Human Anatomy & Physiology Sixth Edition Elaine N. Marieb Icon Identification Guide Word Roots, Prefixes, Suffixes, and Combining Forms HOMEOSTATIC IMBALANCE Prefixes and Combining Forms Homcostatic imbalances are indicated by an orange symbol followed by a heading. The icon is used to reinforce the concept that disease a-, an- absenceorlack acardia, lackofa heart; anaerobic, in the ab- is viewed as a loss of homeostasis, hence an imbalance in body sence ofoxygen functioning. ab- departingfrom; awayfrom abnormal, departingfrom normal acou- hearing acoustics, the science of sound ac-, aero- extremeorextremity-, peak acrodermatitis, inflammation ofthe skin ofthe extremities ad- toortoward adorbital, toward theorbit aden-, adeno- gland adeniform, resembling a gland in shape Keyquestions for selected figures address the concepts orprocesses adren- toward thekidney adrenal gland, adjacent to the kidney aero- air aerobic respiration, oxygen-requiringmetabolism depicted in the illustrations or in related topics covered earlier in the text. Answers to these questions appear upside down on the af- toward afferentneurons,whichcarryimpulsestothecentralner- vous system same page as the figure. agon- contest agonistic and antagonistic muscles, which oppose each other alb- white corpus albicans ofthe ovary, awhite scar tissue InterActive Physiology® icons in the Chapter Summaryof ailment- nourish alimentarycanal, ordigestive tract appropriate chapters refer the student to specific exercises on related allel- ofoneanother alleles, alternative expressions ofagene amphi- on both sides; ofboth kinds amphibian, an organism capa- InterActive Physiology® topics. ble oflivinginwaterand on land ana- apart, up, again anaphase of mitosis, when the chromosomes Critical Thinking and separate anastomos- cometogether arteriovenousanastomosis, aconnection Clinical Application between an artery and avein Questions aneurysm a widening aortic aneurism, a weak spot that causes en- Critical Thinkingand Clinical Applications Questions at the end of largement of the bloodvessel everychapter are identified bya combined stethoscope and question angi- vessel angiitis, inflammationofa lymphvesselorbloodvessel mark icon. angin- choked angina pectoris, a choked feeling in the chest due to dysfunctionof the heart ant-, anti- opposedto-,preventingorinhibiting anticoagulant, asub- stance that prevents blood coagulation ante- preceding; before antecubital, in front of the elbow aort- greatartery aorta ap-, api- tip, extremity apexofthe heart append- hangto appendicular skeleton i aqua-, aque- water aqueous solutions arbor tree arborvitaeofthecerebellum, the treelikepatternofwhite matter areola- open space areolar connective tissue, a loose connective tissue arrect- upright arrector pili muscles of the skin, which make the hairs stand erect arthr-, arthro- joint arthropathy, anyjoint disease artic- joint articularsurfaces ofbones, the points ofconnection atri- vestibule atria, upperchambers ofthe heart auscult- hsten auscultatorymethod for measuringbloodpressure aut-, auto- self autogenous, self-generated ax-, axi-, axo- axis, axle axial skeleton, axis ofvertebral column azyg- unpaired azygousvein, anunpaired vessel baro- pressure baroreceptors for monitoring bloodpressure basal base basal lamina ofepithelialbasement membrane bi- two bicuspid, havingtwo cusps bili- bile bilirubin, abilepigment bio- life biology, the studyoflife and livingorganisms blast- budorgerm blastocyte, undifferentiated embryoniccell brachi- arm brachial plexus of peripheral nervous system supplies the arm brady- slow bradycardia, abnormally slow heartrate brev- short peroneus brevis, a short leg muscle broncho- bronchus bronchospasm, spasmodiccontractionof bronchial muscle bucco- cheek buccolabial, pertaining to the cheekand lip calor- heat calories, a measure ofenergy capill- hair blood and lymphcapillaries caput- head decapitate, remove the head dys- difficult, faulty, painful dyspepsia, disturbed digestion carcin- cancer carcinogen, a cancer-causingagent ec-, ex-, ecto- out, outside, awayfrom excrete, to remove materials cardi, cardio- heart cardiotoxic, harmful to the heart horn the body cameo- flesh trabeculae carneae, ridges of muscle in the ventricles ectop- displaced ectopic pregnancy; ectopic focus for initiation of ofthe heart heart contraction carot- lj carrot, 2) stupor 1) carotene, anorangepigment; 2) carotid edem- swelling edema, accumulation ofwater inbodytissues arteries in the neck, blockage causes fainting ef- away efferent nerve fibers, which carry impulses away from the cata- down catabolism, chemicalbreakdown central nervous system caud- tail caudal (directional term) ejac- toshootforth ejaculationofsemen cec- blind cecum oflarge intestine, a blind-endedpouch embol- wedge embolus, anobstructiveobjecttravelingintheblood- cele- abdominal celiac artery, in the abdomen stream cephal- head cephalometer, an instrument for measuringthe head en-, em- in, inside encysted, enclosed in a cystorcapsule cerebro- brain, especiallythe cerebrum cerebrospinal, pertainingto enceph- brain encephalitis, inflammationofthe brain thebrain and spinal cord endo- within, inner endocytosis, takingparticles into a cell cervic-, cervix neck cervixofthe uterus entero- intestine enterologist, one who specializes in the study of chiasm- crossing opticchiasma, where optic nerves cross intestinal disorders chole- bile cholesterol; cholecystokinin, abile-secretinghormone epi- over, above epidermis, outer layerof skin chondr- cartilage chondrogenic, giving rise to cartilage erythr- red erythema, rednessofthe skin erythrocyte, redbloodcell ; chrom- colored chromosome, so namedbecause theystain darkly eso- within esophagus cili- smallhair ciliated epithelium eu- well euesthesia a normal state ofthe senses circum- around circumnuclear, surroundingthe nucleus excret separate excretory system clavic- key clavicle, a "skeleton key" exo- outside, outerlayer exophthalmos, an abnormal protrusion of co-, con- together concentric, commoncenter, togetherinthecenter the eye from the orbit coccy- cuckoo coccyx, which is beak-shaped extra- outside, beyond extracellular, outside the body cells of an cochlea snailshell the cochlea of the inner ear, which is coiled like organism a snail shell extrins- from the outside extrinsic regulation of the heart coel- hollow coelom, theventral bodycavity fasci-, fascia- bundle, band superficial and deep fascia commis- united gray commissure of the spinal cord connects the fenestr- window fenestrae of the innerear; fenestrated capillaries two columns ofgraymatter ferr- iron transferrin, ferritin, both iron-storage proteins concha shell nasal conchae, coiled shelves of bone in the nasal flagell- whip flagellum, the tail of a sperm cell cavity flat- blow, blown flatulence contra- against contraceptive, agent preventingconception folli- bag, bellows hairfollicle corn-, cornu- horn stratum corneum, outer layer of the skin com- fontan- fountain fontanels of the fetal skull posed of (horny) cells foram- opening foramen magnum ofthe skull corona crown coronal suture ofthe skull foss- ditch fossa ovalis of the heart; mandibular fossa ofthe skull corp- body corpse, corpus luteum, hormone-secreting body in the gam-, garnet- married, spouse gametes, the sexcells ovary gangli- swelling, orknot dorsal rootganglia ofthe spinal nerves cort- bark cortex, the outer layer of the brain, kidney, adrenal gastr- stomach gastrin, a hormone that influences gastric acid glands, and lymph nodes secretion cost- rib intercostal, between the ribs gene beginning, origin genetics crani- skull craniotomy, a skull operation germin- grow germinal epithelium of thegonads crypt- hidden cryptomenorrhea, a condition in which menstrual gero-, geront- oldman gerontology, the study of aging symptoms are experienced butno external loss ofblood occurs gest- carried gestation, the period from conception to birth cusp- pointed bicuspid, tricuspidvalves ofthe heart glauc- gray glaucoma, which causes gradual blindness cutic- skin cuticle ofthe nail glom- ball glomeruli, clusters ofcapillaries in the kidneys cyan- blue cyanosis, blue colorofthe skin due to lackofoxygen glosso- tongue glossopathy, anydisease ofthe tongue cyst- sac, bladder cystitis, inflammation ofthe urinarybladder gluco-, glyco- gluconeogenesis, theproductionofglucose from non- cyt- cell cytology, the studyofcells carbohydrate molecules de- undoing, reversal,loss, removal deactivation, becominginactive glute- buttock gluteus maximus, largest muscle ofthe buttock decid- fallingoff deciduous (milk) teeth gnost- knowing the gnostic sense, a sense of awareness ofself delta triangular deltoid muscle, roughly triangular in shape gompho- nail gomphosis, the term applied to the joint between den-, dent- tooth dentin ofthe tooth tooth and jaw dendr- tree, branch dendrites, telodendria, both branches of a gon-, gono- seed, offspring gonads, the sexorgans neuron gust- taste gustatory sense, the sense of taste derm- skin dermis, deep layerof the skin hapt- fasten, grasp hapten, apartial antigen desm- bond desmosome, whichbinds adjacentepithelial cells hema-, hemato-, hemo- blood hematocyst, a cyst containingblood di- twice, double dimorphism, having two forms hemi- half hemiglossal, pertainingto one-halfof the tongue dia- through, between diaphragm, thewall throughorbetween two hepat- hver hepatitis, inflammation ofthe liver areas hetero- differentorother heterosexuality, sexual desire for aperson dialys- separate, break apart kidney dialysis, in which waste prod- ofthe opposite sex ucts are removed from theblood hiat- gap the hiatus of the diaphragm, the opening through which diastol- stand apart cardiac diastole, between successive contrac- the esophagus passes tions ofthe heart hippo- horse hippocampus ofthe brain, shaped like a seahorse diure- urinate diuretic, a drugthat increases urine output hirsut- hairy hirsutism, excessive bodyhair dors- the back dorsal; dorsum,- dorsiflexion hist- tissue histology, the studyoftissues due-, duct lead, draw ductus deferenswhichcarries spermhornthe holo- whole holocrine glands, whose secretions arewhole cells epididymis into the urethra duringejaculation horn-, homo- same homeoplasia, formationoftissue similartonor- dura hard dura mater, toughouter meninx mal tissue; homocentric, havingthe same center hormon- toexcite hormones meningo- membrane meningitis, inflammation of the membranes humor- afluid humoral immunity, which involves antibodiescircu- of the brain lating in the blood mer-, mero-, apart merocrineglands, thesecretionsofwhichdonot hyal- clear hyaline cartilage, which has novisible fibers include the cell hydr-, hydro- water dehydration, loss ofbodywater meso- middle mesoderm, middlegerm layer hyper- excess hypertension, excessive tension meta- beyond, between, transition metatarsus, the part of the foot hypno- sleep hypnosis, a sleeplike state between the tarsus and the phalanges hypo- below, deficient hypodermic, beneath the skin hypokalemia, metro- uterus metroscope, instrument forexaminingthe uterus ; deficiency of potassium micro- small microscope, aninstrumentusedtomakesmallobjects hyster-, hystero- uterus or womb hysterectomy, removal of the appear larger uterus; hysterodynia, pain in thewomb mictur- urinate micturition, the act ofvoiding the bladder ile- intestine ileum, the last portion ofthe small intestine mito- thread, filament mitochondria, small, filamentlike structures im- not impermeable, not permittingpassage, notpermeable located in cells inter- between intercellular, between the cells mnem- memory amnesia intercal- insert intercalated discs, the end membranes between ad- mono- single monospasm, spasm ofa single limb jacent cardiac muscle cells morpho- form morphology, the study of form and structure or intra- within, inside intracellular, inside the cell organisms iso- equal, same isothermal, equal, orsame, temperature multi- many multinuclear, havingseveral nuclei jugul- throat jugularveins, prominentvessels in the neck mural wall intramural ganglion, a nerve junctionwithin anorgan DNA juxta- near, close to juxtaglomerularapparatus, acellclusternext to muta- change mutation, change in the base sequence of the glomeruli in the kidneys myelo- spinalcord, marrow myeloblasts, cells of thebone marrow karyo- kernal nucleus karyotype, the assemblage of the nuclear myo- muscle myocardium, heart muscle chromosomes nano- dwarf nanometer, one billionth ofa meter kera- born keratin, thewater-repellentprotein ofthe skin narco- numbness narcotic, a drug producing stupor or numbed kilo- thousand kilocalories, equal to one thousand calories sensations kin-, kines- move kinetic energy, the energyof motion natri- sodium atrialnatriureticfactor, asodium-regulatinghormone labi-, labri- lip labial frenulum, the membrane which joins the lip necro- death necrosis, tissue death to the gum neo- new neoplasm, an abnormal growth lact- milk lactose, milk sugar nephro- kidney nephritis, inflammationofthe kidney lacun- space, cavity, lake lacunae, the spaces occupied by cells of neuro- nerve neurophysiology, thephysiologyofthenervoussystem cartilage and bone tissue noci- harmful nociceptors, receptors forpain lamell- small plate concentric lamellae, rings of bone matrix in nom- name innominate artery; innominate bone compact bone noto- back notochord, the embryonic structure that precedes the lamina layer, sheet basal lamina, part of the epithelial basement vertebral column membrane nucle- pit, kernel, httle nut nucleus lat- wide latissimus dorsi, a broad muscle of the back nutri- feed, nourish nutrition laten- hidden latentperiod ofa muscle twitch ob- before, against obstruction, impedingorblockingup later- side lateral (directional term) oculo- eye monocular, pertainingto one eye leuko- white leukocyte, white blood cell odonto- teeth orthodontist, onewho specializes inproperposition- leva- raise, elevate levator labii superioris, muscle that elevates up- ingofthe teeth in relation to each other per lip olfact- smell olfactory nerves lingua- tongue lingual tonsil, adjacent to the tongue oligo- few oligodendrocytes, neuroglial cellswith fewbranches lip-, lipo- fat, hpid lipophage, a cell that has taken up fat in its onco- a mass oncology, studyofcancer cytoplasm oo- egg ocyte, precursoroffemalegamete lith- stone cholelithiasis, gallstones ophthalmo- eye ophthalmology, the study of the eyes and related luci- clear stratum lucidum, clearlayerof theepidermis disease lumen light lumen, centerofa hollowstructure orb- circular orbicularis oculi, muscle that encircles the eye lut- yellow corpus luteum, ayellow, hormone-secretingstructurein orchi- testis cryptorchidism, failureofthe testes todescendinto the the ovary scrotum lymph water lymphatic circulation, return of clear fluid to the org- hving organism bloodstream ortho- straight, direct orthopedic, correction of deformities of the macro- large macromolecule, large molecule musculoskeletal system macula spot macula lutea, yellowspoton the retina osm- smell anosmia, loss ofsense ofsmell magn- large foramen magnum, largest openingofthe skull osmo- pushing osmosis mal- bad, abnormal malfunction, abnormalfunctioningofanorgan osteo- bone osteodermia, bonyformations in the skin mamm- breast mammarygland, breast oto- ear otoscope, adevice forexaminingthe ear mast- breast mastectomy, removal ofa mammarygland ov-, ovi- egg ovum, oviduct mater mother dura mater, pia mater, membranes that envelop the oxy- oxygen oxygenation, the saturation ofa substancewithoxygen brain pan- all, universal panacea, acure-all meat- passage external auditorymeatus, theearcanal papill- nipple dermal papillae, projections of the dermis into the medi- middle medial (directional term) epidermal area medull- marrow medulla, themiddleportionofthekidney, adrenal para- beside, near paraphrenias, inflammation of tissues adjacent gland, and lymph node to the diaphragm mega- large megakaryocyte, largeprecursor cell ofplatelets pect-, pectus breast pectoralis major, a largechestmuscle meio- less meiosis, nuclear division that halves the chromosome pelv- a basin pelvicgirdle, whichcradles thepelvic organs number peni- a tail penis; penileurethra melan- black melanocytes, which secrete the black pigment penna- a wing unipennate, bipennatemuscles,whosefascicleshave melanin a feathered appearance men-, menstru- month menses, the cyclic menstrual flow pent- five pentose, a 5-carbon sugar TM MAPing your course through the best media resources for A&P! )mpass With your purchase ofa new copy of Elaine N. 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Marieb, R.N., Ph.D. Holyoke Community College PEARSON San Francisco Boston NewYork CapeTown HongKong London Madrid Mexico City Montreal Munich Paris Singapore Sydney Tokyo Toronto Publisher: Daryl Fox Art Coordinator: Bettina Borer Sponsoring Editor: Serina Beauparlant Photo Researcher: Diane Austin Development Manager: Claire Brassert Copyeditor: Anita Wagner Project Editor: Mary Ann Murray Compositor: GTS Graphics Managing Editor: Wendy Earl Proofreader and Page Checker: Martha Ghent Production Supervisor: Michele Mangelli Indexer: Karen Hollister Art Development Editor: Laura Southworth Principal Artists: Imagineering STA Media Services Inc. and Text Developmental Editor: Irene Nunes Kristin Mount of Gray Matter Studios Associate Online Project Editor: Michelle Levine Manufacturing Buyer: Stacey Weinberger Editorial Assistant: Sarah Kaminker Executive Marketing Manager: Lauren Harp Text and Cover Designer: tani hasagawa Cover Photograph (from left to right): Naoya Tamura, Japan; Apolo Anton Ohno, USA; Marc Gagnon, Canada,- Steven Bradbury, Australia. Clive Mason/Getty Images. Text, photography, and illustration credits follow the Glossary. Library of Congress Cataloging-in Publication Data Marieb, Elaine Nicpon Human anatomy & physiology/ Elaine N. Marieb.—6th ed. cm. p. Includes index. ISBN 0-8053-5463-8 1. Human physiology. 2. Human anatomy. I. Title: Human anatomy and physiology. II. Title. QP34.5 .M265 2003 612-dc21 2003045951 Copyright © 2004 Pearson Education, Inc. Publishing as Pearson Benjamin Cummings San Francisco, CA94111 All rights reserved. Printed in the United States of America. This publication is protected by Copyright and permission should be obtained from the publisher prior to any prohibited reproduction, storage in a retrieval system, or transmission in any form orby any means, electronic, mechanical, photocopying, recording, or likewise. For information regarding permission(s), write to: Pearson Education, Inc., Rights and Permissions Department. Pearson Benjamin Cummings™ is a trademark of Pearson Education, Inc. ISBN 0-8053-5462-X (For sale copywith CD-ROM) ISBN 0-8053-5474-3 (Professional copywith CD-ROM) PEARSON ISBN 0-8057-1693-9 (NASTA Edition) Benjamin ISBN 0-321-20413-1 (Pearson International Edition) Cummings 5 6 7 8 9 10—QWV—07 06 05 www.aw.com/bc