Harpalini (Insecta: Coleoptera: Carabidae: Harpalinae)

2005·3.5 MB·

by  Larochelle, André

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Fauna of New Zealand
Ko te Aitanga Pepeke o Aotearoa
Number / Nama 53
Harpalini
(Insecta: Coleoptera:
Carabidae: Harpalinae)
A. Larochelle and M.-C. Larivière
Landcare Research, Private Bag 92170, Auckland, New Zealand
[email protected]
[email protected]
Manaaki
Whenua
P R E S S
Lincoln, Canterbury, New Zealand
2005
Copyright © Landcare Research New Zealand Ltd 2005
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Cataloguing in publication
LAROCHELLE, ANDRÉ 1940 Apr. 10
Harpalini (Insecta: Coleoptera: Carabidae: Harpalinae) / A. Larochelle & M.-C. Larivière,
– Lincoln, Canterbury, N.Z. : Manaaki Whenua Press, 2005.
(Fauna of New Zealand, ISSN 0111–5383 ; no. 53).
ISBN 0-478-09369-1
I. Larivière, Marie-Claude – II. Title III. Series
UDC 595.762.12(931)
Suggested citation:
Larochelle, A.; Larivière, M.-C . 2005. Harpalini (Insecta: Coleoptera: Carabidae:
Harpalinae). Fauna of New Zealand 53, 160 pp.
Prepared for publication by the series editor and the authors using computer-based text
processing, layout, and printing at Landcare Research, Private Bag 92170, Auckland, New
Zealand. Frontispiece digital image produced by M.-C. Larivière using the synchroscopy system
Auto-Montage®.
M~ori text by H. Jacob, Auckland.
Published by Manaaki Whenua Press, Landcare Research, P.O. Box 40, Lincoln, Canterbury, N.Z.
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Front cover: Tuiharpalus moorei new species (Illustrator: D. W. Helmore).
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Fauna of New Zealand 53 5
POPULAR SUMMARY HE WHAKARAPOPOTOTANGA
Class Insecta
Order Coleoptera
Family Carabidae
Subfamily Harpalinae
Tribe Harpalini
Harpaline ground beetles
The tribe Harpalini belongs to the subfamily Harpalinae Illustration / Whakaahua: Lecanomerus sharpi (Csiki)
(Coleoptera: Carabidae), which contains over 19 000 taxa (Illustrator / Kaiwhakaahua: D. W. Helmore).
worldwide. Molecular sequence data indicate Harpalinae
evolved in the Cretaceous Period (140–65 million years Ng~ p§tara Harpaline noho papa
ago).
The Harpalini form a diverse group, including over 240 NÇ te wh~nau wh~iti Harpalinae (Coleoptera: Carabidae)
genera and subgenera, and approximately 2000 species a ng~i Harpalini. Puta noa i te ao, 1900 Çna rÇpã. E ai ki
distributed in all biogeographic regions of the world. The ng~ raraunga raupapa r~poi ngota, nÇ te Takiw~ Creta-
present faunal review records 20 genera and 57 species for ceous tÇna kunenga mai (i te 140–65 miriona tau ki muri).
New Zealand. This should constitute nearly all the fauna. He iwi matahuhua tonu a ng~i Harpalini – he nui atu i
Compared with larger or warmer regions of the world, te 240 ng~ puninga me ng~ puninga iti, he ~hua 2000 ng~
e.g., Australia, which has a largely undescribed fauna with momo, kei ng~ takiw~ papawhenua-koiora katoa o te ao. I
over 160 known species, the New Zealand fauna may t‘nei tirohanga hou, kua tuhia he mauranga mÇ ng~ puninga
appear relatively small, but New Zealand is a very special e 20 me ng~ momo e 57 i Aotearoa. Ko te whakaaro ia, he
place – a biodiversity ‘hot-spot’ – with 75% of species (42 ruarua noa iho ng~ momo k~ore i mau i t‘nei tatauranga.
out of 57 species) and 55% of genera (11 out of 20 genera) Ina whakaritea ki ng~ whenua rahi ake, mahana ake o
found nowhere else in the world. The remaining fauna that te ao (hei tauira, ar~ a Ahitereiria me Çna momo 160 e
are not endemic to this country are made up of overseas mÇhiotia ana, tae atu ki te maha noa atu k~ore anÇ i
species introduced mainly from Australia. No native species whakaahuatia ~-kupu) t‘r~ ka whakaarohia he torutoru
is shared with Australia, although three native genera occur noa iho ng~ momo o Aotearoa. Engari he whenua ahurei
on both sides of the Tasman Sea, which suggests the New tonu a Aotearoa, in~ r~, ko t‘tahi 75% o ng~ momo (e 42 o
Zealand harpaline lineages have evolved mostly in isolation ng~ momo e 57) me t‘tahi 55% o ng~ puninga (11 o ng~
following the breakup of eastern Gondwanaland. puninga e 20) i Aotearoa nei, k~ore e kitea i whenua k‘. Ko
Harpalini are rather stout-bodied ground beetles with ‘r~ atu momo ehara nÇ konei taketake ake, he r~waho, ko
relatively short mandibles and other appendages, and a te nuinga i manene mai i Ahitereiria. Karekau he momo
body length of 3–20 mm. They are usually darkly coloured, m~ori o Aotearoa kei Ahitereiria anÇ e noho ana, engari e
have only one hair-bearing puncture above each eye, no toru ng~ puninga m~ori kei ng~ whenua e rua nei. E tohu
such puncture at the posterior angles of the pronotum, and ana t‘nei t‘r~ tonu pea i kukune motuhake mai ng~ t~tai
elytra that are rounded, not twisted, at the sides near the harpaline o Aotearoa i muri i te wehewehenga o te pito
apex. Some species living in caves or exhibiting strong r~whiti o Te Uri M~roa.
burrowing habits are characterised by paler bodies and He ~hua porotaka ng~ tinana o ng~ p§tara noho papa
reduced eyes. Harpalini, he ~hua poto ng~ kauwae me ‘r~ atu w~hanga
As observed in many other carabids, harpaline ground toro whakawaho o te tinana. Ko te roa o te tinana, mai i te
beetles are taxonomically diverse, generally abundant in 3 ki te 20 mm. He uriuri te tae o te nuinga, kotahi anake te
the field, and demonstrate ecological preferences and a m~rua whai weu i runga ake o ia karu, k~ore he m~rua
flexible set of responses to environmental factors. Because p‘nei i ng~ koki whakamuri o te papatua pohomua. He
of these features, the relative ease with which their ~hua kÇpuku ng~ pãkoro parirau i ng~ kaokao, i te takiw~
(continued overleaf) (haere tonu)
6 Larochelle & Larivière (2005): Harpalini (Insecta: Coleoptera: Carabidae)
populations may be sampled by reliable quantitative o te pito, k~ore e kÇrino. Heoi, ar~ ‘tahi momo noho ana,
methods (e.g., pitfall-trapping), and their potential use as kari rua r~nei k~ore e tino uriuri ng~ tinana, he iti ake anÇ
bioindicators and biocontrol agents, they represent an hoki ng~ karu.
attractive study group for biologists investigating P‘r~ anÇ i te maha tonu o ng~ carabid, he matahuhua
evolutionary and ecological hypotheses. ng~ whakarÇpãtanga o ng~ p§tara noho papa harpaline, he
As a result, Harpalini are well represented in New huhua anÇ t~ r~tou noho ki te taiao. He rerek‘ ng~ k~inga
Zealand entomological museums and collections – over noho e pai ana ki t‘n~, ki t‘n~, he t~wariwari anÇ te ~hua
5000 specimens were studied for this project. But despite o t~ r~tou aro atu ki ng~ ~huatanga taiao. N~ ‘nei
such high interest, no taxonomic revision of this group has ~huatanga, n~ te m~m~ anÇ ki te t§pako i ng~ taupori i
been produced until now. runga i ng~ tikanga ine ~-rahi tÇtika (p‘nei i te hopu ki te
Before the present revision, 13 genera and 36 species tomo), me tÇ r~tou pai anÇ pea hei tohu koiora, hei kaipatu
of Harpalini were known from New Zealand, but the koiora r~nei, he rÇpã t‘nei e arohia nuitia ana e ng~ tohunga
authors have found a number of species had been described koiora e whakam~tau ana i ng~ whakapae mÇ te kunenga
more than once under different names, and 23 species and me te taupuhi kaiao.
5 genera are new to science. Me te aha, he autaia tonu te maha o ng~ Harpalini e
The geographical distribution of native species was puritia ana ki ng~ whare pepeke me ng~ kohinga pepeke i
undocumented before this study. We now have a better Aotearoa – he nui ake i te 5000 ng~ tauira i ~ta tirohia mÇ
knowledge of their distribution patterns. The authors have t‘nei rangahautanga. Engari ahakoa te aro nui ki t‘nei iwi,
found, for example, that several species are restricted to k~tahi anÇ ka tirohia anÇ te whakarÇpãtanga o ng~ hanga
specific areas of New Zealand – the South Island northwest nei.
and the far north of the New Zealand appearing to have I mua i t‘nei o ng~ tirohanga, 13 ng~ puninga, e 36 ng~
been reservoirs, in geological time, of much of the genetic momo i mÇhiotia i Aotearoa nei. Engari ko t~ ng~ kaituhi i
diversity in New Zealand Harpalini, with several species kite ai, ar~ ‘tahi momo i tapaina ki ng~ ingoa e rua, neke
currently restricted to these regions. Of the two main atu r~nei, me Çna anÇ kupu whakaahua i te taha. }, e 23
islands of New Zealand, the North Island has the greatest ng~ momo, e 5 ng~ puninga k~ore i mÇhiotia i te ao pãtaiao
number of species (35 compared with 31 for the South i mua atu i t‘nei.
Island), and only 4 native species (Allocinopus sculpticollis, Waihoki, k~ore i tuhia te tohanga o ng~ momo m~ori i
Triplosarus novaezelandiae, Syllectus anomalus, mua atu i t‘nei rangahautanga. N~ t‘nei mahi rangahau kua
Euthenarus puncticollis) are shared between the two islands. kaha ake te mÇhio ki Ç r~tou tauira tohatoha. Hei tauira, i
Three genera (Gaioxenus, Parabaris, and Kupeharpalus) kitea ar~ ‘tahi momo maha tonu e noho motuhake ana ki
are found only on the North Island, whereas two genera ‘tahi rohe wh~iti o Aotearoa – ko te uru-m~-raki o Te
(Hakaharpalus and the cave-dwelling Pholeodytes) are Waipounamu me Muriwhenua ‘tahi tino m~t~puna, ~-w~
confined to the northwest of the South Island. Two genera papawhenua nei, o te matahuhuatanga o te huinga ira o
are restricted to the Three Kings Islands (Maoriharpalus, ng~ Harpalini o Aotearoa. He maha hoki ng~ momo k~ore
Kiwiharpalus). There is no genus endemic to the Chatham e kitea i waho atu o ‘nei takiw~. He maha ake ng~ momo i
Islands. Stewart Island also has no endemic taxa, but shares Te Ika a M~ui (e 35 ng~ momo), t‘n~ i Te Waipounamu (e
2 native species: Triplosarus novaezelandiae (with North 31 ng~ momo), ~, e 4 anake ng~ momo m~ori (Allocinopus
Island and South Island), Euthenarus brevicollis (with South sculpticollis, Triplosarus novaezelandiae, Syllectus
Island). anomalus, Euthenarus puncticollis) kei ng~ moutere e rua
Over 50% of native species (25 out of 42 species) are nei. E 3 ng~ puninga (Gaioxenus, Parabaris me
known from 10 populations or fewer. All but one of these Kupeharpalus) kei Te Ika a M~ui anake, e 2
very special species are new to science, and all are of (Hakaharpalus me Pholeodytes – he momo noho ana) kei
potential conservation concern. They are usually te uru-m~-raki o Te Waipounamu anake. E 2 ng~ puninga
taxonomically highly distinctive species with low dispersal k~ore e kitea i waho atu o Manawa-t~whi (Maoriharpalus,
power, often geographically localised in threatened habitats, Kiwiharpalus). Karekau he puninga e kitea ana i R‘kohu
and represented in collections by relatively few specimens anake. Waihoki, k~ore he rÇpã e kitea ana i Rakiura anake,
collected over many decades, which may indicate rare or engari e 2 Çna momo m~ori, ko: Triplosarus novaezelandiae
highly specialised species. (kei R‘kohu, kei Te Ika a M~ui me Te Waipounamu), me
No formal detailed study of the natural history of Euthenarus brevicollis (kei R‘kohu me Te Waipounamu).
individual species of New Zealand Harpalini has ever been He nui ake i te 50% o ng~ momo m~ori (e 25 o ng~
conducted, although Larochelle and Larivière (2001, Fauna momo e 42), nÇ roto i ng~ taupori 10, iti ake r~nei. E 24 o
of New Zealand 43) summarised information available from ‘nei momo, k~tahi anÇ ka mÇhiotia i te ao pãtaiao, ~, ko te
the literature, material in entomological collections, personal katoa me ~ta tiaki ka tika kei koreh~h~ haere. Ko te nuinga,
communications from carabid collectors, and their own he m~m~ ki te wehewehe, t‘tahi i t‘tahi, i te nui o ng~
personal field observations. rerek‘tanga, k~ore e kaha te marara haere, e noho wh~iti
Most native species are flightless, having vestigial ana ki ng~ ripoinga mÇrearea, ~, he ruarua ng~ tauira o t‘n~,
membranous wings (reduced to small wing buds), and live o t‘n~ kua kohia i roto i ng~ tekau tau, e tohu ana he momo
within the confines of native habitats, mostly forests onge tonu pea, he tino wh~iti r~nei Ç r~tou k~inga noho.
(haere tonu)
(continued overleaf)
Fauna of New Zealand 53 7
(especially along streams) and wet habitats, also tussock K~ore anÇ i ~ta rangahaua ng~ h§tori m~ori o t‘n~, o
grasslands and caves. Most Harpalini species are moisture t‘n~ momo o ng~i Harpalini i Aotearoa, engari i
loving and live at the surface of the soil and in leaf litter; whakar~popotohia e Larochelle r~ua ko Larivière (2001,
they also live in caves, and occasionally on plants and Fauna of New Zealand 43) ng~ p~rongo kua tuhia, ng~
trees. Dispersal in most native species is achieved by rauemi i ng~ kohinga pepeke, ng~ whakawhitinga kÇrero
running over the ground; most species are moderate runners, ki ng~ kaikohikohi carabid, me Ç r~ua ake kitenga i te taiao.
except for the long-legged, fast-running cave species He rere kore te nuinga o ng~ momo m~ori, he tumu
(Syllectus, Pholeodytes). In general, Harpalini have relatively parirau noa iho, he mea hanga ki te kiriuhi, e toe mai ana.
short legs and, sometimes, strongly reduced eyes, indicative Ka mutu, noho ai te nuinga ki ng~ ripoinga m~ori, ar~, ki
of strong burrowing habits. ng~ ngahere m~ori (ko ng~ tahataha o ng~ kÇawa t‘tahi tino
All adventive species are able to fly, having long or k~inga) tae atu ki ng~ k~inga kÇreporepo, ng~ whenua
fully developed membranous wings, and live mostly in p~t§t§ taranui me ng~ ana. He pai ki te nuinga t‘nei mea te
highly modified environments (often around human haukã, ~, t‘r~ ka noho ki te mata tonu o te whenua, ki ng~
dwellings), except for Haplanister crypticus, which has rau popo. Ar~ anÇ ‘tahi ko te ana tÇ r~tou w~hi noho, ~, he
also managed to invade native forests. torutoru ng~ momo ka piri tahi ki ng~ r~kau me ‘r~ atu
The collecting season of newly emerged adults suggests tipu. Ko te tikanga t§tari a te nuinga o ng~ momo m~ori, ko
Harpalini species may mate in the spring or autumn. For te takahi i te nuku o te whenua; he ~hua tere te haere a te
most species, adults are active during all months of the nuinga, engari he tere tonu te momo noho ana, waewae roa
year, but are generally less active during cooler months. (Syllectus, Pholeodytes). He poto ng~ waewae o te nuinga
No data are available on the feeding preferences of o ng~i Harpalini, ~, he tino ngoikore ng~ karu, e tohu ana
Harpalini native to New Zealand. Larochelle (1990, Food he kari rua, he noho rua t~ r~tou mahi.
of carabid beetles of the World) showed that on a world Katoa ng~ momo r~waho, he mÇhio ki te rere, ~, he
basis Harpalini feed on both animal and vegetable matter, roa, he pakari r~nei Ç r~tou parirau kiriuhi. Noho ai te
but tend to favour the latter. The mandibles of nuinga ki ng~ taiao kua kaha rawekehia e te ringa tangata
Hakaharpalus, Kiwiharpalus, Syllectus, Pholeodytes, and (he maha e noho tata ana ki ng~ whare), h~unga anÇ te
Maoriharpalus are unusually long in native Harpalini, which Haplanister crypticus. Kua urutomo anÇ t‘nei n~ i ng~
may suggest a specialised type of feeding. In addition, the ngahere m~ori.
strongly notched labrum (upper mouth part at base of Ko te w~ e kohikohia ai ng~ p§tara pakeke k~tahi anÇ
mandibles) of Maoriharpalus is reminiscent of, although ka puta ake ki te ao e tohu ana t‘r~ pea ko te kÇanga, ko te
not necessarily equivalent to, the condition observed in the ngahuru r~nei te w~ e whakaputa uri ai a ng~i Harpalini.
tribe Licinini, where species feed on hard-bodied MÇ te nuinga o ng~ momo, e oreore ana ng~ mea pakeke i
invertebrates, e.g., snails. ng~ marama katoa o te tau, engari ka ~hua ngoikore ake i
This faunal review was written with a wide audience in ng~ marama makariri ake.
mind. It aims to provide an inventory of New Zealand Karekau he raraunga mÇ ng~ kai e pai ana ki ng~
taxa, a concise treatment of their taxonomy, easy-to-follow Harpalini m~ori. Heoi, n~ Larochelle (1990, Food of carabid
identification keys, and several illustrations and maps, as beetles of the World) i whakaatu kai ai ng~ Harpalini o te
well as a summary of all available information on species ao i te kiko me te ota, engari ko te ota pea t~ r~tou tino kai.
distribution, ecology, biology, and dispersal power. It is He roa ake ng~ kauwae o Hakaharpalus, Kiwiharpalus,
one step in the authors’ goal to to reach an overall Syllectus, Pholeodytes, me Maoriharpalus i ‘r~ o te nuinga
understanding of the carabid fauna within a reasonable time o ng~ momo m~ori, e tohu ana pea he rerek‘ ng~ momo kai
frame, and to make relatively large amounts of information kainga ai e r~tou. I tua atu i t‘r~, ar~ ‘tahi k~niwha nui i te
available for practical use by a wide range of end-users. It ngutu o runga o Maoriharpalus, e ~hua rite ana ki t‘r~ e
is hoped this kind of faunal taxonomy will provide both a kitea ana i te iwi Licinini. Ko t~ t‘r~ iwi, he kai i ng~ hanga
solid foundation for studies of other types and the baseline tuar~-kore, tinana m~rÇ, p‘r~ i te ngata.
information required by systematists, identifiers, ecologists, He mea tuhi t‘nei tirohanga hou kia m~rama ai ng~
and other biologists, as well as by biosecurity and kÇrero ki te tokomaha. E whai ana kia takoto mai he r~rangi
conservation managers. o ng~ rÇpã e noho ana ki Aotearoa nei, he whakam~rama
poto o Ç r~tou t~tai hono, he ara tautohu m~m~, he
whakaahua, he mahere, tae atu ki t‘tahi whakar~-
Contributor André Larochelle was born and educated in
popototanga o ng~ p~rongo e w~tea ana mÇ te tohanga o
Québec, graduating in 1974 with a Brevet d’Enseignement
ng~ momo, te taupuhi kaiao, te koiora, me te kaha ki te
spécialisé from the Université du Québec à Montréal. He
t§tari haere i a r~tou anÇ. He takahanga noa t‘nei i roto i te
taught ecology at the Collège Bourget, Rigaud, Québec, up
to 1990. With the encouragement of the late carabid spe- wh~inga roa a ng~ kaituhi kia wh~nui noa ake te
cialist Carl H. Lindroth, André very quickly becames inter- m~ramatanga ki ng~ carabid i roto i te tekau tau pea e tã
ested to the study of ground beetles. From 1975 to 1979 he mai nei, kia hora he p~rongo huhua hei ~whina i ng~ t~ngata
was the co-editor of two entomological journals, Cordulia tokomaha i roto i ~ r~tou kaupapa maha. Ko te tãmanako
and Bulletin d’inventaire des insectes du Québec. From ia, kia noho t‘nei tãmomo whakarÇpãtanga hei tã~papa
(continued overleaf) (haere tonu)
8 Larochelle & Larivière (2005): Harpalini (Insecta: Coleoptera: Carabidae)
1986 to 1992, he was honorary curator to the Lyman Ento- mÇ ng~ mahi rangahau i ‘tahi atu pepeke, hei puna kÇrero
mological Museum and Research Laboratory, McGill Uni- hoki m~ ng~ kait~tai whakapapa, ng~ kaitautohu, ng~
versity, Québec. In 1992, André moved to New Zealand to tohunga taupuhi kaiao, me ‘tahi atu ringa koiora, tae atu ki
work as a research scientist. Currently, he is a Research ng~ kaiwhakahaere haumaru koiora, tiaki taiao.
Associate with the New Zealand Arthropod Collection,
Landcare Research, Auckland. André has written over 400
papers on the distribution, ecology, biology, and dispersal
power of North American carabids and other insects (in-
cluding two handbooks on the Heteroptera of Québec). In I wh~nau mai t‘r~ atu o ng~ kaituhi, a André Larochelle,
1993 he was co-author of a “Catalogue of Carabidae of i Québec. I reira anÇ ia e kura ana, ~, nÇ te tau 1974 ka
whakawhiwhia ki tana tohu Brevet d’Enseignement
spécialisé, mai i te Whare W~nanga o Québec ki Montreal.
Taka mai ki te tau 1990, e whakaako ana ia i te m~tauranga
taupuhi kaiao i te K~reti Bourget, i Rigaud, Québec. N~
ng~ akiaki a t‘r~ tohunga carabid kua riro nei i te tirohanga
kanohi, a Carl H. Lindroth, ka tere tupu tana hiahia ki te
rangahau i ng~ p§tara noho papa. Mai i te 1975 ki te 1979
ko ia t‘tahi o ng~ ‘tita o ‘tahi hautaka m~tai pepeke e rua,
ar~, o Cordulia me te Bulletin d’inventaire des insectes du
Québec. Mai i te 1986 ki te 1992, ko ia te kaitiaki utu-kore
o te Whare Pupuri, Rangahau Pepeke o Lyman, i te Whare
W~nanga o McGill, i Québec. I te tau 1992, ka neke mai a
André ki Aotearoa, ka mahi hei kaipãtaiao rangahau. I
t‘nei w~, e noho ana ia hei Kairangahau i te Kohinga
Angawaho o Aotearoa, i Manaaki Whenua ki T~maki-
makau-rau. He nui ake i te 400 ng~ kÇrero kua tuhia e
André mÇ te tohanga, te taupuhi kaiao, te koiora, me te
t§taringa o ng~ carabid me ‘tahi atu aitanga pepeke o
Amerika ki te Raki (tae atu ki ‘tahi pukapuka e rua e
whakaahua ana i ng~ Heteroptera o Québec). I te tau 1993
ko ia t‘tahi o ng~ kaituhi i te “R~rangi o ng~ Carabidae o
Amerika ki te raki o M‘hiko”. Ka whakaputaina e r~ua ko
tana hoa wahine, a Marie-Claude, “Ng~ H§tori M~ori o
ng~ Carabidae” mÇ taua takiw~ anÇ (2003), t‘tahi r~rangi
o ng~ Carabidae (2001) me ng~ Heteroptera (2004) o
Aotearoa. Ko te aronga nui o ~na rangahau i t‘nei w~, ko te
America north of Mexico”. With his wife, Marie-Claude, ~hua me te whakarÇpãtanga o ng~ p§tara noho papa o
he published “A Natural History of Carabidae” for the Aotearoa. Ko t‘tahi w~hanga nui o ‘nei, ko tana pukapuka
same region (2003) as well as a catalogue of New Zealand ~whina i te tautohunga o ng~ iwi me ng~ puninga Carabidae
Carabidae (2001) and Heteroptera (2004). His current main o Aotearoa, taihoa nei ka puta.
research interest is the faunistics and taxonomy of New
Zealand ground beetles, which involves a soon-to-be-pub-
lished identification guide to the tribes and genera of
Carabidae from New Zealand. I wh~nau mai a Marie-Claude Larivière i Québec. I
reira anÇ ia e rapu ana i te m~tauranga ~, riro noa i a ia tana
Contributor Marie-Claude Larivière was born and edu- Tohu T~kutatanga mai i te Whare W~nanga o McGill, i te
cated in Québec, graduating with a Ph.D. in systematic tau 1990, ko ng~ whakapapa pepeke te kaupapa. MÇ te
entomology from McGill University in 1990. For the fol- rua tau i muri mai, kei Agriculture Canada, i Ottawa, ia e
lowing 2 years she did postdoctoral research at Agriculture wh~wh~ ana i ‘tahi atu rangahautanga. I te tau 1992, ka
Canada, Ottawa. In 1992, Marie-Claude moved to New neke mai a Marie-Claude ki Aotearoa, ka mahi hei kait~tai
Zealand to work as a full-time Hemiptera biosystematist i ng~ whakapapa o ng~i Hemiptera i Manaaki Whenua.
with Landcare Research. From 1994 to 1997 she led the N~na i ~rahi Te T~tainga o ng~ Whakapapa o ng~ Aitanga
Biosystematics of New Zealand Land Invertebrates pro- Tuar~-Kore a T~ne mai i te tau 1994 ki te 1997, me te
gramme, and from 1999 to 2004, the Koiora-BioAssist™ kaupapa Koiora-BioAssist™ (Te Aromatawai i ng~ Koiora
project (Biodiversity Assessment using Information Tech- i runga i te Whakamahi i te Hangarau MÇhiohio me te
nology and Taxonomy). Marie-Claude is the author of over WhakarÇpãtanga) mai i te tau 1999 ki te 2004. He neke
70 papers and monographs on the taxonomy, distribution, atu i te 70 ng~ tuhinga kua puta i a ia e p~ ana ki te
(continued overleaf) (haere tonu)
Fauna of New Zealand 53 9
and natural history of Hemiptera and Carabidae whakarÇpãtanga, te tohanga me te h§tori m~ori o ng~ Hemi-
(Coleoptera), including four Fauna of New Zealand contri- ptera me ng~ Carabidae (Coleoptera), tae atu ki ‘tahi
butions (Hemiptera—Cixiidae and Pentatomoidea revisions, putanga e wh~ o Te Aitanga Pepeke o Aotearoa. He tuhinga
catalogues—Carabidae and Heteroptera ). She has also anÇ kua puta i a ia mÇ ng~ Orthoptera me ng~ Carabidae o
published on North American Orthoptera and Carabidae. Amerika ki te Raki. Kua mahi tahi anÇ r~ua ko tana hoa
Many of her publications were written in collaboration t~ne, a André, ki te whakaputa i ng~ tuhinga huhua. Ko te
with her husband André with whom she hopes to soon tãmanako, taihoa ka puta i a r~ua he tuhinga hou mÇ ng~
publish new works on New Zealand Hemiptera and Hemiptera me ng~ Carabidae o Aotearoa. Kei te ng~kaunui
Carabidae. Marie-Claude has a keen interest in biodiversity anÇ a Marie-Claude ki te p~rongo-koiora, tae atu ki te
informatics, especially digital taxonomy, computer imaging, whakarÇpãtanga ~-mati, te t~rai whakaahua ki te rorohiko,
interactive identification, and web-publishing. te tautohu i runga i te mahi p~hekoheko, me te p~nui
kÇrero ki te pae tukutuku.
Translation by H. Jacob
T~maki-makau-rau / Auckland
DEDICATION
“Think where man’s glory most begins and ends
And say my glory was I had such friends”
W. B. Yeats 1865-1939: The Municipal Gallery Re-visited (1939)
We are glad to dedicate this revision to our colleague Barry P. Moore (Research Associate, Australian
National Insect Collection, Canberra) in acknowledgement of his continued friendship towards New Zea-
land coleopterists and his generously provided expertise on New Zealand carabids. Over the years Barry
has kindly identified ground-beetles for the New Zealand Arthropod Collection and private collectors, at a
time when a large proportion of the fauna remained undescribed. His publications on the carabids of New
Zealand (e.g., 1980, Anillina; 1996, Haplanister crypticus) and Australia (e.g., 1987, Australian catalogue)
have provided a solid foundation for our New Zealand catalogue (Larochelle & Larivière, 2001) and future
taxonomic revisions. In the preparation of the Harpalini revision, Barry has generously given us much
encouragement and support in the identification of adventive species as well as useful comments on
some difficult taxonomic problems.
10 Larochelle & Larivière (2005): Harpalini (Insecta: Coleoptera: Carabidae)
Frontispiece: Triplosarus novaezelandiae (Laporte de
Castelnau, 1867) (photograph prepared by M.-C. Larivière,
Landcare Research)