Gastropods from the Burdekin Formation, Middle Devonian, north Queensland

13 Pages·1997·5.1 MB·English

1
GASTROPODS FROM THEBURDEKIN FORMATION, MIDDLE DEVONIAN, NORTH
QUEENSLAND
ALEX G. COOK
Cook, Alex G. 1997 06 30: Gastropods from the Burdckin Formation, Middle Devonian,
north Queensland Memoirs ofthe Queensland Museum 42(1): 37-49. Brisbane. ISSN'
0079-8835.
Tweni) onelaxaofgastropodsaredescribedfromtheMiddleDevonianBurdckinFormation.
northQueensland.NewamongstthesearcBurdekinostamaburdekhunensisgen.et. sp.nov.,
EwyzoHfbHfdekinettsissp, now AiiotnphQltisp&jeUisp.nov., Didytnalgiabarthofomai$zn<
et sp. now. Murchisonia(MurchisoniaIjackjelli sp. nov., Pafaeozygopleuramaeheniyi sp.
nov., and Trinema heuleckerigen. ei sp. nov. The fauna contains typical Old World Realm
geneta but is Specifically StrOllgl> endemic. Gastropods, Devonian, Queensland, Burdc-
kin Formation
Me\- G Cook. Queensland Museum. PO Box 3300, South Brisbane, Queensland 4101,
Australia; 10March 1997,
Burdekin Formation is a shallow marine car- SYSTEMATIC PALAEONTOLOGY
bonate sequence exposed within the Burdekin
Subprovince (Wyatt & Jell, 1967; Henderson, PhyluGmAMSOTLRLOUPSOCDAA
Class
1980; ) The formation consists of. inter alia, a OrderARCHAEOGASTROPODA
reelal and biostromal complex, with associated Super-family BELLEROPHONTOIDEA
lagoonal. intcr-recf and offshore facies (Cook. Family BELLEROPHONTIDAEMcCoy. 185
Subfamily TROPIDODISCINAE Knight, 1956
1995). It contains diverse fossil assemblages
which include rugose corals (Zhen 1991, 1994), TropidodiscusMeek & Worthen, 1866
tabulate corals, slromaioporoids (Cook, 1994), TropidodLscus sp.
sponges, molluscs (Cook, 1993a) and conodonts (Fig. 1A-C)
(Talent & Mawson, 1994). Large, thick-shelled MATERIAL F.XAMINED QMF355I3.QMF35514.
gastropods from the underlying and interdigitat-
DESCRIPTION. Shell minute, approximately
ing, siliciclastie-dominated. Big Bend Arkose 2.5mm to diameter, 0.8mm wide; involute with
and basal units of the Burdekin Formation are moderately wide umbilicus. Whorl profile
Bnrdikinia hurdekinensis (Etheridge, 1917), sharply angular, withoutobvious crest. Relics of
A/nphelissa emtnatum (Heidecker, 1959), fine, numerous growth lines present.
Lahrocuspis nodosa Heidecker. 1959, and
Belleruphon de Montfort, 1810
Fleicheniewia septafa Cook, 1993b-
This paper is concerned with a diverse gastro- Betlcrophnn iBellerophon)de Montfort, 1810
pod faunule from a stratigraphically higher Bellerophon (Bellerophon) sp. A
unil in the Burdekin Formation in coralline Fig. 1D.E)
i
packstones interpreted to represent shallow- MATERIAL EXAMINED. QMF34916,QMF34S
water coralline thickets seaward of a biohermal
structure (Cook, 1995). The fauna has been DESCRIPTION. Small to medium-sized, spher-
silicilicd and was recovered by hu!k dissolution tcal. involute. 5.5mm wide; narrow deep umbili-
cus. Whorl profile evenly rounded, margin
in acetic acid. Both large and small gastropods bearing weak crest. Aperture unknown. Orna-
were retrieved, in addition tocorals, brachiopods ment consistsofmany fine growth lines.
aQnMdLb1i09v4a.lvneeasr.LiAtltlleRmoactkesr.i2a7lkmcoNmWesofCfhraorm- cRhEarMacAtRerKiSst.icsThteo ftehwe stpyepec,imBe.n(sB.p)osvsaessuslistiemsildaer
ters Towers. Montfort, 1810 from the Middle Devonian of
38 MEMOIRS OFTHEQUEENSLAND MUSEUM
Germany as figured by Knight (1941). The nu- SuborderPLEUROTOMARIINA
merousgrowthlinesandoverallformaresimilar, Cox & Knight, 1960
but the Burdekin form is much smaller (Knight, SuperfamilyPLEUROTOMARIOIDEA
1941: pi. 11) and the selenizone-bearing crest is Swainson, 1840
weaker, possibly due to imperfect silicification. Family RAPHIOSTOMATIDAE Koken, 1986
Subfamily OPHILETINAE Knight, 1956
Bellerophon (Bellerophon) sp. B
(Fig. 1F,G) Burdekinostoma gen. nov.
MATERIALEXAMINED.QMF34908. TYPE SPECIES. Burdekinostoma burdekinensis sp.
nov.
DESCRIPTION. Spherical, medium-sized, ap-
proximately IOmm in diameter, 9mmhigh; invo- ETYMOLOGY. Forthe Burdekin River,andalluding
lute with deep, wide umbilicus. Whorl profile tothe Raphiostomatidae.
wideand rounded with weakcrest; slight flatten-
ingofinnerwhorl surfaceadjacenttopenultimate DIAGNOSIS. Minute, low-spired to lenticular,
whorl, extending approximately one third across gradate shell, widely phaneromphalous, havinga
the inner whorl profile. commonly channeled suture and prominent,
gently inclined, flattenedareaonthe upperwhorl
REMARKS. The specimen is larger and has a surface bordered by 2 strong cords, inferred to
wider whorl profile than Bellerophon contain the selenizone and also commonly con-
(Bellerophon) sp. A, which is a result of the taining weakthreads; midwhorl profilegenerally
flattened area on the whorl profile abutting the subrounded but stepped with several cords.
penultimate whorl.
REMARXS. Placementofthis striking ifminute
Superfamily EUOMPHALOIDEA genus is difficult, as it superficially resembles
members of the Ophiletinae, Raphiosomatinae
deKoninck, 1881
Family EUOMPHALIDAEde Koninck, 1881 and Euomphalidae. Placement depends on the
interpretation oftheinclinedsurfaceon the upper
whorl face, which is inferred to contain the
Straparollus de Montfort, 1810 selenizone. In particular the raphiostomine
Straparollus (Euomphalus) Sowerby 1814 Wisconsonella Blodgett (1988) from the Eifelian
ofNorthAmericaisa similartaxonto the Burde-
Straparollus (Euomphalus) sp. kin material, sharing the gradate shell form, but
(Fig. 1Q) lacking the additional cords and the channeled
suture. Placement within the Raphiostominae is
MATERIALEXAMINED. QMF34965. restricted by the widely phaneromphalus nature
of this snail. Alternatively the channeled suture
DESCRIPTION. Medium-sized, planispiral, may be interpreted as containing a sinus, which
21.4mm wide, 8.2mm high. Suture in a channel. would ally the genus to the Euomphalidae, of
Whorl profile square with prominent angulation whichPoleumitaClarke& Ruedemann and Cen-
above and below gently convex midwhorl sur- tifugus Bronn are similar cord-bearing forms.
face. Weak spiral line adumbilically to the upper Neitheroftheseplacementsissatisfactory.Place-
angulation which may mark the position of the mentintheOphiletinaebestsatisfiestheinterpre-
sinus. Aperturerounded, butquadrate. Base flat- tation of the selenizone, accomodates the
tened. channelledsutureandthephaneromphalouscon-
dition.
Straparollus (Serpulospira) Cossman, 1916
Straparollus (Serpulospira) sp. Burdekinostoma burdekinensis sp. nov.
(Fig. 1R)
(Fig. 2J-P)
MATERIALEXAMINED. QMF34966.
MATERIAL EXAMINED. HOLOTYPE: QMF34961.
PARATYPES: QMF35522-QMF35530.
DESCRIPTION. Shell planispiral, disjunct,
13.6mm wide, 4.6mm high. Whorl profile ETYMOLOGY. Fromthe Burdekin River.
rounded, aperture rounded. No growth lines pre-
served. DIAGNOSIS. As forgenus.
GASTROPODS FROMTHE BURDEKIN FORMATION 39
^M:
Q
M
>
FIG. 1. A-C. Tropidodiscus sp. SEM images. A,B, QMF35512. side and oblique views respectively, x 16; C,
QMF355I3, side view, x 16. D-E. Bellerophon (Bellerophon) sp. A. QMF34916, apertural and side views
respectively.F,G.Bellerophon(Bellerophon)sp. B.QMF34908,sideandbrokenapertural viewsrespectively,
H-P,Euryzoneburdekinensissp. nov. H-J,HolotypeQMF34917,apicalaperturalandbasal viewsrespectively,
x 4,4; K,L, Paratype QMF 34969, basal and apical views, x 3.2; M,N, Paratype QMF34964 apical andbasal
views x 6.3: O, Paratype. QMF35536 oblique side view, x 22.5, SEM image; P Paratype QMF35536 apical
view x 22.5, SEM image. Q, Straparollus(Euomphalus) sp. QMF34965 apical view, x 1.8. R, Straparollus
(Serpidospira)sp. QMF34966, apical view x 2.3.
J
40 MEMOIRS OFTHEQUEENSLAND MUSEUM
D1E.S5CmRmIwPiTdIeOaNn.dS1h.e5llmsmmahlilghl,omvienruytel,owl-esspsitrehda,n TsApB.LnoEv.i , Measurements for Eury&nebtta4vkiwir$i3
-mewhat gradate, widely phaneromphalous,
with deep umbilicus. Whorl profile series ofan- Specimen Hcieht(nwn) Width(mm)
gularsurfacesbutgiving an overall rounded pro- Holoivpe 16 1,1 3
tile. Suture with adjacent, wide channel. Upper QMI 34909 6 ,10
whorl face bears numerous spiral threads, of at QMF34864 10 4.8
' 2 orders of intensity. Three prominent OMP349IS 3J 7.3
threads on smaller specimens; bordering the su- QMKM921 1 6 5.0
tural channel, another high on the whorl profile
and another at the edge of the upper whorl face the Middle Devonian Siringocephalus Lime-
giving way to the steepermidwhor! surfaces, the
stone. Germany, has a turbiniform shell and a
last 2 inferred to contain the selenizone. At least
widerselenizone.
2 threads on the midwhorl, creating a vertical
surface at the midwhorl periphery; at least 3
ruepvpoelrviwnhgorclordfsacoenbetaherslouwpertowh7ormlinfoarce.spTirhael SuSpuebrofradmeirlyTRAONCOHMIPNHAALCoOxI&DEKAniWghetn,z,| 1938
threads, in addition to the major cords. Family ANOMPHALIDAE Wenz. 1938
REMARKS. This striking taxon is easily differ- Anomphalus Meek & Wonhen, 1H67
entiated by Enrvzone burdektnensis sp. nov. by
the numerous spiral threads and more angular TYPESPEC1ES Anomphalusrotuius IroniiheMiddle
whorl profile. Perms!yvanian,SiDavidsLimestone.Illinois,USA.by
originaldesignation.
Family GOSSELETIN1DAE Wenz, 1938
Subfamily COELOZONINAE Knight, 1956 Anomphalus pajelJi sp nov.
(Fig.2A-J)
Euryzone Koken, 1896
Euryzone burdekininesissp. nov. MATERIAL EXAMINED. HOLOTYPE: QMF34nsh
(Fig. IH-P) PARATYPES:QMF34S51-QMF34S5S.
MATERIAL EXAMINED HOLOTYPE: QMFS4917. ETYMOLOGY. ForPeterA.Jell.
PARATYPES: QMF34918-QMF34927.QMF34964.
QMF34969. DIAGNOSIS. Smooth-shelled, medium-sued
DIAGNOSIS. Small to minute memberofgenus member of genus with very weak sutuics
withsinus on upper whorl profilebouded by fine prominentthickeningofinnerlipwhichcreates a
hemiomphalus base.
threads,
DESCRIPTION, Shelldiscoidal,minuteLosmall, DESCRIPTION- Medium-sized, low-spircd,
up to 11.9mm wide, 3.6mm high, generally rmomielliform hemmiommphalus gastropod, up to 1 1
smaller (Table 1). Whorls abut, with impressed high, 13.5 wide(Table 2). Sutures flush,
suture. Base with moderately deep umbilicus. commonly difficult to detect, whorls embrace
Whorl profile rounded, bearing two fine cords high on upper whorl surface. Umbilicus narrow
whichborderthesinus.Growthlinesfine,numer- obscured oroccluded by thickening at the innci
ous. Shell repair in evidenceon holotype. labrum, otherwise base rounded. Whorl profile
rounded with periphery at midwhorl. Apeiiurc
REMARKS. The species is characterised by its rounded. Shell thick, inner labrum particularly
discoidal shape and the striking presence or the thickened forming partial umbilical plug. Shell
sinus on the upper whorl face and its bordering smooth, very faint growth lines on the base oj
cords. The most similar memberofthe genus to QMF3485 1. There is some variation in the taxon
the Burdekin material is Eu/yzone penliwmom with respect to where the whorls embrace. On
Linsley 1968 from the Middle Devonian An- QMF3485 the whorlsembracefurtherdowndie
1
dcrdon Limestone, Michigan, but its selenizone whorl face, producing a slightly more naticir. im
is slightly lower on the whorl profile, and it is a shell shape. Despite silica replacement the
pianispiral than the Burdekin species. The smooth shell has an almost polished appearance
type species E. delphinuloides(Schothcim) from on some specimens.
GASTROPODS FROMTHE BURDEKIN FORMATION 41
FIG, 2. A-I.Anomphaluspajellisp.nov. A-C,HolotypeQMF34850,apertural,apical andbasal views,x9.D-F,
Paraiype QMF34852. apertural, basal andapical views, x 2.2. CM, Paratype QMF34851 apical, apertural and
basalViews, x 2.5. J-P, Burdekinostomaburdekinensis gen. et sp. nov. J,K, Paratype QMF35523. basal and
apicalviews,SEMimages.xl3.5;L,M.ParatypeQMF35522,obliqueaperturalandapicalviews,SEMimages,
x 15; N,QMF35525 side view, x 13.5; O.P. Holotype QMF34961, x 5.4, apical andside viewsrespectively.
.
42 MEMOIRS OFTHEQUEENSLAND MUSEUM
I'ABLE 2. Measurm«,nts for Anomphalux pajrlli sp from the Middle Devonian at Lummaton, En-
nov gland, but the (alter possesses strong collabral
, --.{M,,l_. i'Ml!L.-',ll! 1 Heigh10t.(7mm) Widt1h3.(1mm) 1 csMopirddd(sRloeollniDneesav,rolEnylidwarhniodrglMesa.r&cNeaSlptikulosleprsF,iors1m9af7ft1Ni)ao,tnfi,rCooNpmseitwsh}e
QMH34H51 11.5 l?.5 York, shows a similarornaIment, but the threads
QMF34852 7.8 13.1 are Hue and inclined on the whorl 1.
QMF34S53 7.8 1 1 ; Straparollus vorrugatas (Stauffer. 1909), which
Linsley (1968] placed wuhm llsonema, and Rol
REMARKS. Both Knight (1941 and Knight et lins. Eldridge & Spiller (1971) assigned 10
)
al. (I960) noted the variable umbilicus in this Natiropsis (Naticopsish fromtheMiddleDevon
genus. The Burdekin materia! is assigned to the ianofOhioalsoshowsstrongcollabral ornament
genus on the basis ofgross shape, lack oforna- but not restricted tothe mature whorls. yisontnui
ment and the hemiomphalus condition. The type cotrugatus (Stauffer) from the Middle Dew
ics A rvtulus Meek & VVonhen as figured by Anderdon Limestone (Linsley, 19680 is a mi-
Knight (1941) has a slightly lower whorl profile nutely illustrated taxon. Linsley (1968) dcsn
and more prominently incised sutures. An- the early whorls as smoothsuggesting affinity to
wnphalusumbilicoliratus Batten(1966) fromthe the Burdekinspecimen. Positive generic identifi-
Lower Carboniferous HotweLls Limestone, En- cation of Linsley*s taxon is impossible withuut
gland is clearly phaneromphalous and has spiral viewing his material. 1 place Naiica nexicosta
ornament on the base. Atwmphalus (Philips) of Whidborne and Straparollus
helicinaeformes (Schlotheim) from the Middle corrugatus Stauffer both wiihin NaticapsU
onian Stringocephalus Limestones near fNatiropsis)alongside thisdistinct, ifpoorly pre-
Gladbach,Germanyhasa fullydevelopedumbil- served and unresolved, taxon from the Buidekin
ical plug (see Knight 1941 pi 63). but in overall Formation,
:
aspect is most similar to the Burdekin species,
and is closely related. Specimens illustrated by SuperfamilvORIOSTONIATOIDEA Wenz. I
Knight (1941) show mutOf variation in spire Family ORIOSTOMATIDAE Wenz. 1938
height for A. helicanaefonni:* (Schlotheim).
NaticopsU margheriti Mansuy (1912), may also Didymalgia gen nov.
natiIeHitAbnrommsphehlal.hi; albeit with a Slightly more TYPESPECIES. Dutvmalgiabartholamaisp. nov
ETYMOLOGY
For'Didymalgia' hill,informal (Utile
SuborderNERITOPSINAECox & Knight, 1960 oflateriteprofilenearthe Flctchcrviow locality
Superfamily NERUOIDEA Rahnesquc, 1815
Family NERITOPSIDAE Gray, 1847 Didymalgia bartholomai gen. et sp. nov
(Fig 3)
NaticopsisM'Coy, 1844
MATER!ALEXAMINED. HOLOTYPB QMF34HW.
PARATYPES: QMF34876-QMF34895, QMF349I5
Naticopsis iNaticopsis) sp
Fig.4A) ETYMOLOGY. ForAlan Bartholomai.
•
MATERIALEXAMINED.QMF349 DIAGNOSIS. Turbiniform, widely phan-
13
eromphalous shell with numerous revolvinu
DESCRIPTION. Very small, 4.9mm wide, 5mm cords and strong comarginal cords forming and
high, natieitbrm, anompbalous; suture adpressed caneellate structure, whorl profile angular,
toslightlyimpressed;whorl profilerounded, with formed by series ofinclined surfaces.
periphery well above midwhorl. LFpper whorl
surface flatterthat lowerwhorl face. Final whorl DESCRIPTION. Medium-sized, lurhimtorm,
dominated by strong collabral cords, increasing widely phaneromphalus shell, up to 9.5mm high
mm
in intensity towards the aperture. Early whorls and 11.1 wide (Table 3). Whorl face adorned
lack this ornament. Parietal and inner lip heavily with strong revolving cords and slightly less
thickened; aperture ovate Base rounded. strong comarginal ribs, producing a caneellate
shell decoration. Whorl profile series ofangu-
REMARKS. The specimen strongly resembles lar surfaces with a subsutural shelf, steeply
Xaticanexicosta(Phillips)ofWhidborne(1892), inclined upper whorl \\\cq, angular periphery.
:
GASTROPODS FROMTHE BURDEKIN FORMATION 43
1
- Lr:r sg
;:>
FIG, 3. Didymalgiahanholomaigen. etsp. nov. A-D, Holotype QMF34896, x 3 apertural, basal (x2.8), apical
andside views(x 2.8). E-G, Paratype QMF34875. E, apertural viewx 5; F, apical views x5; G.side view x4.
I,J, Paratypes QMF34K76, x 3 apical andside views.
inclined lower surface leading to the phan- ostomatidae is problematic, in that it extends the
eromphalous base, sutures embrace just below range of the family beyond their supposed
periphery which is situated slightly below mid- Eifelian demise (Blodgett et al. 1990), but the
whorl. Slight subsutural shelf, 4 majorrevolving strong ornament, lack of selenizone on the
cordson the upperwhorl face, amajorcordatthe whorl faceandtrochiform,phaneromphalusshell
periphery, and 7 to 8 cords on the lower whor! form make the placement appropriate, ifcontro-
face and base. Two orders ofcords on the lower versial.
whorl face. Aperture rounded to subrounded,
slightly widerthan high.Thickcolabral cords are
numerous almost orthocline, and are not de-
flected by asinus orselenizone,
TABLE3.DataforDidymalgiabartholomaigen.etsp.
REMARKS. Oriostoma sp. aff O. gerbaulti nov.
&
Oehlert of Blodgett Johnson (1992) from the
EifelianofNevadaisgrosslysimilartothe Burde- Specimen Height(mm) width(mm)
kin material but has achanneled subsuturalzone. QMF34896 9.5 1LI
Gyronemamuhinodosa Blodgett&Johnson from QMF34S76 s.s 8 8
the Eifelian of Nevada, which could be QMF34S75 ; 8,(
accomodated in Kitikamispira, has fewer cords
and more prominent nodes on the shell surface.
Placement of the genus within the Ori-
1
MEMOIRS OFTHEQUEENSLAND MUSEUM
SuborderMURCHISONTINACox & Knight, >960 Murchisonia (Murch«>oni;0 -
Superfamily MURCHISONlOIDEAKoken. 18% (Fig. 41-Jj
Family MURCHISONIIDAE KOKEN, S96
I MATERIALEXAMINED.QMP3493Q
Vlurchisonia d*Archaic & de Vcrncuil 184 DESCRIPTION. Small, 10.5mm high, 5.3mm
wide, high-spired, with eight whorls present. Su-
MurehLsonia (Vlurchisonia)jackjelli sp. rtov. ture strongly impressed, whorls embrace well
(Fig. 4B-H) below midwhorl, angular whorl profile with 2
prominent spiral cords bordering a selenizone.
MATERJAL EXAMINED. HOLOTYPE: QMF34905. Otherdecoration not preserved.
PARATYFfiS: QMF34897-QMF34900, QMF349G2,
(JMF34903, QMF34929, QMF34932, QMF34933, REMARKS. Aselenizone on the midwhorl, bor-
QMF34935-QMF34940, QMF34942-QMF34943,
dered by 2 cords is theclassic murefvisoniid
J4946^MF34948, QMF34952-QMF34953.
dition and the specimen is here allocated to the
LI YMOLOGY. ForJ.S.Jell. subgenuson this basis. The species resemblesA7
(M )fermioni Tassel!, 1982. from the Emsian of
DIAGNOSIS. Medium-sized member of subge- New South Wales, and also known from (he
nuswitharelatively wideselenizone, subrounded Givetian in the Broken River Province, but the
I profile and up to 5 threads on Ihe adult material is loo poorly preserved to be confident
lower whorl surface. ofany specific assignment
DESCRIPTION. Medium-sized to large, fugh- Vlurchisonia iMurchisomul sp H
Spiredshell, up to41.5mm longand 1 1.6mtn wide ,Fig.4K>
01 base. Suture impressed, whorls embrace at
approximately lower one third ot the whorl MATERIALEXAMINED.QMF35520.
periplhue.rWahlocrolrdpsrobfoilredesruibnrgoaunwdieddebsuerlewniitzho2new.eUapk DESCRIPTION. Shell small, 6mm high, 2mm
basal width, high-spired, with 7 whorls present.
to 5 additional weaker spiral threadson the lower
Sutures impressed, whorls embrace just below
whorl face with 2 extreme!) faint threads on the
periphery. Whorl profile angular with 2 strong
upper surface. In juvenile specimens cords bor-
dering the selenizone are stronger on early cords borderingaselenizone at the periphery,the
upper cord slightly stronger. Upper whorl face
whorls, and the lower whorl face lacks the multi-
gently sloped with a weak spiral cord closetothe
ptlhereatdhsreaardesmoofrethperoamduilntenwth.orAlpse,rtiunrseteoavdat1e worith2 main ornament on the margin. Lower whorl pro-
minorthickening ofthe inner lip. fileobscured, aperture and ba.se unknown
RSiEgnMiAfiRcaKnStl.y iPmopoerdepsresiehrevasttiuodny ooff tthhies msapteecriieasl. tRhEeMsuAbRgeKnSus..TIthies hbiogrhdeerr-esdpirseedletnhiaznonMeuriddeuntsiofimeos
;rly The presence of the wide midwhorl ifMeurrecn!tusownhioar)lsp,pAr.ofainldehasthaasnignMiufricchatmshonfjiila
seienizonc denotesthe subgenus, and the numer-
ous weak threads on adult whorls distinguish a (Mttrchisanta)jackjelli sp. nov.
new taxon. The width of the seleni/one and the Superfamily CRASPEDOSTOMIDOIDEA
numerous threads may suggest Carboniferous
Wenz, 1938
vcaelia, buttheselenizone isat the periphery Family CODONOCHEILIDAE Miller. lUtS
negating that assignment. In the Middle Devon-
ian Murchisonia (Muniusonia) underwent sig-
MiichelliadeKonuuk.
nificantradiationintoamultitudeofunusual shell 18
forms (Knight et al. I960; Blodgeti et al., 1990)
and the included species described to dale are ?Mitche11ia ftp
numerous (e.g., Whidborne, 1892). Taxa with (Fig. 4L..MI
s(iMm.i)larvmiuclatripilaentahreWahdsidonbotrhenewh,orl fMa.ceare(.A/M.. MATERIALEXAMINED. QMF35515. QMF35516.
ioKunemoides Whidborne, both from the Middle) DESCRIPTION. Two fragmcntal specimens,
Devonian of Southern England, but the former neitherofwhichhaveagerontic Stagsoraperture
has nodose threads andthe latterathinner, lower preserved. Small high-spired gastropod with im-
set selenizone. pressed suture; whorls embrace josr below mid-
GASTROPODS FROM THE BURDEKJN FORMATION 45
FIG. 4. A, Naticopsis(Naticopsis) sp. QMF34913, oblique apical view, x 5. B-H, Murchisonia (Murchisonia)
jackjellisp. nov. B, C, Holotype QMF34905. side views x 2.8; D. Paratype QMF34900, side view x 1.5; E,
ParatypeQMF34897,sideviewx2;F.ParatypeQMF34903,sideviewx2.6;G,ParatypeQMF34899apertural
view x 1.6; H, Paratype QMF35520, side view xl2. IJ,Murchisonia(Murchisonia)sp. A. QMF34930, x3.5.
I, side view; J. apertural view. K Murchisonia {Murchisonia) sp. B, side view x 10. L, M, ?Mitchellia sp, L,
QMF35515, x 20; M, QMF35516 x 20.
whorl. Whorl profile rounded, face bearing nu- lian Emsian genus on the basis of gross shell
merous, (up to 10) spiral threads ofequal inten- shape. It lacks the characteristic aperture needed
sity. Aperture not preserved, but inner whorl formoredefinite assignmenttothe taxon.
profile ovate with long axis vertical.
REMARKS. Generic assignment is impossible
without more material, but the material is ques-
tionably assigned to the common eastern Auslra-
46 MEMOIRS OFTHEQUEENSLAND MUSEUM
Order CAENOGASTROPODACox, 1959 ETYMOLOGY. Tri-,three, nema(Greek)thread.
Superfamily LOXONOMATOIDEAKoken, 1889
Family PALAEOZYGOPLEURIDAEHorny 1955 DIAGNOSIS. Smalltomedium-sized,anompha-
lous,high-spired,manywhorledshell; whorlpro-
Palaeozygopleura Horny, 1955 file rounded, bearing three prominent midwhorl
cords. Suture impressed; a subsutural surface is
boundedbelow byaweakthread whichformsthe
Palaeozygopleura machenryi sp. nov upper limit of the deep sinus, in turn bounded
(Fig. 5A-F) belowbythehighestofthethreemidwhorlcords.
MATERIAL EXAMINED. HOLOTYPE; QMF35531. Finerthreadsarepresenton the lowerwhorl face.
PARATYPES: QMF35532-QMF35525,QMF35539. Growth lines are prosociine above the sinus and
opisthocline below. Whorls embracejust below
ETYMOLOGY. ForColin McHenry. thehigherofthe weakthreadsonthe lowerwhorl
face.
DIAGNOSIS. Small to minute memberofgenus
with moderately impressed suture and only REMARKS. Placement inthe familyis indicated
slightly sigmoidal ribs. by the overall shell form and the position of the
DESCRIPTION. Shell high-spired, many sinusabovethemidwhorlcords (see Knightetal.
whorled, up to 8 present, up to 4mm long and 1960: 1317). The taxon is differentiated by the
1.6mm wide. Sutures impressed, whorlsembrace AonclayntohtohenremDaevSohneirzaenrt&urrGirtealbiadu,(Li1n9s0l8eyby19l7a8c)k,-
well below midwhorl; whorl profile rounded. Pe-
ing the distinctive nodose cords. In the Late Car-
riphery at midwhorl. Whorl face bears many boniferous Orthonema Meek & Worthen, 1861
slightly sigmoidal ribs, obvious on all adult (seeKnight 1934, 1941,Knightetal., 1960) there
whorls.Protoconchpoorlypreserved,butsmooth
are 2 prominent midwhorl cords, widely sepa-
and dextral.
rated below the similarly positioned sinus. In
REMARKS. The material is most similar to P Callispira Nelson,1947, the sinus is also posi-
joanni Linsley from the Middle Devonian An- tioned high on the whorl face, but the whorl face
derdon LimestoneofMichigan,but in theBurde- is lessrounded and there are many more stronger
kin species the ribs are less numerous and the cords (5-6) on the whorl face. There is a consid-
suture less impressed. It differs from the single erable gap in the taxonomic record of the
sbpyetchiemmenorPealiameporzeysgsoepdlseuutruaressp.adnedsccorairbseedrboerlnoaw- ATucrarnittheolnleimdaaeinbtehtewEeaerlny Dtheevonaipapnea(rSahenrczeero&f
ment. Grabau, 1908), their Middle Devonian record
(herein)andthenumerousCarboniferousrecords
Palaeozygopleura sp. ofOrthonema(Knight, 1934). Suchadiscontinu-
(Fig 5G) ity in the record may be resolved by analysis of
Late Devonian age faunas.
MATERIALEXAMINED. QMF355
19.
Trinema heideckeri sp. nov.
DESCRIPTION. Minute, 3.6mm high, 1.5mm (Fig.5H-K)
wide, high-spired, nearly pupiform with shal-
lowly impressed suture, gently rounded whorl MATERIAL EXAMINED. HOLOTYPE: QMF34954.
profile. Ornament consists of fine, prosociine PARATYPES: QMF34941,QMF34945, QMF34955.
threads on the whorl surface.
DIAGNOSIS. As forgenus.
REMARKS. The single specimen has weaker
sutures than P. machenryi sp. nov, and finerpro- DESCRIPTION. Material somewhat fragmental.
sociine ornament. Shell small- to medium-sized, anomphalous,
high-spired, up to 12 whorls present. Whorl pro-
Family TURRITELLIDAEWoodward, 85 file rounded overall, slightly more convex on
1 1 lower whorl face; periphery at midwhorl. Suture
impressed with subsutural face bordered below
Trinema gen. nov.
by weak spiral thread. Midwhorl face dominated
TYPESPECIES. Trinemaheideckerisp.nov.fromthe by 3 strong spiral cords the middle of which is
Middle Devonian Burdekin Formation, north Queens- slightly weaker. Between the uppermost of the
land. strong peripheral cords and the weak cord below