedumcampestre(Mull.Hal.) Schimp. Dicranellaceriiculata(Hedw) Schimp. Brachythedumoxycladon Dicranellaheteromalla(Hedw) Schimp. GR1MM1ACEAE & (Brid.)Jaeg. Sauerb.var. oxycladon Dicranellarufescens(With.) Schimp. GrimmiapiliferaP.Beauv. Brachythedumplumosum(Hedw.) Schimp. DicranumcondensatumHedw. Racomitriumadculare(Hedw)Brid. Brachythedumpopuleum(Hedw) Schimp. DicranumflagellareHedw Schislidiumapocarpum(Hedw)Bruch&Schimp. Long Island Botanical Society Vol. 20 No. 1 Page 7 HEDWIGIACEAE NECKERACEAE SPHAGNACEAE Hedndgiadliata(Hedw) P.Beauv. NeckerapennataHedw SphagnumaffineRenauld&Cardot HELODIACEAE ORTHOTRICHACEAE Sphagnumangustfolium(C.E.O.Jensenex Helodiumpaludosum(Sull.)Austin OrthotrichumcupulatumBrid. Russow) C.E.O.Jensen & OrthotrichumohioenseSull. Lesq. SphagnumaustiniiSull. HYLOCOMIACEAE OrthotrichumpumilumSw SphagnumcompactumDC. Hylocomiastrumumbratum(Hedw)Fleisch. OrthotrichumpusillumMitt. SphagnumcuspidatumEhrh.exHoffm. Ilylocomiumspkndens(Hedw.) Schimp. OrthotrichumstellatumBrid. Sphagnumfallax(Klinggr.)Klinggr. Tkuroyiumschreberi(Brid.)Mitt. OrthotrichumstrangulatumP.Beauv. Biotacrispa(Hedw) Brid. Sphagnumflancomans(Cardot)Warnst. HYPNACEAE Biotahutchinsiae(Sm.)Hammar SphagnumfkxuosumDozy&Molk. Callicladiumhaldanianum(Grev.)H.A. Crum Sphagnumfuscum(Schimp.)Klinggr. Ctenidiummolluscum(Hedw.)Mitt. PLAGIOIHECIACEAE SphagnumgirgensohniiRussow Heryogiellastriatella(Brid.) Z.Iwats. Plagiotheciumcavifolium(Brid.) Z.Iwats. SphagnumhenryenseWarnst. Heryogiellaturfacea(Lindb.) Z.Iwats. Plagiothedumdenticulatum(Hedw.) Schimp. SphagnumlescuriiSull. I\omomalliumadnatum(Hedw) Broth. PlagiothedumlatebricolaSchimp. SphagnummacrophyllumBrid. HypnumcupressiformeHedw SphagnummagellanicumBrid. HypnumcurvfoliumHedw. POLYTRICHACEAE SphagnummolkSull. HypnumfertileSendtn. Atrichumangustatum(Brid.)Bruch&Schimp. SphagnumpalustreL. HypnumimponensHedw Atrichumcrispum(James) Sull. Sphagnumpapillosumlindb. HypnumlindbergiiMitt. Atrichumundulatum(Hedw.)P.Beauv. Sphagnumplatyphyllum Hypnumpallescens(Hedw)P.Beauv. PolytrichumcommuneHedw. (Lindb.exBraithw) Sull.exWarnst. Hypnumpratense(Rabenh.) KochexSpruce PolytrichumformosumHedw SphagnumportoricenseHampe & Isopterygiumekgans(Brid.)Lindb. PolytrichumohioenseRenauld Cardot SphagnumrecurmmP.Beauv. Isopterygiumtenerum(Sw)Mitt. PolytrichumjuniperinumHedw SphagnumrubellumWilson Tlatydictyasubtilis(Hedw)H.A.Crum PolytrichumpiliferumHedw SphagnumrussowiiWarnst. Platygyriumrepens(Brid.)B.S.G. Pogonatumbrachyphyllum(Michx.)P.Beauv. SphagnumsquamosumCrome Ptiliumcrista-castrensis(Hedw.)DeNot. Pogonatumpensilvanicum(Hedw)P.Beauv. SphagnumstrictumSull. Taxiphyllumdeplanatum SphagnumsubsecundumNees (Bruch&Schimp.exSull.)Fleisch. POTTIACEAE Sphagnumsubtik(Russow)Warnst. Astomummuhlenbergianum(Sw) Grout SphagnumtenerumSull.&Lesq. LESKEACEAE BarbulaunguiculataHedw Sphagnumteres(Schimp.)Angstr. Bryohaplocladiummicrophyllum Desmatodonobtusifolius(Schwagr.) Schimp. (Hedw)R.Watan.&Z.Iwats. Hymenostyliumrecurmrostre(Hedw)Dix. SphagnumtrinitenseMull.Hal. Bryohaplocladiumlirginianum PhascumcuspidatumHedw (Brid.) R.Watan.&Z.Iwats. Pottiatruncata(Hedw.) Fiirnr. TETRAPHIDACEAE BeskeagracilescensHedw Tortellafragilis(Hook.&Wilson)Iimpr. TetraphispelluddaHedw LeskeaobscuraHedw Tortellahumilis(Hedw.)Jenn. BeskeapolycarpaHedw BortulamucronifoliaSchwagr. 1HELIACEAE Bortulapagorum(Milde)DeNot. ThedaasprellaSull. LEUCOBRYACEAE BortulapapillosaWilson Theliahirtella(Hedw) Sull. Beucobryumglaucum(Hedw)Angstr. WeissiacontroversaHedw ThedakscuriiSull. LEUCODONTACEAE PTYCHOMITRIACEAE T1IUTD1ACEAE Beucodonjulaceus(Hedw.) Sull. Ptychomitriumincurvum(Schwagr.) Spruce Haphcladiummicrophyllum(Hedw)Broth. Helodiumpaludulosum(Sull.)Austin MNIACEAE SEMATOPHYLLACEAE Rauiellasdta(P.Beauv.) Reim. MniumhornumHedw Brotherellarecurvans(Michx.) Fleisch. Thuidiumdelicatulum(Hedw.) Schimp. Tlagiomniumdliare(Mull.Hal.)T.Kop. Sematophyllumadnatum(Michx.) E.Britton Thuidiumrecognitum(Hedw.) lindb. Tlagiomniumcuspidatum(Hedw)T.Kop. Tlagiomniumellipticum(Brid.)T.Kop. T1MM1ACEAE Rhfomniumpunctatum(Hedw)T.Kop. LongIslandBotanicalSociety POBox507 Aquebogue,NY11931 Page 8 Long Island Botanical Society Vol. 20 No.1 Upcoming Programs April 13, 2010* Tuesday, 7:30 PM Meg McGrath: “Determining the Impact January and February: No meeting! on Plants of Ground-level Ozone on Long Island.” Dr. McGrath will talk about her March 9, 2010* Tuesday, 7:30 PM work investigating the impact of ambient ozone on Bill Titus: “Biology Today - Beyond agricultural plants growing on Long Island. Each Darwin” This talk will be about biology and summer this ozone reaches concentrations which are high enough to require the issuance of health evolution with a botanical slant. Bill is a retired Coordinator of Cornell Cooperative Extension in advisories for people. Meg is an Associate Professor Nassau County. He has an interest in the natural with a research/extension appointment in the Department of Plant Pathology and Plant-Microbe sciences and, specifically, evolution and its evidence in ourworld today. He recently attended a shipboard/ Biology at Cornell University. She is stationed at the Long Island Horticultural Research and Extension cruise course entitled “Evolution Emanations” Center where she has been working since 1988 which consisted of about twenty-four hours of on optimizing management of diseases affecting presentations by researchers working on evolution. Location: Bill Paterson Nature Center, vegetable cropswithin organicaswell asconventional production systems, and also on determining impact Muttontown Preserve, EastNorwich ofambient ozone on plant productivity. Location: Museum ofLong IslandNatural Sciences, Earth andSpace Science Building, GilHanson Room (Room 123), SUNYat StonyBrook, StonyBrook * Refreshments and informal talk begin at 7:30 p.m. Formal meeting starts at 8:00 p.m. Directionsto MuttontownorStonyBrook: 516-354-6506

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