Important calcareous foraminifera of the type- locality of the genus Quadratobuliminella Quadratob de Klasz, region of Eisenärzt, Upper Bavaria

11 Pages·1993·4.6 MB·English

by  Ivan De Klasz

Zitteliana
312
Germany
Wunich•Salzburghighway
X
Ruhpolding
• /
Austria B«.xhtesgadan
Fig.1a
Hereunder we mention and illustrate the most important
calcareousspecies,includingtheplanktonicones,topotypesof
Quadratobuliminellapyramidalisandformsalsoencountered
forinstancebyvonHillebrandt(1962)inthePaleoceneof
the Bad Reichenhall and Salzburg basins and by diverse
authorsinvariousregionsoftheSouthAtlantic.Forreasonof
space, apart from of the original description, only a few
referencesfromtheseareasaregivenforeachspecies.Amore
detailedbibliographyisincludedforinstanceindeKlaszetal.
(1988).
The figured specimens are deposited in the Bayerische Fig.1b
Staatssammlung für Paläontologie und historische Geologie, Fig.1 aand1 b:RegionalandpreciselocalisationoftheQuadrato-
Munich,Germany. b(uFlluirmkianretlel)a1t:y50p0e0,loschaeleittySO(x)1.6-(3R5epwritohdupcetrimoinssoifonpoafrtthoefBaLyaenrdis-cmhaeps
Landesvermessungsamt,n°7705/89).
ThepreparationofthispaperwasencouragedbyProf.Dr.K.F.
Weidich, who also kindly arranged the SEM photography ofour forustousethelaboratory'sdarkroomtomakepnntsofthenegati-
specimensbeforehisuntimelvdeath.WeareindebtedtotheInstitute ves. Miss Sophie Pagnotta and Mr.Jean Pierre Laugier of the
ofPaleontologyandhistoricalGeologyoftheUniversityofMunich laboratorystaffsecondedusineverypossibleway.D.Schnitkerof
tfoorokentahbeliSngEoMurpiwcoturrkestoanbedckaerprtieidnocuot,ntaancdttwoitMhrsu.sRd.uLriienbgrePirocfhewsshoor Wmaatleproilael,ofMec.o,mpUaSriAs,ona.nWdeJ.arWeidmomsatrkgraotfefuUlpptsoaDlra.kWi.ndWlyersneentruosf
Weidich'sillness. Prof. G. NicaiseDirectorofthe Laboratoryfor the Bayerische Staatssammlung für Paläontologie und historische
AppliedMicroscopytheUniversityofNicekindlymadeitpossible GeologieaswellastoMrs.M.Dallosforrevisingthetext.
THE QUADRATOBULIMINELLA TYPE LOCALITY
2.
2.1REMARKSCONCERNINGTHE 2.2FAUNALAFFINITIESOFTHECALCAREOUS
QUADRATOBULIMINELLATYPELOCALITY FAUNAFROMTHEGRUBLOCALITY
The structural position of the type Stratum has been A considerable part of the calcareous foraminifers of the
discussedamongothersbydeKlasz&deKlasz(1990).They Grub material was also encountered by von Hillebrandt
consideredit,asdiddeKlasz(1953b),asatectonicandnot (1962)intheBadReichenhallandSalzburgbasins,whichare
stratigraphicintercalationintheBucheckSchichten.Therecent only 30 to 40 km away, as well as by various authors in
findingofasampleresiduecollectedbyoneofus(I.K.)in1953 numerousSouthAtlanticlocalitiesandtheCarribeanregion.
about20mSofthe Quadratobuliminellatypelocality(con- The planktonic foraminifers indicate Upper Danian age.
taininga rieh Campanian fauna, with numerous Globotrun- BecauseofthecoexistenceofMorozovellatrinidadensisandM.
canidsof„Buchecktype")confirmsthisStatementdecisively:it uncinata,deKlaszetal.(1990)attributedthemtothelower
proves that Senonian Bucheck Layers exist both North and partofBlow'sP2zone.ThebathymetricInterpretationofthe
SouthoftheQuadratobuliminellabearinglevels.Thesection Grubmaterialandtherelationshipofthelatterwithgeological
figured by de Klasz & de Klasz (1990) has been modified featuresoftheareaisshortlygivenindeKlasz&deKlasz
accordingly(Fig.2). (1990)wherefurtherreferencesconcerningthesearealsoin-
Quadratobuliminella
type level
SI^CSDORF UNT.-SCHARAM EISÜNARZT
Wei«e'Triun
ek f>i
FEiigs.en2ä:rGzetolaongdicSailegssedcotrifonin1d:i2c5at0i0n0gbtehtewteeecntotnhiccspeodsiimteinotnoolfogtihce-tDeacntionaincluenvietlsoffrGormuFblys(cp)h((affgtte)rtoGamnoslsass19e5(6,omm)odiinftiheed)r.egAirornoowf
pointstoBuchecklocalityencountreredtothesouthoftheQuadratobuliminellatypelocality(p).Forabbreviationsseealso
deKlasz&deKlasz,1990:fig.4.
cluded. The composition of the calcareous assemblage con- Morozovellapraecursoria(Morozova,1957)
firms the data obtained from agglutinating forms. Although PI. 1,Fig.6
manyspeciesarealsofoundintheDanianfaunaofSenegal(de 1957 Acarininapraecursorian.sp.-Morozova:1111,fig.1.
Klaszetal. 1988(PI zone)),theirrelativeabundanceisrather 1975 Globorotaliapraecursoria (Morozova). -Stainforth
different. It recalls rather the associations considered by
Nyong & Olsson (1986) as being characteristic of meso- etal.:214-215,fig.74.
bathyalconditions. 1985 Morozovellapraecursoria(Morozova).-Toumarkine
&Luterbacher:110,fig.13/5a-c.
Hagnetal.(1981)mentionthatnohiatuscanbeobserved
between the Late Cretaceous and Early Paleogene in the
Ultrahelvetic zone in South-Eastern Bavaria. In the Grub Morozovellapseudobulloides(Plummer,1926)
localitythepresenceofsomeGlobotruncanidsintheDanian PI. 1,Fig.4a-b
faunashowshoweverthatsomereworkingoccurred. 1926 Globigerina pseudobulloides n. sp. - Plummer: 33,
pl.8/9.
TAXONOMICREFERENCESANDREMARKS 1975 Globorotaliapseudobulloides(Plummer).-Stainforth
2.3 etal.:216-217,fig.76.
Morozovellapseudobulloides(Plummer).-
2.3.1Planktonicforaminifera Toumarkine&Luterbacher:110,fig.14/1a-2c.
ChiloguembelinasubtriangulansBeckmann,1957
PI.1,Fig.1
Morozovellatrinidadensis(Bolli,1957)
1957 Chiloguembelina subtriangularis n. sp. - Beckmann: PI. 1,Fig.5a-c
91,pl.21/5,text-fig.15/39-42.
1957 Globorotaliatrinidadensisn.sp.-Bolli:73,pl.16/19-23.
1975 Globorotalia trinidadensis Bolli. - Stainforthet al.:
GlobigerinatriloculinoidesPlummer,1926 p.235,237,fig.93.
PI. l,Fig.2a-c 1985 Morozovella trinidadensis (Bolli). - Toumarkine &
1926 Globigerinatriloculinoidesn.sp.-Plummer:134,pl.8/ Luterbacher:110,fig.13/3a-4c.
10.
1975 GlobigerinatriloculinoidesPlummer. -Stainforthet Morozovellauncinata(Bolli,1957)
1985 aGll.:ob2i34g,erfiign.a9t2r.iloculinoidesPlummer.-Toumarkine& PI.1,Fig.7a-c
Luterbacher:117,fig.19/1a-2c. 1957 Globorotaliauncinatan.sp.-Bolli:74,pl.17/13-15.
1975 GloborotaliauncinataBolli.-Stainforthetal.:239,
fig.95.
Morozovellainconstans(Subbotina,1953) 1985 Morozovella uncinata (Bolli). - Toumarkine &
PI. 1,Fig.3a-c Luterbacher:110,fig.14/3a-4c.
1953 Globigerinainconstansn.sp.-Subbotina:58,pl.3/1-2.
1975 Globorotaliainconstans(Subbotina). - Stainforthet Planorotalitescompressa(Plummer,1926)
al.:193,fig.55. PI. 1,Fig.8a,b
1985 Morozovellainconstans(Subbotina).-Toumarkine& 1926 Globigerinacompressan.sp.-Plummer:135,pl.8/11.
Luterbacher:110,fig.13/1a-2c. 1975 Globorotaliacompressa(Plummer).-Stainforthetal.:
178,fig.43.
1985 Planorotalitescompressa(Plummer). -Toumarkine&
Luterbacher:107,fig.12/1a-2c.
314
2.3.2Benthicforaminifera Thetaxonomyofthisspecieshasbeendiscussedindetailby
vanMorkhovenetal.(1986).
Anomalinoidesacutus(Plummer,1926)
PL2,fig. 11a-c
Bolivinoidescf.paleocenicus(Brotzen,1948)
1926 Anomalina ammonoides (Reuss) var. acuta n. var. - PI.1,Fig.11a,b
Plummer:149,pl.X/2a-c.
1976 Anomalinoidesacuta(Plummer).-Salajetal.:161,1X7 cf.1948 Bolivinapaleocenican.sp.-Brotzen:66,pl.9/5.
3a-c,6a-b. ? 1962 BolivinoidesdelicatulusCushman.-von
1983 Cibicidoidesdayi(White). -Tjalsma&Lohmann:9, Hillebrandt:71,pl.V/13(non 14).
pl.6/6-7. cf.1970 Bolivinoides paleocenicus (Brotzen). - Barr: 650,
1988 Anomalinoidesacutus(Plummer).-deKlaszetal.:66, pl.99/7a-b.
pl. 17/3a-c. Our specimens are narrower than the specimens of B.
paleocenicus(Brotzen)figuredbyBrotzenandbyBarr.The
sutures show generally a crenulation and most specimens
Aragoniaäff.velascoensis(Cushman,1925) exhibitsomeelevatedlobesonthecentralpartofthetest.
PI. 1,Fig. 12a,b
r1925 Textulariavelascoensisn.sp.-Cushman:18,pl.3/1. Buliminacf.cooperensisCushman,1933
?1988 Aragoniaäff.velascoensis(Cushman).-deKlaszet PI. 1,Fig.15a,b
al.:63,pl.14/12a-c.
cf. 1933 Bulimina cooperensisn. sp. - Cushman: 12,pl. 1/32
Becauseofsubstantial recrystallisationandstronglyadher-
a-b.
pionsgsimbalteriwxitahnceerxtaacitntdye.termination at the species level is not cf. 1947 Bulimina cooperensisCushman. -Cushman&Par-
ker:98,pl.22/19.
cf.1992 BuliminatrinitatensisCushman&Jarvis.-Widmark
BolivinoidesdelicatulusCushman, 1927 &Malmgren:pl.1/7.
PI. I,Fig. 10a-c Ourspecimens are lessslenderthanthe Eocenespecies B.
1927 Bolivinoidesdecorata(Jones)var.delicatula.- cooperensisCushmanandthelamellarornamentationusually
Cushman:90,pl.4/8. reachesthetoppartoftheChambers,evenontheterminalpart
1983 BolivinoidesdelicatulusCushman.-Tjalsma& ofthetest.1tdiffersfromBuliminatrinitatensisinnothavinga
Lohmann:5,pl.4/7. lamellar (and sometimes perforated) lower edge of the
1986 BolivinoidesdelicatulusCushman.-vanMorkhoven Chambersasdescribedbytheauthorsinthelatter.(Seealsothe
etal.:337-339,pl.110/2a-b. SEM photograph fig. 16onpl.2 inProto-Decima&Bolli
1988 Bplo.li1v2i/n9oia-dce.sdelicatulusCushman.-deKlaszetal.:57, t(r1i9n7i8t)atewnshiis)c.hOins vtehreyoctlhoesrehtaondthWeiodrmiagirnkal&draMwailnmggroefn'Bs.
pictureofB.trinitatensisisverysimilartoourform.
Plate1
Fig.
Fig.
Zitteliana,20,1993
Klasz,deI.& deKlasz,S.:ForaminiferaoftheQuadratobuliminellatypelocality Plate1
BuliminataylorensisCushman&Parker,1935 }FrondiculariajarvisiCushman,1939
PI.1,Fig.16 PL1,Fig.9
1935 Buliminataylorensisn. sp. -Cushman& Parker: 96, 1939 Frondiculariajarvisin.sp.-Cushman:91,pl.16/6.
pl. 15/3a-b. 1962 FrondiculariajarvisiCushman.-vonHillebrandt:62,
1962 Bulimina trinitatensis Cushman & Jarvis. - von pl.IV/14.
Hillebrandt:78-79,pl.V/27a-b. 1988 FrondiculariajarvisiCushman. - de Klasz et al.: 46,
1982 Bulimina trinitatensis Cushman & Jarvis. - pl.8/12a-b.
BOLTOVSKOY&WATANABE:pl.2/21. Inouropinion,thefiguredspecimenistheinitial,evenmore
1986 Bulimina trinitatensis Cushman & Jarvis. - van juvenile stage of Frondicularia jarvisi Cushman, than the
Morkhovenetal.,partim:300-303,98B/2;non98A/1 specimenillustratedindeKlaszetal.(1988),pl.8,fig.12b.
a-2c,non98B/l,3,4.
1988 BuliminataylorensisCushman&Parker.-deKlaszet
al.:60,pl.13/5a-c. Gavelinellavelascoensis(Cushman,1925)
PI.2,Fig.14a-c
InastudyontheDanianofSenegal,deKlaszetal.(1988)
discussed the differences between Bulimina taylorensis and 1925 Anomalinavelascoensisn.sp.-Cushman:21,pl.3/
Bulimina trinitatensis. Van Morkhoven et al. (1986) inter- 3a-c.
preted the morphological differences in their figures of B, 1983 Gavelinella velascoensis (Cushman). - Tjalsma &
trinitatensisasbeingduetoevolutionarychange. Lohmann:14,pl.5/8a-b.
1988 Gavelinellarubiginosa(Cushman).-deKlaszetal.:
66,pl.17/2a-c.
Coryphostomaäff.midwayense(Cushman,1936) 1992 Gavelinellavelascoensis(Cushman). -Widmark&
PI.1,Fig.17a,b Malmgren:pl.5/5a-c.
1936 Bolivinamidwayensisn.sp.-Cushman:50,pl.7/12a-b, non 1989 Gavelinellavelascoensis(Cushman). -Boltovskoy
(fidevanMorkhovenetal.1986). &Boltovskoy:305,pl.111/13-14.
1978 Coryphostoma cf. limonense (Cushman). - Proto-
Decima&Bolli:792,pl.2/1.
1983 Coryphostomacf.midwayensis(Cushman). -Tjalsma Gavelinellaäff.sp.AdeKlasz,deKlasz&Ausseil-Badie,
&Lohmann:12,pl.2/7-8. 1988
1985 Bolivina midwayensis (Cushman). - Brun et al.: 25, PI.2,Fig.15a-c
pl.III/7a-f. non1928 Rotaliabeccariiformisn.sp.-White:187,
1988 BolivinamidwayensisCushman.-deKlaszetal.:57, pl.39/2a-c.
pl.12/8a-b. 1982 Gavelinella beccariiformis (White). - Boltovskoy
Intheliterature,severalforaminiferaidentifiedandfiguredas &Watanabe:31,pl.3/60-61.
BolivinalimonensisandCoryphostomalimonenserespectively 1988 Gavelinellasp.A.-deKlasz,deKlasz&Ausseil-
(seereferencesabove)correspondwellwiththeoriginalfigure Badie:67,pl.17a-c.
ofCmidwayense.
Plate2
Zitteliana,20,1993
Klasz,deI.&deKlasz,S.:ForaminiferaoftheQttadratobuliminellatypelocality Place2
318
The specimens of Grub are very similar to the species taxonomyandstratigraphicdistributionofN.ßorealis.Inview
describedinopennomenclarurebyDEK.LASZetal.(1988).For ofthetaxonomicproblemsconnectedwiththelatterasshown
theSenegalformseeremarksinthelatterpaper. byWhiteandvanMorkhovenetal.andtheforms figured
underthis namebyvariousotherauthors,astudyofalarge
numberofthiscosmopolitanindexforminthetypeareawould
Gyroidinoidesglobosus(vonHagenow,1842) bemostuseful.Suchastudywouldmakeitpossibletofindout
PL2,Fig.12a,b whether the specimens observed by some authors in the
1842 Nonioninaglobosan.sp.-vonHagenow:574. Bavarian Alps and the Southern Atlantic fall within the Va-
1962 Gyroidinoidesglobosus(vonHagenow).-von riationspectrumofthespeciesN.ßorealisorrepresentanew
Hillebrandt:107,pl.IX/2a-3c. speciesorsubspecies.
1988 Gyroidinoidesglobosus(vonHagenow).-deKlaszet
al.:66,pl.16/4a-c. PleurostomellapaleocenicaCushman,1947
PI.2,Fig.4
NodosarellakuglenCushman&Renz,1946 1947 Pleurostomellapaleocenican.sp.-Cusiiman:68,pl.18/
PI.2,Fig.2 14-15.
1946 Nodosarellakuglerin.sp.-Cushman&Renz:42,pl.6/ 1962 PleurostomellapaleocenicaCushman.-von
30,33. Hillebrandt:95,pl.VII/27.
1962 Nodosarella kugleri Cushman & Renz. - von 1988 PleurostomellapaleocenicaCushman.-dlKlaszetal.:
Hillebrandt:96,pl.VIII/7,17,18a-b. 61,pl. 13/8a-b.
Nodosarellasubnodosa(Gurpv,1894) Pleurostomellasp.AdeKlasz,
PI.2,Fig.3 deKlasz&Ausseil-Badie1988
1894 Ellipsoidinasubnodosan.sp.-Guppv:640,pl.41/12. PI.2,Fig.5
1962 Nodosarellasubnodosa(Guppy).-vonHillebrandt: 1988 Pleurostomellasp.A.-deKlasz,deKlasz&Ausseil-
97,pl.7/14-16. Badie:61,pl. 14/2a-b.
? 1978 Nodosarella subnodosa Guppy. - Proto-Decima &
Born:795,pl.3/18. PiillemacoryelliWhite,1929
PI.2,Fig.8a,b
Nuttallinellaäff.ßorealis(White,1928) 1929 Pullemacoryellin.sp.-White:56,pl.5/22a-b.
PI.2,Fig.7 1962 PullemacoryelliWhite.-vonHillebrandt:94,pl.6/
? 1928 Gyroidinaßorealisn.sp.-White:293,pl.4/3a-c. 34a-b.
1951 GyroidinaßorealisWhite.-Noth:70-71,pl.7/8a-c. 1988 PullemacoryelliWhite.-deKlaszetal.:64,pl. 15/4
1962 Osangulariaßorealis (White). - von Hillebrandt: a-b
109,pl.IX/13a-14c. Thisspecieshasbeenfiguredinvariouspublicationsonthe
1978 Charltoninaßorealis (White). - Proto-Decima & EarlyandMiddlePaleoceneinCentralandSouthAmerica,as
Bolli:79,pl.4/17-18. well as in the Southern Atlantic and in Europe. Van
1982 Charltonina ßorealis (White). - Boltovskoy & Morkhovenetal.(1986)consideritasacharacteristicelement
Watanabe:31,pl.2/31-32. of„lowerbathyal"(1000-1500m)waterdepths.
? 1986 Nuttallinellaßorealis(White),-vanMorkhovenet
al.:356-358,pl. 115/1-3. Pulleniacf.cretaceaCushman,1936
1988 Charltoninaßorealis(White). - deKlaszetal.: 65, PI.2,Fig.9a,b
pl.16/2a-c. cf.1936 Pulleniacretacean.sp.-Cushman1936(a):75,pl.13/
1992 Nuttallinella ßorealis (White). - Widmark & 8a-b.
Malmgren:pl.2/4a-c. 1983 PulleniaeocenicaCushman&Siegfus. -Tjalsma&
Ourform, like theones observed in approximatelycoeval Lohmann:36,pl.16/1.
W(le1hv9ie8lt6s)eofforroSemtnheetghaeslpGeaucrliefmeloensfsfMihegixugirhceodt.hbaWynevthafenololMroiowgrintkahhleodevrxeaanwmipnlegteaoolff. 1988 PPul.ll1e5n/i5aa-cbr.etacea Cushman. - de Klasz et al.: 64,
various authors in calling this form „Nuttallinella (or Someforaminifers identifiedand figuredasP. eocenica by
Osangularia, orCharltonina)ßorealis",while admitting that various authors (see for instance references above) are very
thisdeterminationisquestionable.Duringthedraftingofour similartoP.cretacea.Onlythestudyoflargepopulationsfrom
paperontheDanianofSenegal(deKlaszetal. 1988)wedid the type locality and from intermediate levels could show
not have van Morkhoven et al.'s book, which gives SEM whether they are different species or varieties of the same
picturesofaformobviouslyidenticalwiththespecimendrawn taxon.
by White; van Morkhoven et al. discuss in detail the
319
}Pyramidinacf.rudita(Cushman&Parker,1936) (18.6.1992)hewrotethathefoundthisforminseveralDSDP
PL 1,Fig. 13 boreholes(sites167-NPacific;sites363,364and525A-South
cf.1936 Buliminaruditan.sp.-Cushman&Parker:45,(fide Atlantic)inlevelsvaryingfromPaleocenetoMiddleEocene.
Cushman&cParker1947). Boltovskoy &Watanabe (1982) figured verytypical Qua-
cf.1947 BPaurlkiemri:na82r-u8d3i,taplC.1u9s/h26m.an&Parker.-Cushman& d-kroMayitdo&dbulBleioEmliotncoeevlnlsea)k.opyWyri(ad1m9mi8da9ar)lkifso&ufnMrdoaimltimDngSsriDteePn52(s51iAt9'e9(23l)6a3te;encPBoaoullnettooecvresen-de
cf.1983 PyramidinaruditaCushman&Parker.-Tjalsma& Quadratobulimmella spp. in DSDP sites 465A (Central
Li»iima\\: 19,pl.4/2,7/2. Pacific),525Aand527(SouthAtlantic)inbothMaastrichtian
1988 Pyramidina cf. rudita Cushman & Parker. - m andDaniandeposits(withapronouncedincreaseintheDanian
K.LASZetal.:60,pl.13/4a-b. samples at site 465A). Specimens from the above three sites
1992 PWylrDaMmAiRdiKn&acMf.ArLuMdGitRaENC:usplh.m1a/n12.&Parker.- ftkooinprdomltsyywpsheesincthotfwoeQuusoabdbtryaaiJtn.oebWduifldirmmoaimrnDekl.,lSaUcphppnysriaatlmaki,edraalroiefs,Widaelanpstoilcaaerle,wMaielt.sh,o
Althoughsubtriangularinsectionthisformdoesnotshow USA,fromtheN.Atlanticsite401(GulfofBiscay).Widmark
the strong overlapping of the last few Chambers, nor the & Malmgren consider Quadratobuliminella as an infaunal
apertural characteristics of Pyramidina, as described by genus.
Loeblich&Tappan(1988).
Qiiadrimorpbinatrocboides(Reuss,1846)
Pyramidinaäff.triangularisCushman&Parker,1935 PI.2,Fig. 10a,b
PI. 1,Fig.14a-c 1846 Globigerinatrocboidesm.-Reuss:36,pl.12/22.
1935 Buliminatriangularisn.sp.-Cushman&Parker:97, 1962 Allomorphina conica Cushman & Todd. - von
pl. 15/4a-b. Hillebrandt:90,pl.VI/21a,b.
1947 B&ulPiamriknear:tr8i2a,ngplu.la1r9i/s2C5UaS-bH.MAN&Parker.-CUSHMAN 1988 6Q3u,apdlr.i1m5o/1rpah-ib.natrocboides(Reuss).-deKlaszetal.:
Thestrongrecrystallisationofthe Grub specimensdo not 1989 AllomorphinaconicaCushman&Todd.-
allowamoreprecisedetermination. Boltovskoy&Boltovskoy:299,pl.V/l-2.
QuadratobuliminellapyramidalisdeKlasz,1953 Stensioeinabeccariiformis(White,1928)
PI.2,Fig.1a-f PI.2,Fig. 13a-c
Type reference: I.deKlasz(1953): Quadratobuliminella 1928 Rotaliabeccariiformisn.sp.-White:27,pl.39/2a-4c.
n.gen.,eineneueForaminiferengattungvonderWendeKrei- 1962 Gavelinellabeccariiformis(White).-von
de-Tertiär. - N. Jb. Geol. Paläont., ML, 10: 434-436, fig. 1 Hillebrandt:101,pl.VIII/2a-c.
(holotype),fig.2(paratvpe),(x76). 1978 Gavelinella beccariiformis (White). - Proto-Decima
Type specimen: holotype:Prot.321;paratvpe:Prot.322, &Bolli:793,pl.6/3-4.
depositedintheBayerischeStaatssammlungfürPaläontologie 1986 Stensioinabeccariiformis(White). -vanMorkhoven
undhistorischeGeologie,Munich,Germany. etal.:346-353,pl. 113A-D.
Type level: DarkgreyishbrownshalesofDanianage(P2 1992 Gavelinella beccariiformis (White). - Widmark &
fzooontep)atthecWtoniocfaltlhye irnatielrwcaaylabterdidgienoBfuWciheescekn"B,edNsE„iaselnoänrgzt,a Malmgren:pl.5/3a-c.
southofthehamletofGrub,50mNofthecrossingofrailway Thetaxonomic position and distribution ofthis species is
(Fig. 1b). dsipseccuismseendsianredsetoamilewbhyatvlaenssMthoirckkhthoavnetnhosetefail.gu(r1e9d86b)y.tOheusre
Paratvpe: FromtheDanianofBidart,SofBiarritz,Pyre- authorsorbyWidmark&Malmgren(1992).
neesAtlantiques,SWFrance.
Stratigraphicränge:EarlyPaleocene(PI)toMiddleEoce-
neI(nPIad1d)i(tfiiodnevtoanthMeotyrpkehloocvaelintyetanald.t1h9e86D:a3n2i2a).nofBidart,SW deStKillaosstzom&elAluassspe.iAl-BdaedKiel,as1z9,88
PI.2,Fig.6
France, Quadratobuliminella has been recorded in widely
dispersedareas(vanMorkhovenetal. 1986,de Klaszetal. 1983 Stilostomellasp.-Tjalsma&Lohmann:5,pl.5/2.
1990).Boltovskoy(1989)mentioneditatDSDPsite305from 1988 Stilostomellasp.A. - de Klasz,deKlasz&Ausseil-
the Late Oligocene. However in a personal communication Badie:59,pl. 12/17.
CONCLUSIONS
3.
Thispapercompletesrecentstudies(deKlaszetal.1990,de The calcareous foraminiferal assemblage shows numerous
Klasz&DEKlasz1990)ontheonlyexistingmaterialfromthe commonformswithvariousEarlyPaleogenefaunasdescribed
nolongeraccessibletypelocalityofthecosmopolitanmainly fromtheBad Reichenhall and theSalzburgBasin,theSouth
EarlyPaleogenedeepwaterbenthicforaminiferalgenusQua- Atlanticandthecircum-Caribbeanregion,asdoestheagglu-
dratobuliminelladeKlasz,1953. tinatedfaunaoftheGrublevels.
Theplanktonicforaminiferausedforagedeterminationare ThefindingofasampleresidueofBucheckLayerscollected
illustratedinthispaper,asaretheprincipalcalcareousbenthic nearbysouthoftheQuadratobuliminellatypelocalitycorro-
species(aftertheagglutinatedformswerepreviouslyfiguredin boratesearlierobservations,i.e.thatthedepositsrepresenta
deKlasz&deKlasz1990).TheyindicateaUpperDanianage tectonic and not stratigraphic intercalation in the Bucheck
(lowerpartofBlow'sP2zone). Layers.
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