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446 Novon but also were unable to verify the identity of the Variety remotispora differs from typical P. no- other parental taxon. bilis only in its venation pattern, with copious mar¬ ginal anastomoses, rather than the free veins found Paratypes. MEXICO. CHIAPAS: 13 kin N of Berriozabal, in variety nobilis. The two varieties are sympatric 2 Nov. 1971, Breedlove & Smith 2/652(DS, F, MICH, NY), oaxaca: trail from San Pedro Nolasco N to the in much of Mexico, although they are seldom en¬ Llano Verde, 5 Oct. 1969, Alickel & Hellwig 3786B countered in mixed populations. Rare specimens ot (NY). VERACRUZ: ca. 12 kin S of Misantla, 3 Jan. 1974, intermediate morphology (veins with few, irregular Conant 726 (GH, MEXU). anastomoses) originate from Michoaean (Arsene 5961; GH, MO, US) and Veracruz (Fisher s.n Phanerophlebia nobi I is (Schlechtendal & MO), Mexico, and are apparently fully lertile. Chamisso) C. Presl var. remotispora (Four¬ nier) Yatskievych, comb, et stat. nov. Pha¬ Literature Cited nerophlebia remotispora Fournier, Mexic. PI. Maxon, W. R. 1912. Notes on the North American 1: 100, 1872. Cyrtomium remotisporum species of Phanerophlebia. Bull. Torrey Bot. Club (Fournier) C. Morton, Amer. Fern J. 47: 54, 39: 23-28. Mickel, J. T. & J. M. Beitel. 1988. Pteridophyte flora 1957. TYPE: Mexico. Veracruz: Orizaba, of Oaxaca, Mexico. Mem. New York Bot. Card. 46: Izhuatlancillo, Aug. 1805-1866, Bourgeon 1-568. 23 19(holotype, P; isotypes, BM, GH, k, MICH, Tryon, R. M. & A. F. Tryon. 1982. Ferns and Allied MO, NY, I’, US). Plants, With Special Reference to Tropical America. Springer-Verlag, New York. Previous authors (cl. Tryon & Tryon, 1982) uni¬ Underwood, L. M. 1899. American ferns—II. The formly have treated variety remotispora as a distinct genus Phanerophlebia. Bull. Torrey Bot. Club 26: species. These authors emphasized anastomosing 205-215, 2 pi. Yatskievych, G. 1990. Systematics of the Fern Genus versus free venation patterns as a critical character Cyrtomium in the New World. Ph.D. Dissertation, in subdivision of Phanerophlebia into two species Indiana University, Bloomington. groups. Yatskievych et al. (1988) showed that P. - & G. J. Gastony. 1987. Hybridization and n obi Us and P. remotispora are genetically very polyploidy in the fern genus Phanerophlebia. Amer. J. Bot. 74: 715. [Abstract.] closely related, based on analysis of ehloroplast-DNA -, D. B. Stein & G. J. Gastony. 1988. Chloro- restriction-site variation, and suggested that anas¬ plast DNA evolution and systematics of Phanero¬ tomosing veins were probably derived twice inde¬ phlebia and related fern genera. Proc. Natl. Acad. pendently within the genus. U.S.A. 85: 2589-2593.