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ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES 1 I I G A L LLUSTRATED DENTIFICATION UIDE TO DULTS AND ARVAE N N A OF ORTHEASTERN ORTH MERICAN G B (C : C ) ROUND EETLES OLEOPTERA ARABIDAE Yves Bousquet 2 ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES This page intentionally left blank ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES 3 Illustrated Identifi cation Guide to Adults and Larvae of Northeastern North American Ground Beetles (Coleoptera: Carabidae) Y B VES OUSQUET Agriculture and Agri-Food Canada Ottawa, Ontario K1A 0C6 S –M OFIA OSCOW 2010 4 ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES (COLEOPTERA: CARABIDAE) Yves Bousquet Cover illustration: Cicindela lepida, photo taken by Olivier Lalonde. First published 2010 ISBN 978-954-642-522-5 Pensoft Series Faunistica No 90 ISSN 1312-0174 © PENSOFT Publishers All rights reserved. No part of this publication may be reproduced, stored in a retrieval system or transmitted in any form by any means, electronic, mechanical, photocopying, recording or otherwise, without the prior written permission of the copyright owner. Pensoft Publishers Geo Milev Str. 13a, Sofi a 1111, Bulgaria Fax: +359-2-870-42-82 [email protected] www.pensoft.net Printed in Bulgaria, March 2010 ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES 5 To Henri For introducing me to the world of carabids For the color habitus photographs pictured in this book and For his contagious love for entomology 6 ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES This page intentionally left blank CONTENTS 7 CONTENTS Introduction ...........................................................................................................11 Acknowledgments ................................................................................................11 Generalities 1. – Biology .......................................................................................................12 2. – Suprageneric classifi cation ......................................................................13 3. – Lists of northeastern species ..................................................................15 SECTION A – ADULTS Generalities 1. – Information included .............................................................................42 2. – Identifi cation ...........................................................................................45 3. – Structures and terms ..............................................................................47 4. – Abbreviations used ................................................................................57 Key to genera .........................................................................................................58 Comments and keys to species Tribe Pelophilini ...............................................................................................79 Tribe Nebriini ...................................................................................................80 Tribe Notiophilini ............................................................................................82 Tribe Cychrini ...................................................................................................85 Tribe Carabini ...................................................................................................87 Tribe Cicindelini ...............................................................................................92 Tribe Loricerini .................................................................................................98 Tribe Elaphrini ..................................................................................................99 Tribe Omophronini .......................................................................................105 Tribe Scaritini ..................................................................................................106 Tribe Clivinini .................................................................................................107 8 ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES Tribe Broscini .................................................................................................118 Tribe Trechini..................................................................................................118 Tribe Bembidiini .............................................................................................120 Subtribe Bembidiina ................................................................................121 Subtribe Xystosomina .............................................................................154 Subtribe Tachyina ....................................................................................154 Tribe Patrobini ................................................................................................162 Tribe Psydrini ..................................................................................................165 Tribe Brachinini ..............................................................................................165 Tribe Pterostichini ..........................................................................................170 Tribe Zabrini ...................................................................................................188 Tribe Oodini....................................................................................................209 Tribe Chlaeniini ..............................................................................................211 Tribe Licinini ...................................................................................................217 Tribe Harpalini ................................................................................................224 Subtribe Anisodactylina ..........................................................................225 Subtribe Stenolophina ............................................................................233 Subtribe Harpalina ..................................................................................247 Tribe Pentagonicini ........................................................................................264 Tribe Platynini .................................................................................................264 Subtribe Sphodrina ..................................................................................265 Subtribe Platynina ...................................................................................268 Tribe Atranini ..................................................................................................292 Tribe Perigonini ..............................................................................................293 Tribe Odacanthini ..........................................................................................294 Tribe Cyclosomini ..........................................................................................294 Tribe Lebiini ....................................................................................................294 Subtribe Cymindidina .............................................................................296 Subtribe Dromiusina ...............................................................................298 Subtribe Lebiina .......................................................................................300 Subtribe Calleidina ..................................................................................304 Tribe Galeritini ...............................................................................................305 Tribe Helluonini .............................................................................................306 CONTENTS 9 SECTION B – LARVAE Generalities 1. – Information included ...........................................................................374 2. – Identifi cation .........................................................................................374 3. – Structures and terms ............................................................................375 4. – Abbreviations used ..............................................................................379 Key to genera .......................................................................................................380 Comments and keys to species for some genera Tribe Pelophilini .............................................................................................390 Tribe Nebriini .................................................................................................390 Tribe Notiophilini ..........................................................................................394 Tribe Cychrini .................................................................................................396 Tribe Carabini .................................................................................................398 Tribe Cicindelini .............................................................................................403 Tribe Loricerini ...............................................................................................408 Tribe Elaphrini ................................................................................................410 Tribe Omophronini .......................................................................................415 Tribe Scaritini ..................................................................................................418 Tribe Clivinini .................................................................................................418 Tribe Broscini .................................................................................................423 Tribe Trechini..................................................................................................426 Tribe Bembidiini .............................................................................................428 Subtribe Bembidiina ................................................................................429 Subtribe Xystosomina .............................................................................430 Subtribe Tachyina ....................................................................................430 Tribe Patrobini ................................................................................................434 Tribe Psydrini ..................................................................................................437 Tribe Brachinini ..............................................................................................438 Tribe Pterostichini ..........................................................................................440 Tribe Zabrini ...................................................................................................455 Tribe Oodini....................................................................................................459 Tribe Chlaeniini ..............................................................................................461

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Illustrated Identification Guide to Adults & Larvae of Northeastern North America Ground Beetles: Coleoptera Carabidae (Faunistica) PDF

563 Pages·2010·18.93 MB·English
by  Yves Bousquet
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Preview Illustrated Identification Guide to Adults & Larvae of Northeastern North America Ground Beetles: Coleoptera Carabidae (Faunistica)

ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES 1 I I G A L LLUSTRATED DENTIFICATION UIDE TO DULTS AND ARVAE N N A OF ORTHEASTERN ORTH MERICAN G B (C : C ) ROUND EETLES OLEOPTERA ARABIDAE Yves Bousquet 2 ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES This page intentionally left blank ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES 3 Illustrated Identifi cation Guide to Adults and Larvae of Northeastern North American Ground Beetles (Coleoptera: Carabidae) Y B VES OUSQUET Agriculture and Agri-Food Canada Ottawa, Ontario K1A 0C6 S –M OFIA OSCOW 2010 4 ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES (COLEOPTERA: CARABIDAE) Yves Bousquet Cover illustration: Cicindela lepida, photo taken by Olivier Lalonde. First published 2010 ISBN 978-954-642-522-5 Pensoft Series Faunistica No 90 ISSN 1312-0174 © PENSOFT Publishers All rights reserved. No part of this publication may be reproduced, stored in a retrieval system or transmitted in any form by any means, electronic, mechanical, photocopying, recording or otherwise, without the prior written permission of the copyright owner. Pensoft Publishers Geo Milev Str. 13a, Sofi a 1111, Bulgaria Fax: +359-2-870-42-82 [email protected] www.pensoft.net Printed in Bulgaria, March 2010 ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES 5 To Henri For introducing me to the world of carabids For the color habitus photographs pictured in this book and For his contagious love for entomology 6 ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES This page intentionally left blank CONTENTS 7 CONTENTS Introduction ...........................................................................................................11 Acknowledgments ................................................................................................11 Generalities 1. – Biology .......................................................................................................12 2. – Suprageneric classifi cation ......................................................................13 3. – Lists of northeastern species ..................................................................15 SECTION A – ADULTS Generalities 1. – Information included .............................................................................42 2. – Identifi cation ...........................................................................................45 3. – Structures and terms ..............................................................................47 4. – Abbreviations used ................................................................................57 Key to genera .........................................................................................................58 Comments and keys to species Tribe Pelophilini ...............................................................................................79 Tribe Nebriini ...................................................................................................80 Tribe Notiophilini ............................................................................................82 Tribe Cychrini ...................................................................................................85 Tribe Carabini ...................................................................................................87 Tribe Cicindelini ...............................................................................................92 Tribe Loricerini .................................................................................................98 Tribe Elaphrini ..................................................................................................99 Tribe Omophronini .......................................................................................105 Tribe Scaritini ..................................................................................................106 Tribe Clivinini .................................................................................................107 8 ILLUSTRATED IDENTIFICATION GUIDE TO ADULTS AND LARVAE OF NORTHEASTERN NORTH AMERICAN GROUND BEETLES Tribe Broscini .................................................................................................118 Tribe Trechini..................................................................................................118 Tribe Bembidiini .............................................................................................120 Subtribe Bembidiina ................................................................................121 Subtribe Xystosomina .............................................................................154 Subtribe Tachyina ....................................................................................154 Tribe Patrobini ................................................................................................162 Tribe Psydrini ..................................................................................................165 Tribe Brachinini ..............................................................................................165 Tribe Pterostichini ..........................................................................................170 Tribe Zabrini ...................................................................................................188 Tribe Oodini....................................................................................................209 Tribe Chlaeniini ..............................................................................................211 Tribe Licinini ...................................................................................................217 Tribe Harpalini ................................................................................................224 Subtribe Anisodactylina ..........................................................................225 Subtribe Stenolophina ............................................................................233 Subtribe Harpalina ..................................................................................247 Tribe Pentagonicini ........................................................................................264 Tribe Platynini .................................................................................................264 Subtribe Sphodrina ..................................................................................265 Subtribe Platynina ...................................................................................268 Tribe Atranini ..................................................................................................292 Tribe Perigonini ..............................................................................................293 Tribe Odacanthini ..........................................................................................294 Tribe Cyclosomini ..........................................................................................294 Tribe Lebiini ....................................................................................................294 Subtribe Cymindidina .............................................................................296 Subtribe Dromiusina ...............................................................................298 Subtribe Lebiina .......................................................................................300 Subtribe Calleidina ..................................................................................304 Tribe Galeritini ...............................................................................................305 Tribe Helluonini .............................................................................................306 CONTENTS 9 SECTION B – LARVAE Generalities 1. – Information included ...........................................................................374 2. – Identifi cation .........................................................................................374 3. – Structures and terms ............................................................................375 4. – Abbreviations used ..............................................................................379 Key to genera .......................................................................................................380 Comments and keys to species for some genera Tribe Pelophilini .............................................................................................390 Tribe Nebriini .................................................................................................390 Tribe Notiophilini ..........................................................................................394 Tribe Cychrini .................................................................................................396 Tribe Carabini .................................................................................................398 Tribe Cicindelini .............................................................................................403 Tribe Loricerini ...............................................................................................408 Tribe Elaphrini ................................................................................................410 Tribe Omophronini .......................................................................................415 Tribe Scaritini ..................................................................................................418 Tribe Clivinini .................................................................................................418 Tribe Broscini .................................................................................................423 Tribe Trechini..................................................................................................426 Tribe Bembidiini .............................................................................................428 Subtribe Bembidiina ................................................................................429 Subtribe Xystosomina .............................................................................430 Subtribe Tachyina ....................................................................................430 Tribe Patrobini ................................................................................................434 Tribe Psydrini ..................................................................................................437 Tribe Brachinini ..............................................................................................438 Tribe Pterostichini ..........................................................................................440 Tribe Zabrini ...................................................................................................455 Tribe Oodini....................................................................................................459 Tribe Chlaeniini ..............................................................................................461

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