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; Forest management options for carbon stock and soil reha- bilitation in Chilimo dry afro -montane forest, Ethiopia DOCTORAL THESIS Mehari Alebachew Tesfaye February, 2015 Palencia, Spain Forest management options for carbon stock and soil rehabilitation in Chilimo dry afro-montane forest, Ethiopia ESCUELA TĖCNICA SUPERIOR DE INGENIERĺAS AGRARIAS INSTITUTO UNIVERSITARIO DE INVESTIGACIȮN EN GESTIȮN FORESTAL SOSTENIBLE TESIS DOCTORAL: FOREST MANAGEMENT OPTIONS FOR CARBON STOCK AND SOIL REHABILITATION IN CHILIMO DRY AFRO-MONTANE FOREST, ETHIOPIA Presentada por Mehari Alebachew Tesfaye por optar al grado de doc- tor por la Universidad de Valladolid Dirigida por: Dr. Andrés Bravo Oviedo Dr. Felipe Bravo Oviedo Forest management options for carbon stock and soil rehabilitation in Chilimo dry afro-montane forest, Ethiopia Dedication This thesis is dedicated to my beloved children (Kidus Mehari and Yohanan Mehari) Tesfaye (2015) Table of contents Contents LIST OF ORIGINAL WORKS .................................................................................................................................. 1 ACKNOWLEDGEMENTS ...................................................................................................................................... 3 ABSTRACT ....................................................................................................................................................... 5 RESUMEN ........................................................................................................................................................ 9 1. INTRODUCTION ....................................................................................................................................... 13 1.1. African forest resources, the case of Ethiopia ....................................................................................... 13 1.1.1. Ethiopian forests ............................................................................................................................ 15 1.3. Carbon stock in tropical and Ethiopian forests ..................................................................................... 20 1.4. Chilimo dry afro-montane forest of Ethiopia ........................................................................................ 23 1.5. Forest management tools available for African tropical forests ........................................................... 26 2. OBJECTIVES OF THE THESIS ................................................................................................................. 27 2.1. Aims and specific objectives................................................................................................................. 27 2.2. Conceptual model ................................................................................................................................. 29 3. MATERIAL AND METHODS ........................................................................................................................... 33 3.1. Study sites ............................................................................................................................................. 33 3.2. Experimental design and sampling ....................................................................................................... 34 3.3. Data gathering ....................................................................................................................................... 35 3.3.1. Tree data ........................................................................................................................................ 35 3.3.2. Biomass data .................................................................................................................................. 37 3.3.3 Coarse wood debris and stump data collection .............................................................................. 38 3.3.4. Soil data (study I and study III) ..................................................................................................... 38 3.3.5. Wood density and carbon concentration (study V) ........................................................................ 39 3.4. Laboratory methods .............................................................................................................................. 40 3.4.1. Soil analysis ................................................................................................................................... 40 3.4.2. Wood density and carbon concentration........................................................................................ 41 3.5. Statistical analysis ................................................................................................................................. 42 3.5.1. Mixed models ................................................................................................................................. 42 3.5.2. General linear models .................................................................................................................... 43 3.5.3. R- software ..................................................................................................................................... 44 4. RESULTS ........................................................................................................................................................ 47 4.1. Performance and soil condition of selected species in degraded highlands of Ethiopia (study I) ........ 47 ii Forest management options for carbon stock and soil rehabilitation in Chilimo dry afro-montane forest, Ethiopia 4.2. Biomass model, partitioning and comparison (study II) ....................................................................... 54 4.3. SOC and SON concentration and bulk density along elevation, land use and species (study III) ........ 55 4.4. Stand density management model evaluation and fitting (study IV) .................................................... 57 4.5. Carbon concentration and wood density for five most commonly tree species (study V) .................... 58 5. DISCUSSION .................................................................................................................................................. 61 5.1. Performance and soil condition of selected species in degraded highlands of Ethiopia (study I) ........ 62 5.2. Biomass model, partitioning and comparison (study II) ....................................................................... 64 5.3. SOC and SON concentration and bulk density along elevation gradient, land use and species (study III) ................................................................................................................................................................ 65 5.4. Stand density management model evaluation and fitting (study IV) .................................................... 68 5.5. Carbon concentration and wood density for five most commonly tree species (study V) .................... 68 6. GENERAL CONCLUSIONS .............................................................................................................................. 73 6. CONCLUSIONES GENERALES ........................................................................................................................ 75 REFERENCES ..................................................................................................................................................... 77 Sas codes used for data analysis ................................................................................................................ 102 Study I ............................................................................................................................................................ 127 Selection of tree species and soil management for simultaneous fuelwood production and soil rehabilitation in the Ethiopian Central Highlands ................................................................................................................ 127 Abstract ...................................................................................................................................................... 127 Introduction ................................................................................................................................................ 128 Material and Methods ................................................................................................................................ 129 Results ........................................................................................................................................................ 135 Discussion................................................................................................................................................... 148 Conclusions ................................................................................................................................................ 152 Acknowledgements .................................................................................................................................... 153 References ................................................................................................................................................. 153 Study II ........................................................................................................................................................... 163 Aboveground biomass equations for sustainable production of fuel wood in a native dry tropical afro- montane forest of Ethiopia ............................................................................................................................ 163 Abstract ...................................................................................................................................................... 163 Introduction ................................................................................................................................................ 164 Material and Methods ................................................................................................................................ 166 Results ........................................................................................................................................................ 173 Discussion .................................................................................................................................................. 180 Tesfaye (2015) Conclusions ................................................................................................................................................ 182 Acknowledgements .................................................................................................................................... 183 References .................................................................................................................................................. 183 Study III .......................................................................................................................................................... 191 Impact of changes in land use, species and elevation on soil organic carbon and total nitrogen in Ethiopian Central Highlands ........................................................................................................................................... 191 Abstract ...................................................................................................................................................... 191 Introduction ................................................................................................................................................ 192 Material and Methods ................................................................................................................................ 194 Results ........................................................................................................................................................ 200 Discussion .................................................................................................................................................. 210 Conclusions ................................................................................................................................................ 214 Acknowledgements .................................................................................................................................... 214 References .................................................................................................................................................. 215 Study IV .......................................................................................................................................................... 223 Stand density management diagram for Chilimo dry afro-montane forest using species proportion in Central Highlands of Ethiopia ........................................................................................................................ 223 Abstract ...................................................................................................................................................... 223 Introduction ................................................................................................................................................ 224 Material and Methods ................................................................................................................................ 225 Results ........................................................................................................................................................ 230 Discussion .................................................................................................................................................. 239 Conclusions ................................................................................................................................................ 241 Acknowledgements .................................................................................................................................... 241 References .................................................................................................................................................. 242 Study V ........................................................................................................................................................... 247 Variation in carbon concentration and wood density for five most commonly grown tree species in Ethiopia ....................................................................................................................................................................... 247 Abstract ...................................................................................................................................................... 247 Introduction ................................................................................................................................................ 248 Material and Methods ................................................................................................................................ 250 Results ........................................................................................................................................................ 253 Discussion .................................................................................................................................................. 262 Conclusions ................................................................................................................................................ 264 iv Forest management options for carbon stock and soil rehabilitation in Chilimo dry afro-montane forest, Ethiopia Acknowledgements .................................................................................................................................... 265 References .................................................................................................................................................. 265 Photographs ................................................................................................................................................... 271 Forest management options for carbon stock and soil rehabilitation in Chilimo dry afro-montane forest, Ethiopia LIST OF ORIGINAL WORKS This thesis is based on five original works which are presented in the text with Roman nu- merals (I, II, III, IV, V) i.e. one published, one accepted with major revision, one submitted and two manuscripts. I. Tesfaye, A. Mehari, Bravo-Oviedo, A., Bravo, F., Kidane, B., Bekele, K., Sertse, D., 2014. Selec- tion of tree species and soil management for simultaneous fuelwood production and soil rehabili- tation in the Ethiopian Central Highlands. Land Degradation & Development (2014). Doi: 10.1002/idr/.2268 (Wiley online library.com). II. Tesfaye, A. Mehari, Ruiz-Peinado, R., Bravo-Oviedo, A., Bravo, F., 2014. Aboveground bio- mass equations for sustainable production of fuelwood in a native dry tropical afro-montane for- est of Ethiopia. Submitted to Annals of Forest Science, Springer. III. Tesfaye, A. Mehari, Bravo, F., Ruiz-Peinado, R., Pando, V., Bravo-Oviedo, A., 2015. Impact of changes in land use, species and elevation on soil organic carbon and total nitrogen in Ethiopi- an Central Highlands. Accepted with major revision, Geoderma, Elsevier. IV. Tesfaye, A. Mehari, Bravo, F., Bravo-Oviedo, A., 2015. Stand density management diagram for Chilimo dry afro-montane forest using species proportion in Central Highlands of Ethiopia. Manuscript. V. Tesfaye, A. Mehari, Bravo-Oviedo, A., Bravo, F., Herrero, C., 2015. Variation in carbon con- centration and wood density for five most commonly grown tree species in Ethiopia. A paper presented on 28th and 29th of January 2015: IX young researchers meeting on conservation and sustainable use of forest systems 2015, Valsaín-Segovia-Spain (Abstract published). 1

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