2 2, 0 O Thirteenth Edition BD H 00 VE E R HENDERSON’S IIC E NTRI O ES T DICTIONARY OF N I O BIOLOGY L D N OA E Edited by Eleanor Lawrence R R G Y S Thirteenth Edition The thirteenth edition of this classic dictionary has O been updated in response to changes in the core Y O HENDERSON’S life-sciences fields, and now includes new terms F from bioinformatics, genomics, and proteomics. N DICTIONARY OF Key Features ’ BIOLOGY S (cid:2) over 22,000 entries (cid:2) clear definitions and additional explanations Edited by Eleanor Lawrence (cid:2) covers a wide range of topics (cid:2) includes new terms since the last edition, e.g. chameleon sequence, metabolomics, protein chip, boundary elements, lipid raft, L LUCA, RNA interference, ontology A W (cid:2) includes simplified, up-to-date classification tables for the kingdoms of living organisms R E N Henderson’s Dictionary of Biologycontinues to provide C an essential reference for students of any of the E biological sciences and for teachers and researchers. Eleanor Lawrenceis a freelance science writer and editor, and co-author of Longman Dictionary of Environmental Thirteenth ISBN 0-131-27384-1 Science. Edition 9 780131 273849 www.pearson-books.com HENDERSON’S DICTIONARY OF Biology Untitled-2 1 5/10/06, 11:56 AM We work with leading authors to develop the strongest educational materials in science, bringing cutting-edge thinking and best learning practice to a global market. Under a range of well-known imprints, including Prentice Hall, we craft high quality print and electronic publications which help readers to understand and apply their content, whether studying or at work. To find out more about the complete range of our publishing, please visit us on the World Wide Web at: www.pearsoned.co.uk Untitled-2 2 5/10/06, 11:56 AM HENDERSON’S DICTIONARY OF Biology Thirteenth Edition Eleanor Lawrence Untitled-2 3 5/10/06, 11:56 AM Pearson Education Limited Edinburgh Gate Harlow Essex CM20 2JE England and Associated Companies throughout the world Visit us on the World Wide Web at: www.pearsoned.co.uk First published 1920 Eighth edition 1963 Ninth edition 1979 Tenth edition 1989 Eleventh edition 1995 Twelfth edition 2000 Thirteenth edition 2005 © Oliver and Boyd Limited 1963 © Pearson Education Limited 1979, 1989, 1995, 2000, 2005 The right of Eleanor Lawrence to be identified as author of this work has been asserted by her in accordance with the Copyright, Designs and Patents Act 1988. All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted in any form or by any means, electronic, mechanical, photocopying, recording or otherwise, without either the prior written permission of the publisher or a licence permitting restricted copying in the united Kingdom issued by the Copyright Licensing Agency Ltd, 90 Tottenham Court Road, London W1T 4LP. ISBN-10: 0-13-127384-1 ISBN-13: 978-0-13-127384-9 British Library Cataloguing-in-Publication Data A catalogue record for this book is available from the British Library Library of Congress Cataloging-in-Publication Data Henderson’s dictionary of biology / [edited by] Eleanor Lawrence. — 13th ed. p. cm. Rev. ed. of: Henderson’s dictionary of biological terms. 12th ed., 2000. ISBN 0-13-127384-1 (pbk.) 1. Biology–Dictionaries. I. Title: Dictionary of biology. II. Lawrence, Eleanor, 1949– III. Henderson’s dictionary of biological terms. QH302.5.H45 2005 570′.3—dc22 2004043190 10 9 8 7 6 5 4 3 10 09 08 07 06 Typeset in 8/9pt Times NR by 35 Printed and bound in Malaysia The publisher’s policy is to use paper manufactured from sustainable forests. Untitled-2 4 5/10/06, 11:56 AM CONTENTS Preface to the Thirteenth Edition vii Abbreviations ix Units and conversions ix SI prefixes xi Greek alphabet xi Common Latin and Greek noun endings xii THE DICTIONARY 1–718 Appendices 1. An outline of the plant kingdom (Domain Eukarya) 719 2. An outline of the Fungi (Domain Eukarya) 723 3. An outline of the animal kingdom (Domain Eukarya) 725 4. An outline of the kingdoms protoctista and stramenopila (Domain Eukarya) 734 5. An outline of the Domain Bacteria 736 6. An outline of the Domain Archaea 739 7. Virus families 740 8. Etymological origins of some common word elements in biology 742 v Untitled-2 5 5/10/06, 11:56 AM Allie Untitled-2 6 5/10/06, 11:56 AM PREFACE TO THE THIRTEENTH EDITION Since the publication of the Twelfth Edition of Henderson’s Dictionary of Biological Terms new words have entered the biological vocabulary and old ones have acquired new or more precise usages. Progress in genetics, cell biology and biotechnology continues to be rapid, driven by the genome-sequencing programmes, bioinformatics, the use of gene manipulation and advances in experimental cell biology. The classification of the living world followed in this edition reflects the divi- sion of all living things into three domains or superkingdoms. Entries in the body of the dictionary are given for all the main phyla, divisions and classes of plants, fungi, animals, protists and prokaryotes (Bacteria and Archaea), with some orders being included for groups such as the insects, birds, mammals and flowering plants. There are also entries under many common names of organisms. The appendices at the back give a fuller outline of the various kingdoms. Viruses are covered by entries for the main groups and in an appendix. Terms are arranged in strict alphabetical order, disregarding hyphenation and spaces between words, with abbreviations and acronyms included in theirappropriate place within the body of the dictionary. Numbers, Greek letters, andconfigurational letters at the beginning of chemical names are ignored for alphabetization pur- poses. Within an entry, different meanings of a term are numbered and separated by semicolons. The abbreviations (bot.), (zool.), etc. have been used in some cases to indicate more clearly which subject area a definition refers to. Almost all the technical terms that may be used as part of a definition are defined within the dictionary; to avoid complicating the text with excessive cross-referencing, such terms are not generally indicated by (q.v.) within the body of an entry. Common suffixes and prefixes derived from Latin and Greek are entered in the body of the dictionary, along with their usual meanings, and Appendix 8 gives etymological origins of some common word elements. I should like to thank the staff of Longmans for their help and encouragement throughout the project. Comments concerning errors or omissions in this editionwill be greatly appreciated, so that they may be rectified in future reprints or editions. Eleanor Lawrence London, 2004 vii Untitled-2 7 5/10/06, 11:56 AM Allie Untitled-2 8 5/10/06, 11:56 AM ABBREVIATIONS a. adjective immunol. immunology adv. adverb L. Latin alt. alternative (synonym) microbiol. microbiology anat. anatomy mol. biol. molecular biology appl. applies or applied to mycol. mycology biochem. biochemistry n. noun bioinf. bioinformatics n haploid no. of chromosomes bot. botany neurobiol. neurobiology ca. circa (approximately) pert. pertaining to cf. compare plu. plural dev. biol. developmental biology p.p.m. parts per million EC Enzyme Commission number q.v. see (1978) r.n. Enzyme Commission e.g. for example recommended name, where it differs esp. especially from that used as the headword et al. and others sing. singular etc. and so forth sp. species (sing.) evol. evolution spp. species (plu.) genet. genetics v. verb Gk Greek virol. virology i.e. that is zool. zoology UNITS AND CONVERSIONS Basic SI units ampere, A (electric current) candela, cd (luminous intensity) kelvin, K (temperature) kilogram, kg (mass) metre, m (length) mole, mol (amount of substance) second, s (time) ix Untitled-2 9 5/10/06, 11:56 AM