Key To Species Of Omadius (Coleoptera: Cleridae) From Manado, Sulawesi

3 Pages·1996·1 MB·English

by  J R Mawdsley

Vol. 107, No. 5,November& December, 1996 329
KEY TO SPECIES OF OMADIUS (COLEOPTERA:
CLERIDAE) FROM MANADO, SULAWESI (INDONESIA)
WITH NEW RECORDS AND SYSTEMATIC NOTES
1
Jonathan R. Mawdsley^
ABSTRACT: A key is providedtothe six species ofOmadiusrecorded from Manado, Sulawesi
(Indonesia): O. bivulneratus; O. fasciipes (NEW RECORD); O. indicus (NEW RECORD);
O. prioceroides; O. mediofasciatus', and O. radulifer. Lectotypes are designated for Omadius
bivulneratusand Omadiusprioceroides. Omadiusfemoralis and O. posticalisare placed in syn-
onymywithO. mediofasciatus(NEWSYNONYMIES).
Four species ofthe checkered beetle genus Omadius were described from
specimens collected by Alfred Russel Wallace in the vicinity of Manado,
Sulawesi, during Wallace's celebrated voyage through the islands of present-
dayIndonesia: O. bivulneratusThomson; O.prioceroidesThomson;O.femoralis
Gorham (a synonym of O. mediofasciatus Westwood; see below); and O.
radulifer Gorham. During a recent visit to the Museum National d'Histoire
Naturelle (Paris), Iexamined the holotype orsyntype series ofall ofthese spe-
cies as a part of my ongoing systematic investigations of this genus. These
investigations have also enabled me to synonymize O.femoralis Gorham and
O.posticalisGorham(describedfromthePhilippines)underO. mediofasciatus
Westwood.
Tropical Indonesiahas been greatly altered sinceWallace'svisit, and there-
fore Wallace's collection of Omadius species serves as a useful benchmark
against which more recentcollections ofthese beetles may be compared. Col-
lections made during 1986 by P. M. Hammond (1990) ofthe Natural History
Museum (London) in the Dumoga-Bone National Park near Manado captured
onlyoneofWallace's species(O.prioceroidesThomson),aswell asthecrypti-
cally colored and more inconspicuous O. fasciipes Westwood and O. indicus
LaportedeCastelnau, which were probably overlooked by Wallace. These two
species are therefore reported here as NEW RECORDS. I have prepared the
following key to species of Omadius known to occur on the northern arm of
Sulawesitoaid futureworkersin identifyingtheselargeandconspicuousmem-
bers ofthe forest clerid fauna.
1 ReceivedNovember 14, 1995.AcceptedMarch 15, 1996.
2DepartmentofEntomology,ComstockHall,Cornell University,Ithaca, NY 14853,USA.
ENT. NEWS 107(5)329-331, November& December, 1996
330 ENTOMOLOGICALNEWS
GenusOmadius Laporte deCastelnau (1836)
Type Species: Omadiusindicus LaportedeCastelnau (1836).
(Subsequentdesignation by Desmarest (1860).
Diagnosis: Body elongate, nearly cylindrical; overall length 10-19 mm; antennae filiform near
base, stronglyserratetowardsapices;eyesfinelygranulateand very large, fronscorrespondingly
reduced;terminalmaxillarypalpomeressubulate;terminallabialpalpomeresbroadenedatapices;
postgenalprocessesstronglyacuminateandmeetingmediallyabovegula;elyuralhumerielevated;
femoraand tibiae elongate, slender; tarsal pads present on protarsomeres 1-4. mesotarsomere 4,
andmetatarsomere4;pretarsalclawsappendiculate.
Key tospecies ofOmadius recorded from Manado, Sulawesi
1.) Elytralbaseswithoutrowsofsmalltubercles 2
Elytralbaseswithrowsofsmalltubercles 3
2.) Integumentnearlyglabrousandstronglyshining;legsreddish-orange,withoutblackmaculae;
elytrablackwithredbasesandwhitetransversefasciae 0.priocewidesThomson(1860)
Integumentdenselypubescent,dull;legsyellowwithblackmaculae;elytrabrownwithtrans-
versepiceousfasciae O.fasciipesWestwood(1855)
3.) Pronotalarchtransverselystrigose,atleastinpart 4
Pronotalarchnottransverselystrigose,usuallyminutelyandsparselypunctate 5
4.) Elytra nearly glabrous except forwhite setae along base ofsuture and lateral ovate orange
patchesofpubescence;integumentalcolorationpredominantlymetallicblueblack;metatibiae
planate O. bivulneratusThomson(1860)
Elytradenselypubescent,patterndistinctive:groundcolorgreyish-brownwithabroadtrans-
verseblackbandandoneortwosmallblackpatchesatbasalandapicalthird;metatibiaemore
cylindrical O.mediofasciatusWestwood(1852)
5.) Elytra yellowish brown with dark brown fasciae; elytra tapering uniformly from humeri to
apices;elytralapicessharplyacuminate O. indicusLaportedeCastelnau(1836)
Elytrablack with white fasciae;elytraparallel-sided with apicesseparately rounded; apices
broadlyrounded O. raduliferGorham(1876)
SPECIMENS EXAMINED
Iexaminedholotypeorsyntypematerialofallspeciesdescribedorrecorded
fromManado,aslistedbelow; allofthesespecimensarecontainedeitherinthe
collectionsoftheHopeEntomologicalDepartment,OxfordUniversity(HCOX),
orinthecollectionoftheMuseumNationald'HistoireNaturelle,Paris(MNHN).
O. bivulneratusThomson-2syntypes,Manado(MNHN),includingLECTOTYPEmale
(here designated).
O. fasciipesWestwood - Holotype, Singapore (HCOX).
O.femoralis Gorham - Holotype, Manado (MNHN).
O. indicus Laporte de Castelnau - 5 syntypes from India (MNHN). I do not plan to
designatealectotypeforthisvariablespeciesuntil thepatternsofvariationwithinthecom-
plex "indicusgroup"are betterunderstood,andtherelationshipsofall speciesinthisgroup
can beevaluated simultaneously.
O. mediofasciatus Westwood - Holotype, Khasyah Hills, India(HCOX).
O.prioceroidesThomson-2syntypes, Manado(MNHN),includingLECTOTYPEmale
(here designated).
O. raduliferGorham - Holotype, Manado (MNHN).
Vol. 107, No. 5,November&December, 1996 331
Ialsoexamined five individualseachofO. prioceroides, O. indicus, and O.
fasciipescollected by P. M. Hammond in the Dumoga-Bone National Forestin
1986 (MNHN).
SYSTEMATIC NOTES
I examined the unique holotypes ofO.femoralis Gorham (1876:104) from
Manado and O. posticalis Gorham (1876:105) from "Philippines" (both in the
H. S. Gorham Collection, Museum National d'Histoire Naturelle, Paris) and
concluded that both are conspecific with O. mediofasciatusWestwood (1852),
aspecieswhoseholotypeIexamined in 1990(HopeEntomological Collection,
Oxford University). Gorham used characters of size and coloration to distin-
guish these two species from O. mediofasciatus; these characters vary consid-
erably in a series of 320 specimens of O. mediofasciatus from Laos which I
examined in the Bernice P. Bishop Museum, and undoubtedly only a single
widespreadspeciesisinvolved.Thefollowingnewsynonymy isthuspresented:
Omadius mediofasciatusWestwood, 1852.
O.femoralisGorham, 1876. NEW SYNONYMY.
O. posticalis Gorham, 1876. NEW SYNONYMY.
ACKNOWLEDGMENTS
I thank J. J. Menier(Museum National D'Histoire Naturelle, Paris) for his many courtesies
duringmyvisitin 1993,includingpermissiontoexaminespecimenscollectedbyP.M. Hammond
inDumoga-BoneNationalForest.G.A.Samuelson(BerniceP. BishopMuseum)andG. McGavin
(OxfordUniversity)bothprovidedmuchcourteousassistanceduringmyvisitstotheirrespective
collections.FundingfortravelassociatedwiththisprojectwasprovidedbytheBernice P. Bishop
Museum, two Ford Foundation Undergraduate Research Grants, an Emst Mayr Grant from the
MuseumofComparativeZoology,aNationalScienceFoundationGraduateResearchFellowship,
andaGraduateFellowshipfromtheOlinFoundation.J. K. LiebherrandE. R. Hoebekeprovided
usefulcommentsonthemanuscript.
LITERATURECITED
Desmarest, H. 1860. Cleridae. pp. 251-279. In Chenu, D., Coleopteres, deuxieme partie.
EncyclopedicD'HistoireNaturelle. Maresqcetco., Paris.
Gorham,H.S. 1876. NotesontheColeopterousFamilyCleridae.Cist.Entomol.2:57-106.
Hammond, P. M. 1990. Insectabundanceand diversity in the Dumoga-Bone National Park, N.
Sulawesi,withspecialreferencetothebeetlefaunaoflowlandrainforestintheTorautregion.
pp. 197-254. In Knight,W.J.and Holloway, J. D. (eds.) InsectsandtheRainforestsofSouth-
eastAsia.TheRoyalEntomologicalSocietyofLondon,London.
Laporte,F.L.de,ComtedeCastelnau. 1836.EtudesEntomologiques,oudescriptionsd'insectes
nouveauxetobservationssurlasynonymic. Rev.Entomol.4:5-60.
Thomson,J. 1860. Matenauxpourservieraunemonographicnouvelledelafamilledesderides.
MuseeScientifique 1:46-67.
Westwood,J.O. 1852.DescriptionsofnewspeciesofCleridae,fromAsia,Africa,andAustralia.
Proc.Zool. Soc.Lond. 1852:34-55+pis.24-27.
Westwood,J.O. 1855. Descriptionsofsome newspeciesofCleridae,collectedat Singapore by
Mr.Wallace. Proc.Zool. Soc.Lond. 1855:19-26+pi. 38.