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Hydroclimate: The Influence of Water Movement in Freshwater Ecology PDF

295 Pages·1992·16.827 MB·English
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HYDROCLIMATE The Influence of Water Movement on Freshwater Ecology HYDROCLIMATE The Influence of Water Movement on Freshwater Ecology IAN R. SMITH Edinburgh, Scotland, UK ELSEVIER APPLIED SCIENCE LONDON and NEW YORK ELSEVIER SCIENCE PUBLISHERS LTD Crown House, Linton Road, Barking, Essex IGll 8JU, England Sole distributor in the USA and Canada ELSEVIER SCIENCE PUBLISHING CO., INC. 655 Avenue of the Americas, New York, NY 10010, USA WITH 17 TABLES AND 135 ILLUSTRATIONS © 1992 ELSEVIER SCIENCE PUBLISHERS LTD British Library Cataloguing in Publication Data Smith, Ian R. Hydroclimate: the influence of water movement on freshwater ecology. I. Title 574.52632 ISBN 1-85166-724-5 Library of Congress Cataloging-in-Publication Data Smith, Ian R. Hydroclimate: the influence of water movement on freshwater ecology/Ian R. Smith. p. cm. Includes bibliographical reference and index. ISBN 1-85166-724-5 1. Freshwater ecology. 2. Hydrometeorology. I. Title. QH541.5.F7S65 1992 91-31565 574.5'2632--dc20 CIP No responsibility is assumed by the Publisher for any injury and/or damage to persons or property as a matter of products liability, negligence or otherwise, or from any use or operation of any methods, products, instructions or ideas contained in the material herein. Special regulations for readers in the USA This publication has been registered with the Copyright Clearance Centre Inc. (Ccq, Salem, Massachusetts. Information can be obtained from the CCC about conditions under which photocopies of parts of this publication may be made in the USA. All other copyright questions, including photocopying outside the USA, should be referred to the publisher. All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted in any form or by any means, electronic, mechanical, photo copying, recording, or otherwise, without the prior written permission of the publisher. Preface Sun, wind and water draining from the land interact with the morphological features of a water body to create the environment experienced by freshwater plants and animals. The result of this interaction can be considered as the freshwater hydro climate and this plays the same role as that of conventional climate in terrestrial ecology. Agriculture, for example, has long been supported by specialist meteorological services which not only provide farmers with a sound interpretation of weather and climate without excessive technicality but which also consider relations between climate and the growth of crops and stock. There is a need for a similar service in freshwater ecology and applied biology. This book is the result of a number of years devoted to developing part of that service. It concentrates on the influence of all forms of water movement on the ecology of fresh waters. Water movement implies interest in both the quantity of water moving through river basins which reflects the climate of the catch ment as well as the nature of the fluid motion within the rivers and lakes of the basin. The book is not so much a review of recent research as an attempt to establish a logic-how knowledge of water movement can contribute to understanding the ecology of fresh waters. Two points follow directly. One is the need for the book to be comprehen sible to non-specialists and I hope this is achieved by emphasising simple physical arguments despite the rather formidable list of symbols. There is also the difficulty that, once you start discussing freshwater ecology, when do you stop? The chapter on freshwater ecosystems is, inevitably, over-simplified and the intention is simply to provide a basis for assessing the influence of physical conditions. The general theme, however, is that of stimulating ecologists to consider v vi Preface the implications of the environmental features described than to suggest that these are fully understood already. I am very conscious that there is a gap between this account of some of the physical features of rivers and lakes and the interests and activities of practising ecologists. I hope this book will help to bridge that gap. I hope the book will appeal to all with an interest in rivers and lakes-undergraduates, graduate students and practising professionals in both management and research. I also like to think that the book will be of interest to naturalists. Understanding processes is always satisfying and, to me, the motion of eddies in a river is just as delightful as the flight of a dipper. Engineers and hydrologists may find the account of the freshwater environment presents their own interests in a new light and give them an insight into the ecological conse quences of their activities. I would like to thank Peter Maitland, Chris Gibson and Alan Steel. All read parts of the book and made a number of helpful suggestions. Above all, I would like to thank Alec Lyle who read the whole draft, supplied most of the photographs and who, over the years, made working with rivers and lakes such a pleasure. The responsibility for the errors is, obviously, mine. I am also grateful to Ken Morris and David Jones for allowing me to use their photographs. IAN R. SMITH 18 Tarpin Road, Edinburgh, EH13 OHW, UK Contents Preface .. v Chapter 1 Hydroclimate . . . . . . . . . . . . . . . . . 1 Chapter 2 The structure and dynamics of river basins . 4 2.1 River basin structure . . . . . 4 2.1.1 Introduction . . . . . . . 4 2.1.2 Longitudinal river sections . 6 2.1.3 Catchment form . 8 2.1.4 River networks. . 12 2.2 Climate and streamflow 16 2.2.1 Runoff generation. 16 2.2.2 Climate classification 19 2.2.3 Average flow. . . . 22 2.2.4 Seasonal variation in flow 25 2.2.5 Discharge frequency. 26 2.2.6 Floods and low flows . . 31 Chapter 3 Outline of fluid dynamics . . 39 3.1 Basic features of fluid motion .. 40 3.1.1 Characteristics of fluid motion 40 3.1.2 Methods of analysis. 42 3.2 The nature of turbulence 43 3.2.1 Basic features . . . 43 3.2.2 Boundary layer flow. 48 3.2.3 Turbulent structure . 51 3.2.4 Advection-diffusion equation. 54 3.2.5 Free turbulence. . . . . . . 56 3.3 Hydrodynamic properties of particulate matter. 58 3.3.1 Introduction .. 58 3.3.2 Static properties . . 60 vii viii Contents 3.3.3 Bed deposits . . . . . . . 62 3.3.4 Particles in suspension. . . 62 3.4 The dynamics of particulate matter. 64 3.4.1 Introduction . . . . . . . 64 3.4.2 The onset of bed movement 66 3.4.3 Particles in suspension. 67 3.4.4 Shields' diagram . 70 3.4.5 Cohesive sediments. . 72 Chapter 4 The dynamics of hydraulic systems 74 4.1 Linear systems theory. . . . . 74 4.1.1 Introduction . . . . . . . . 74 4.1.2 Response to standard inputs . 77 4.1.3 Mass balance of a mixed basin 79 4.2 Incomplete mixing . . . . . . 82 4.2.1 Mixing processes . . . . . 82 4.2.2 Residence time distributions 83 4.2.3 Reaction efficiency 87 4.3 Compartment models . 88 4.3.1 Introduction . . 88 4.3.2 Parallel systems. . 89 4.3.3 Series systems . . 91 4.3.4 Systems with backflow . 93 4.3.5 Seasonal stratification 95 4.4 Final remarks . . . . . . 97 Chapter 5 Hydraulic characteristics of rivers . 98 5.1 River morphology . . . . . . . . . 98 5.1.1 Introduction . . . . . . . . . . 98 5.1.2 Classification of river morphology. 98 5.1.3 Relations between channel dimensions and flow 101 5.1.4 Particle size variation along a river 108 5.1.5 Micro-topography of river beds. 111 5.2 River mechanics . . . . . . . . 113 5.2.1 River as a boundary layer flow. 113 5.2.2 Stage discharge curves. . . . . 123 5.2.3 Dissolved and suspended loads. 124 5.2.4 Numerical example . . . . . . 126 5.3 Hydrodynamic features of river habitats. 131 5.3.1 Introduction . . . . . . . . . . . 131 Contents ix 5.3.2 River stability 132 5.3.3 Micro-hydraulics 136 5.3.4 Dispersion and mixing in rivers . 139 Chapter 6 Hydraulic characteristics of lakes . 142 6.1 Lake morphology. 142 6.1.1 Introduction 142 6.1.2 Standard measures 142 6.1.3 Hydrodynamic measures. 146 6.2 Wind characteristics. 148 6.2.1 Introduction 148 6.2.2 Features of land station data . 150 6.2.3 Wind over water 151 6.3 The hydraulic structure of lakes 155 6.4 Wind-water interactions. 157 6.4.1 Basic processes . 157 6.4.2 Empirical observations. 158 6.4.3 Langmuir circulations . 159 6.4.4 Hydrostatic balance . 161 6.4.5 Vertical current structure 162 6.5 Motion in isothermal lakes. 163 6.5.1 Introduction 163 6.5.2 Two dimensional steady circulation 165 6.5.3 Circulation in irregularly shaped lakes . 167 6.5.4 Surface seiches . 169 6.6 Motion in stratified lakes 173 6.6.1 The occurrence of stratification . 173 6.6.2 Properties of a stratified profile . 176 6.6.3 Thermocline tilt and internal seiches 178 6.6.4 Regimes of stratified motion . 179 6.6.5 Steady state circulation in stratified lakes 183 6.7 Surface waves 183 6.7.1 Wave characteristics. 183 6.7.2 Deep water waves. 186 6.7.3 Shore zone waves . 189 6.8 Other features of lake motion 191 6.8.1 Inflow and outflow dynamics . 191 6.8.2 Turbulence in lakes . 195 6.8.3 Exchanges between sediment and water 199 6.9 Final remarks 202 x Contents Chapter 7 River basin modification. . 204 7.1 Reservoirs and flow regulation . 204 7.1.1 Types of reservoir . 204 7.1.2 Storage theory . . . . . 206 7.1.3 Reservoir features. . . . 208 7.2 Modifications to river habitats 209 7.2.1 Introduction . . . . . . 209 7.2.2 Effects of flow changes . 210 7.2.3 Changes in river structure 211 Chapter 8 Freshwater Ecosystems . 215 8.1 Idealised ecosystems . . . . 215 8.1.1 Ecosystems and ecosystem models 215 8.1.2 Idealised river ecosystems 219 8.1.3 Idealised lake ecosystems 222 8.2 Rate controlled growth 224 8.2.1 Introduction 224 8.2.2 Resources . . 224 8.2.3 Regulators. . 226 8.2.4 Physical losses 228 8.2.5 General remarks 229 8.3 Event controlled growth. 229 8.3.1 The nature of events 230 8.3.2 Species response to events . 231 8.3.3 Hydrodynamic stress and population loss 234 8.3.4 Population loss due to a sudden discharge of toxin 235 8.4 Space controlled growth. . . . . . . . 237 8.4.1 Spatial variation in plankton growth. 237 8.4.2 Habitat availability 239 8.5 Final remarks 240 Chapter 9 Synthesis 242 9.1 Background . 242 9.1.1 Interests involved in river basins 242 9.1. 2 The logic of hydro dim ate investigations 244 9.1.3 Modelling aquatic systems 246 9.2 River models. . . . . . . 246 9.2.1 River description . . . 247 9.2.2 Transport and storage. 249 9.2.3 River ecology. . . . . 253 Contents xi 9.3 Lake models. . . . . . . . . 256 9.3.1 Introduction ...... . 256 9.3.2 Basic features of lake hydroclimate 256 9.3.3 Lake ecology. 259 References . 261 Appendix 1: The properties of water . 271 A.l Viscosity-temperature relationship. 271 A.2 Density-temperature relationship 271 Appendix 2: Notation 273 Index ... 279

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