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Springer Biographies Imperial Biologists The Imperial Family of Japan and Their Contributions to Biological Research HIDEO MOHRI Springer Biographies The books published in the Springer Biographies tell of the life and work of scholars,innovators,andpioneersinallfields oflearningandthroughout theages. Prominent scientists and philosophers will feature, but so too will lesser known personalitieswhosesignificantcontributionsdeservegreaterrecognitionandwhose remarkable life stories will stir and motivate readers. Authored by historians and other academic writers, the volumes describe and analyse the main achievements of their subjects in manner accessible to nonspecialists, interweaving these with salient aspects of the protagonists’ personal lives. Autobiographies and memoirs also fall into the scope of the series. More information about this series at http://www.springer.com/series/13617 Hideo Mohri Imperial Biologists The Imperial Family of Japan and Their Contributions to Biological Research 123 HideoMohri University of Tokyo Bunkyo-ku,Tokyo,Japan Translated by Editorial advisor Yoko Kawazoe NaotoUeno Tsukuba,Ibaraki, Japan National Institute for BasicBiology Okazaki,Aichi, Japan ISSN 2365-0613 ISSN 2365-0621 (electronic) SpringerBiographies ISBN978-981-13-6755-7 ISBN978-981-13-6756-4 (eBook) https://doi.org/10.1007/978-981-13-6756-4 LibraryofCongressControlNumber:2019932827 TranslationfromtheJapaneselanguageedition:TennoketoSeibutsugakubyHideoMohri©2015Hideo Mohri,publishedbyAsahiShimbunPublicationsInc.AllRightsReserved. ©SpringerNatureSingaporePteLtd.2019 Thisworkissubjecttocopyright.AllrightsarereservedbythePublisher,whetherthewholeorpart of the material is concerned, specifically the rights of translation, reprinting, reuse of illustrations, recitation, broadcasting, reproduction on microfilms or in any other physical way, and transmission orinformationstorageandretrieval,electronicadaptation,computersoftware,orbysimilarordissimilar methodologynowknownorhereafterdeveloped. The use of general descriptive names, registered names, trademarks, service marks, etc. in this publicationdoesnotimply,evenintheabsenceofaspecificstatement,thatsuchnamesareexemptfrom therelevantprotectivelawsandregulationsandthereforefreeforgeneraluse. The publisher, the authors and the editors are safe to assume that the advice and information in this book are believed to be true and accurate at the date of publication. Neither the publisher nor the authorsortheeditorsgiveawarranty,expressorimplied,withrespecttothematerialcontainedhereinor for any errors or omissions that may have been made. The publisher remains neutral with regard to jurisdictionalclaimsinpublishedmapsandinstitutionalaffiliations. Cover illustration: Front cover—Emperor Akihito engaging in research on gobioid fish in 1984 (CourtesyoftheImperialHouseholdAgency). Backcover—Above:EmperorShowaexaminingaspecimenofseaurchinattheShimodaImperialVilla in1985(CourtesyoftheImperialHouseholdAgency);Below:PrinceAkishinoinfrontofthepanelof fowlsatSOKENDAI(TheGraduateUniversityforAdvancedStudies)(CourtesyofAkifumiOikawa) ThisSpringerimprintispublishedbytheregisteredcompanySpringerNatureSingaporePteLtd. The registered company address is: 152 Beach Road, #21-01/04 Gateway East, Singapore 189721, Singapore o n Hisako koAyak mesinJapa uriko do= Nori usna Y a o = m o m nceMikasakahito) ako PrinceTaka(Norihito) Tsuguk edbyposthu Pri(Ta Mas call e o ar Tree n eimei Takamatsu=KikukPrince(Nobuhito) hi=HanakoTakako PrinceKatsura(Yoshihito)=Nobuko Yoko erEmperorsandEmpresses Imperial Family 1Jinmu 122Meiji=Shoke 123Taisho=T 24EmperorShowaEmpressKojun=Prince=SetsukoChichibu(Hirohito)(Nagako)(Yasuhito) 125PrinceHitachikoKazukoAtsukoEmperorEmpress=(Masahito)()()AkihitoMichiko =PrinceKikoceMasakoAkishino=SayakoYasuko(Fumihito)PrinceMikasa(Tomohito) AikoMakoKakoHisahitoAkiko Form 1 gekoSac CrownPrin(Naruhito) hi S Emperor Hirohito (Showa) and his family in 1972. Upper left to lower right. Crown Princess Michiko, Corwn Pince Akihito, Emperor, Empress Nagako (Kojun), Prince Hitachi, Princess Hanako,PrincessYoriSayako,PrinceHiroNaruhito,andPrinceAyaFumihito.SourceCourtesy oftheImperialHouseholdAgency EmperorAkihitoandhisfamilyin2016.Lefttoright.CrownPrincessMasako,PrincessMako, Crown Prince Naruhito, Princess Aiko, Emperor, Empress Michiko, Prince Hisahito, Prince Akishino,PrincessKako,PrincessKiko.SourceCourtesyoftheImperialHouseholdAgency Preface to the English Edition It was decided that Emperor Akihito would abdicate the throne on April 30, 2019. He had expressed his intention to step down primarily as a result of difficulties in fulfillingofficialdutiesathisadvancedage.EmperorAkihito’ssincereandearnest commitment to his role can also be seen in his work as a biologist in his taxo- nomical researches on gobies. His father, the late Emperor Showa (Hirohito), also conducted outstanding research as a biologist in the taxonomy of marine inverte- brates, including hydrozoans and slime molds (myxomycetes), even during those difficultyearsbeforeandafterWorldWarII.PrinceHitachi,theyoungerbrotherof Emperor Akihito, is an expert in comparative oncology, Prince Akishino, his second son, is carrying out phylogenetical researches on catfish and fowl and researchonbiosophiastudies,andSayakoKuroda,hisfirstdaughter,isresearching birds, including the kingfisher. It is highly unusual that members of three genera- tionsofonemonarchicalfamilylinehavedevotedthemselvestobiologicalstudies. We are now hoping that His Majesty the Emperor will be able to enjoy his bio- logical researches to the fullest after his retirement as Emperor. The fact that the Emperor is a biologist is only vaguely known, even to the Japanese people. When I, as a biologist, prepared to publish Tennoke to seibut- sugaku (The Imperial Family and Biology, Asahi Shimbun Publications, 2015), I realized that there were only a few works that depicted the Imperial Family members as biologists. With such limited resources, it is even harder for non-Japanese people to obtain a clear understanding of the Imperial biologists of Japan.Giventhesecircumstances,IdecidedtotranslatethisoriginalJapanesebook intoEnglishinthehopesthatitwillintroducethesincerecontributionsofmembers oftheJapaneseImperialFamilytothefieldofbiologytotheworld.Thisbookalso devotes some pages to the International Prize for Biology, which was set up to commemorate the contributions of the two generations of Emperors to biology. vii viii PrefacetotheEnglishEdition ThecoverimageoftheJapaneseversion,Tennoketoseibut- sugaku (The Imperial Family and Biology, Asahi Shimbun Publications,2015) IampleasedtohavehadthegoodfortunetohavetheEnglishversionpublished by Springer, thanks to Ms. Yoko Kawazoe, an excellent translator. Several years have passed since the Japanese version was published; therefore, some additional material is included. For readers’ reference, the family tree of the Imperial Family andtwophotographsdepictingmembersoftheImperialFamilyarenewlyaddedat the beginning. The content and wording have been put forward in as simple a manner as possible for readers who are unfamiliar with Japanese history and culture. Iexpressmythankstothemanypersonsconcerned,includingProf.NaotoUeno of the National Institute for Basic Biology, Prof. Tetsuji Nakabo of Kyoto University, and Dr. Hiroshi Namikawa of the National Museum of Nature and Sciencefortheirhelpwiththetranslation.IalsoexpressmydeepestthankstoMses. Alexandrine Cheronet, Aiko Hiraguchi, Fumiko Yamaguchi and Momoko Asawa of Springer, and Yumiko Nara of Asahi Shimbun Publications for their support in publication. Tokyo, Japan Hideo Mohri Preface In November 2011, Emperor Akihito was hospitalized for two weeks at The University of Tokyo Hospital for treatment for mycoplasma pneumonia. Deeply concerned by the disaster of the Great East Japan Earthquake of March 11, 2011, Emperor Akihito and Empress Michiko had continued visiting the quake victims andafflictedareasalmosteveryweek,inadditiontotheiralreadybusyscheduleof official duties. One can imagine how tough this must have been, and these events must have caused fatigue to build up in the Emperor.In February of thefollowing year, Emperor Akihito had to undergo heart bypass surgery. On March 11, 2012, still not fully recovered after the surgery, he attended the memorial service to commemorate the first anniversary of the Great East Japan Earthquake. The fol- lowing May, he visited the UK to celebrate the Diamond Jubilee of Her Majesty Queen Elizabeth II. The Emperor and the Empress were scheduled to attend the award ceremony of the 27th International Prize for Biology, to be held at the Japan Academy in Ueno,Tokyo,onNovember28,2011,asoneoftheirofficialduties.Asdescribedin detail later, this prize was founded in 1985 to commemorate the 60-year reign of Emperor Showa (Hirohito) and his longtime devotion to biology and to promote biologicalresearches.Moreover,ontheoccasionofits25thanniversaryin2009,it was decided that the prize would also honor the path toward the further advance- mentofbiology,includingincommemorationoftheachievements oftheEmperor at that time, His Majesty Emperor Akihito, who had strived over many years to advance the study of the taxonomy of gobioid fishes while contributing continu- ously to the development of the prize. Emperor Akihito was discharged from the hospitalonlytwodaysbeforetheawardceremony.Presumably,hisearlydischarge wasbecauseheintendedtoattendtheceremony.Helookedforwardtoattendingit everyyearasoneofthebiologists.Unfortunately,hiswishdidnotcometrue,ashe had still not fully recovered his health, and the ceremony was conducted with the hastily arranged attendance of the Crown Prince Naruhito on behalf of Emperor Akihito. Surprisingly, however, the Empress was present at the buffet party cele- brating the prize winner, as she had been in past years. The Empress told me that ix

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