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Introductory botany : plants, people, and the environment PDF

649 Pages·2008·81.31 MB·English
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S E C O N D E D I T I O N I N T R O D U C T O R Y B O TA N Y P L A N T S , P E O P L E , A N D T H E E N V I R O N M E N T LINDA R. BERG, Ph.D. Former Affiliations: University of Maryland, College Park St. Petersburg College Australia • Brazil • Canada • Mexico • Singapore • Spain • United Kingdom • United States 0000--WW44111166--FFMM..iinndddd ii 22//2211//0077 88::4455::0022 AAMM Introductory Botany: Plants, People, and the Environment, Second Edition Linda R. Berg Acquisitions Editor: Peter Adams Permissions Editor: Bob Kauser Development Editor: Kari Hopperstead Production Service: Newgen–A ustin Assistant Editor: Lauren Oliveira Text Designer: Lisa Buckley Editorial Assistant: Rose Barlow Photo Researcher: Abigail Baxter Technology Project Manager: Keli Amann Copy Editor: Cynthia Lindlof Marketing Manager: Kara Kindstrom Cover Designer: Bill Stanton Marketing Communications Manager: Bryan Vann Cover Image: Tim Gainey/Alamy Project Manager, Editorial Production: Belinda Krohmer Back Cover Image: DK. Khattiya/Alamy Creative Director: Rob Hugel Cover Printer: Courier Kendallville Art Director: John Walker Compositor: Newgen–A ustin Print Buyer: Doreen Suruki Printer: Courier Kendallville © 2008, 1997. Thomson Brooks/Cole, a part of The Thomson Higher Education Thomson Corporation. Thomson, the Star logo, and 10 Davis Drive Brooks/Cole are trademarks used herein under license. Belmont, CA 94002-3098 USA ALL RIGHTS RESERVED. No part of this work covered by the copyright hereon may be reproduced or used in any form or by any means—graphic, electronic, or mechanical, including photocopying, recording, taping, web distribution, information storage and retrieval systems, or in any other manner—without the written permission of the publisher. Printed in the United States of America 1 2 3 4 5 6 7 11 10 09 08 07 For more information about our products, contact us at: Thomson Learning Academic Resource Center 1-800-423-0563 For permission to use material from this text or product, submit a request online at http://www.thomsonrights.com. Any additional questions about permissions can be submitted by e-mail to [email protected]. ExamView® and ExamView Pro® are registered trademarks of FSCreations, Inc. Windows is a registered trademark of the Microsoft Corporation used herein under license. Macintosh and Power Macintosh are registered trademarks of Apple Computer, Inc. Used herein under license. Library of Congress Control Number: 2006939406 ISBN-13: 978-0-534-46669-5 ISBN-10: 0-534-46669-9 0000--WW44111166--FFMM..iinndddd iiii 22//2211//0077 88::4455::0044 AAMM This book is dedicated to botany professors and students everywhere. Crescat scientia vita excolatur. 0000--WW44111166--FFMM..iinndddd iiiiii 22//2211//0077 88::4455::0055 AAMM This page intentionally left blank Brief Contents Chapter 1 An Introduction to the Science Chapter 14 The Molecular Basis of Botany 2 of Inheritance 274 Chapter 15 Genetic Frontiers 292 Part 1 The Plant Cell Chapter 16 Continuity through Evolution 310 Chapter 2 The Chemical Composition of Cells 22 Chapter 17 The Evolution of Populations and Species 328 Chapter 3 Plant Cells 44 Chapter 4 Metabolism in Cells 64 Part 4 Diversity Chapter 18 The Classifi cation of Plants Part 2 Plant Structures and Life and Other Organisms 348 Processes Chapter 19 Viruses and Prokaryotes 364 Chapter 5 Plant Tissues and the Multicellular Plant Body 90 Chapter 20 Kingdom Protista 382 Chapter 6 Plant Organs: Roots 110 Chapter 21 Kingdom Fungi 404 Chapter 7 Plant Organs: Stems 128 Chapter 22 Introduction to the Plant Kingdom: Bryophytes 428 Chapter 8 Plant Organs: Leaves 150 Chapter 23 The Plant Kingdom: Seedless Chapter 9 Flowers, Fruits, and Seeds 174 Vascular Plants 444 Chapter 10 Mineral Nutrition and Transport Chapter 24 The Plant Kingdom: in Plants 198 Gymnosperms 466 Chapter 11 Growth Responses and Regulation Chapter 25 The Plant Kingdom: Flowering of Growth 220 Plants 484 Part 3 The Continuity of Plant Life Part 5 Plant Ecology Chapter 12 Mitosis, Meiosis, and Life Chapter 26 Ecosystems 514 Cycles 240 Chapter 27 Global Ecology and Human Chapter 13 Patterns of Inheritance 254 Impacts 538 0000--WW44111166--FFMM..iinndddd vv 22//2211//0077 88::4455::0055 AAMM vi • Brief Contents Appendix A Metric Equivalents and Appendix G Broadleaf Evergreens 580 Temperature Conversion 566 Appendix H Deciduous Trees 582 Appendix B Some Important Biological Events in Geologic Time 568 Appendix I Plant Selection for Various Conditions 587 Appendix C USDA Plant Hardiness Zones 569 Appendix J Biological Control of Insects 589 Appendix D Annual Flowers 570 Appendix E Flowering Perennials 574 Glossary 590 Index 604 Appendix F Narrowleaf Evergreens 578 0000--WW44111166--FFMM..iinndddd vvii 22//2211//0077 88::4455::0066 AAMM Contents Preface xxiii Plant DNA transmits information from one generation to the next 12 Plant populations undergo genetic changes over CHAPTER 1 time 12 An Introduction to the Science Biological Diversity 13 of Botany 2 Many biologists assign organisms into six Learning Objectives 2 kingdoms 13 The Human Population and Plants 4 Many biologists also assign organisms into three Plants and People: Taxol 5 domains 15 Plants and People: Spices: A Brief History 6 Classifi cation is hierarchical 15 Each species is named using a binomial system 15 What Is Botany? 6 The Scientific Method 16 Botany comprises many disciplines 7 An experiment must have a control 16 Characteristics of Plants 8 Scientists use inductive and deductive reasoning 16 Plants are highly organized 8 Focus On: Using the Scientific Method 17 Plants take in and use energy 10 Many observations and experiments support Plants respond to stimuli 11 theories 18 Plants grow and develop 11 Practicing science requires honesty 19 Plants reproduce 12 Study Outline 19 Review Questions 20 Photo Researchers, Inc. TCHhAouPTgEhRt Q2uestions 20 Jeffrey Lepore/ TLehaern iCnhg eOmbjeicctaivl eCs o 2m2position of Cells 22 An Introduction to Basic Chemistry 24 Atoms are the smallest particles of elements 24 Each atom has an atomic number and an atomic mass 24 Electrons move around the nucleus, but their exact positions are uncertain 25 Chemical bonds hold atoms together 25 Atoms gain or lose electrons to form ionic bonds 26 Atoms share electrons to form covalent bonds 26 Hydrogen bonds are attractions between adjacent molecules 27 Inorganic Compounds 27 Focus On: How Did Life Begin? 28 Water, an inorganic compound, is essential to plants 28 Water’s polarity causes many of its properties 28 vii 0000--WW44111166--FFMM..iinndddd vviiii 22//2211//0077 88::4455::0077 AAMM viii • Contents Acids and Bases 30 Biological Membranes 56 Acids and bases dissociate when dissolved The fl uid mosaic model describes membrane in water 30 structure 56 A solution’s acidity or alkalinity is expressed Membranes perform many functions 57 in terms of pH 30 Passage of Materials across Biological Membranes 58 Plants and the Environment: Acid Rain 31 Diffusion is the movement of a substance from a Organic Compounds 31 region of higher concentration to a region of lower Plants and People: Phytochemicals and Human concentration 58 Health 32 Osmosis is the diffusion of water across a selectively Sugars, starches, and cellulose are examples permeable membrane 59 of carbohydrates 32 Some substances cross membranes by facilitated Fats and oils are examples of lipids 34 diffusion or active transport 60 Proteins are large organic molecules composed Study Outline 61 of amino acids 36 Enzymes are proteins that accelerate chemical Review Questions 62 reactions 37 Thought Questions 63 DNA and RNA are nucleic acids 38 Energy is temporarily stored in ATP 38 CHAPTER 4 Energy and Biological Work 39 Metabolism in Cells 64 The laws of thermodynamics govern energy transformations 40 Learning Objectives 64 Study Outline 41 Metabolism 66 Oxidation– reduction reactions occur Review Questions 42 in metabolism 66 Thought Questions 43 Electrons associated with hydrogen are transferred to electron acceptor molecules 66 CHAPTER 3 Plant Cells 44 mages Learning Objectives 44 Getty I MetPhloadnst btoio Slotugdisyt sC teolldsa y 4u6se a variety of methods Ward/Life File/ to study cells 46 w Eukaryotic and Prokaryotic Cells 48 Andre The Structure of Plant Cells 48 The plasma membrane is the outer boundary of the cell 48 The nucleus is the cell’s control center 48 Chloroplasts convert light energy to chemical energy 49 Mitochondria convert the chemical energy in food molecules to ATP 51 Focus On: The Evolution of Eukaryotic Cells 52 Ribosomes are sites of protein synthesis 52 The endoplasmic reticulum has many functions 53 The Golgi apparatus is the cell’s collecting and packaging center 53 Vacuoles are large, membrane-bounded sacs 54 The cytoskeleton is composed of protein fi bers 54 Cell walls surround plant cells 55 Plant cells and animal cells are more alike than different 56 0000--WW44111166--FFMM..iinndddd vviiiiii 22//2211//0077 88::4455::0099 AAMM Contents • ix Photosynthesis 66 Light exhibits properties of both waves and particles 66 Plants and the Environment: The Environmental Effects of Using Biomass for Fuel 68 In plants and algae, photosynthesis takes place in chloroplasts 68 Chlorophyll is the main photosynthetic pigment 70 In photosynthesis, plants convert light energy to chemical energy stored in carbohydrate molecules 72 The light-dependent reactions capture energy 72 The carbon fixation reactions produce carbohydrates 72 Photosystems I and II are light-harvesting units of the light-dependent reactions 72 ATP synthesis occurs by chemiosmosis 75 Most plants use the Calvin cycle to fix carbon 76 Photorespiration reduces the effi ciency of the C pathway 76 3 Many plants with tropical origins fi x carbon using the C pathway 78 4 Many desert plants fi x carbon using the CAM pathway 79 Aerobic and Anaerobic Pathways 80 mited Aerobic Respiration 80 Unli Aerobic respiration occurs in four stages: Visuals AeArIPnony bo rigvcul eyvrrceavostilpeey iwsirsias ct, iogo8nln1uv ciesor staee cdios t mcoo panlcveexetr yptler Codc oteoAs s p y 8r8u32v ate 83 © David Cavagnaro/ The citric acid cycle oxidizes acetyl CoA 83 The electron transport system produces most of Collenchyma cells have unevenly thickened the ATP 85 primary cell walls 95 Anaerobic Pathways 85 Sclerenchyma cells have both primary cell walls Plants and the Environment: Electron Transport and thick secondary cell walls 95 and Heat in Plants 86 The vascular tissue system consists of two complex tissues 95 Fermentation is ineffi cient 87 The conducting cells in xylem are tracheids and Study Outline 87 vessel elements 99 Review Questions 89 Sieve-tube elements are the conducting cells of phloem 99 Thought Questions 89 The dermal tissue system consists of two complex tissues 99 CHAPTER 5 Epidermis is the outermost layer of an herbaceous Plant Tissues and the Multicellular plant 99 Plants and People: Fibers and Textiles 100 Plant Body 90 Periderm replaces epidermis in woody plants 102 Learning Objectives 90 Plant Meristems 105 Plant Structure and Life Span 92 Primary growth takes place at apical meristems 105 Plants and the Environment: Plant Life History Secondary growth takes place at lateral Strategies 93 meristems 107 Cells and Tissues of the Plant Body 93 Study Outline 107 The ground tissue system is composed of three simple Review Questions 108 tissues 94 Parenchyma cells have thin primary cell walls 95 Thought Questions 109 0000--WW44111166--FFMM..iinndddd iixx 33//99//0077 1122::1177::5566 PPMM

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