Flora of Siberia, Vol. 9: Fabaceae (Leguminosae)

282 Pages·2006·57.014 MB·\282

by  L I Malyschev (Editor)

Flora of Siberia
Volume 9
Fabaceae (Leguminosae)
48 60 72 84 96 108 120 132 144 156
Ta Olenek Lena Ar Indigirka NolymKao
Yan 64
Ob Yenisey Pu Ol
Yam
Khm
Ob Vi Yakutsk
KR
KKUurganTyuImrTtyebsnhOOmMsk NOTO TKomemskerovoVPeYeonidseykamAennngaaTyranaTunguska Pr Nizh.Tunguska Vilyui Ka Al 56
Novosibirsk
Se
Barnaul KE Kha An
Ba Abakan Chita
Corno-Altavsk TU Irkutsk Ulan-Ude Shi 48
CO Kynzl Yuzh
72 96 108 120
(cid:1)(cid:1)(cid:1)(cid:1)(cid:2)(cid:2)(cid:2)(cid:2)(cid:3)(cid:3)(cid:3)(cid:3)(cid:4)(cid:4)(cid:4)(cid:4)(cid:5)(cid:5)(cid:5)(cid:5)(cid:6)(cid:6)(cid:6)(cid:6)(cid:7)(cid:7)(cid:7)(cid:7)(cid:5)(cid:5)(cid:5)(cid:5)(cid:8)(cid:8)(cid:8)(cid:8)(cid:9)(cid:9)(cid:9)(cid:9)(cid:10)(cid:10)(cid:10)(cid:10)(cid:11)(cid:11)(cid:11)(cid:11)(cid:12)(cid:12)(cid:12)(cid:12)(cid:5)(cid:5)(cid:5)(cid:5)(cid:3)(cid:3)(cid:3)(cid:3)(cid:13)(cid:13)(cid:13)(cid:13)(cid:6)(cid:6)(cid:6)(cid:6)(cid:9)(cid:9)(cid:9)(cid:9)(cid:3)(cid:3)(cid:3)(cid:3)(cid:14)(cid:14)(cid:14)(cid:14)(cid:9)(cid:9)(cid:9)(cid:9)(cid:15)(cid:15)(cid:15)(cid:15)(cid:5)(cid:5)(cid:5)(cid:5)(cid:16)(cid:16)(cid:16)(cid:16)(cid:11)(cid:11)(cid:11)(cid:11)(cid:4)(cid:4)(cid:4)(cid:4)(cid:5)(cid:5)(cid:5)(cid:5)(cid:17)(cid:17)(cid:17)(cid:17)
Flora of Siberia
Volume 9
Fabaceae (Leguminosae)
Principal Editors
A.V. Polozhij, Dr. Biol. Sci.
and
L.I. Malyschev, Dr. Biol. Sci.
Science Publishers
Enfield (NH) Jersey Plymouth
CRC Press
Taylor & Francis Group
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Boca Raton, FL 33487-2742
© 2006 by Taylor & Francis Group, LLC
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Preface
Leguminosae represents one of the largest groups of dicotyledons, the tax-
onomy of which continues to evoke discussion. Many taxonomists (Jussieu,
Engler, Prantl, and Takhtadzhyan) combine these plants into a single family
Fabaceae or Leguminosae, with 3 subfamilies: Mimosoideae, Caesal-
pinioideae, and Papilionatae or Faboideae. Some others regard it more
appropriate to treat Mimosoideae, Caesalpinioideae, and Papilionatae as
families.
Considering the significant morphological differences among members
of these 3 taxa and the isolation of their distribution ranges (mainly of
Papilionatae), the authors of Flora evropeiskoi chasti SSSR [Flora of Euro-
pean USSR] (No. 6, 1987) and Siberian botanists regard Papilionatae as a
family of Fabaceae (Leguminosae).
The species of Leguminosae (Fabaceae) are distributed in all the vegeta-
tive zones and belts of Siberia. They enter the composition of almost all
phytocoenoses, many of them dominating. As a result of symbiosis with
bacteria which synthesize atmospheric nitrogen, legumes enrich the soil
with nitrogen compounds and, unlike all other plants, accumulate mainly
proteins and amino acids as reserves.
This unique group of flowering plants plays an important role in the life
of plant communities and is of great practical importance. Many species are
valuable fodder plants and have been introduced in cultivation as edible
plants. Some members of Leguminosae contain alkaloids while some others
contain biologically active principles and find application in medicine.
The work is based on a study of all the data collected at the P.N. Krylov
Herbarium as well as the collections of the herbaria of the Botanical Insti-
tute of the Russian Academy of Sciences, Moscow State University, Central
Siberian Botanical Garden of the Russian Academy of Sciences, and some
specimens at the Herbarium of the Institute of Biology of the Russian Acad-
emy of Sciences, Yakutsk. Moreover, special field investigations were car-
ried out and additional material collected in Gorno-Altay and regions of
eastern Siberia.
Based on the investigations carried out in Siberia, 351 species and sub-
species of Leguminosae of 27 genera have been included, the most abun-
dant of them being Oxytropis (120 species) and Astragalus (96 species).
The authors have described 4 new species. Twenty species are being
cited for Siberia for the first time and 6 species, formerly treated as
vi
synonyms, have been restored to the rank of species. For each species, a
brief morphological description and information on ecology and geography
have been provided. Original data on chromosome numbers have been given
for some species to the extent available in the literature.
The compilers are grateful to T.V. Revushkina, M.V. Olonova, and O.
Timofeeva for preparing the original drawings and to G.A. Kopaneva for
assistance in plotting the maps.
A.V. Polozhij
* * *
The following abbreviations have been used in describing the diagnostic
features of plants:
auct. non…—auctores, non … (authors, not …)
class. hab.—classic habitat
comb. nova—combinatio nova (new combination)
diam.—diameter
f.—forma (form)
s.l.—sensu lato (in a broad sense)
sp.—species
s. str.—sensu stricto (in a narrow sense)
subsp. —subspecies
syn.—synonym
var.—varietas (variety)
The distribution range of plants has been depicted in coded form for the
28 administrative divisions of the nominal floristic regions of Siberia (see
Fig. 1) as follows:
(cid:1)(cid:9)(cid:19)(cid:25)(cid:6)(cid:26)(cid:25)(cid:4)(cid:23)(cid:18).(cid:4)(cid:25)(cid:9)(cid:6)-(cid:10)(cid:19)6(cid:4)%(cid:9)(cid:26)(cid:4)(cid:19)(cid:6)#(cid:10)(cid:19)(cid:18)(cid:7)(cid:4)(cid:29)(cid:7)(cid:6)(cid:14)(cid:10)(cid:25)%(cid:10)(cid:17)(cid:4)(cid:14)(cid:26)6(cid:10)(cid:6)(cid:19)(cid:25)(cid:4)(cid:6)(cid:29)(cid:4)(cid:16)(cid:10)?(cid:26)(cid:14)(cid:10)(cid:18)(cid:2)
vii
West. Sib.—Western Siberia
TYU—Tyumensk Province
Yam—Yamal-Nenets Republic
Khm—Khanty-Mansi National District
Tb—Tobol Floristic Region
KU—Kurgan Province
OM—Omsk Province
TO—Tomsk Province
NO—Novosibirsk Province
KE—Kemerovo Province
AL—Altay Territory
Ba—Barnaul Floristic Region
Go—Gorno-Altay Republic
Cen. Sib.—Central Siberia
KR—Krasnoyarsk Territory
Ta—Taimyr Floristic Region
Pu—Putoran Floristic Region
Tn—Tunguska Floristic Region
Kha—Khakass Republic
Ve—Verkhneenisei (Upper Yenisey) Floristic Region
TU—Tuva Autonomous Soviet Socialist Republic
East. Sib.—Eastern Siberia
IR—Irkutsk Province
An—Angara-Sayan Floristic Region
Pr—Fore Lena-Katanga Floristic Region
BU—Buryat Autonomous Soviet Socialist Republic
Se—Severo-Buryat (Northern Buryat) Floristic Region
Yuzh—Yuzhno-Buryat (Southern Buryat) Floristic Region
ChI—Chitin Province
Ka—Kalar Floristic Region
Shi—Shilko-Argun Floristic Region (Dauria)
YAK—Sakha Republic (Yakut Autonomous Soviet Socialist Republic)
Ar—Arctic Floristic Region
Ol—Olenek-Nizhnelensk (Lower Lena) Floristic Region
Vi—Vilyui-Verkhnelensk (Upper Lena) Floristic Region
Al—Aldan Floristic Region
Ko—Kolyma Floristic Region
Yan—Yano-Indigirka Floristic Region
TThhiiss ppaaggee iinntteennttiioonnaallllyy lleefftt bbllaannkk
Contents
(cid:27)
Preface—A.V. Polozhij
8
Key to Genera—A.V. Polozhij
0
1. Sophora L.—V.I. Kurbatskij
2
2. Lespedeza Michaux—V.I. Kurbatskij
+
3. Sphaerophysa DC.—V.I. Kurbatskij
1
4. Caragana Lam.—V.I. Kurbatskij
80
5. Astragalus L.—S.N. Vydrina
<0
Subgenus Phaca (L.) Bunge
<0
Section Cenantrum Koch
<1
Section Komaroviella Gontsch.
0/
Section Orobella Gontsch.
0<
Section Hemiphragmium (Koch) Bunge
01
Section Hemiphaca Gontsch.
28
Section Melilotopsis Gontsch.
2<
Section Glycyphylla (Steven) Bunge
2<
Subgenus Astragalus
2<
Section Myobroma (Steven) Bunge
22
Subgenus Hypoglottis Bunge
22
Section Hypoglottioidei DC.
2:
Subgenus Trimeniaeus Bunge
2:
Section Cycloglottis Bunge
2+
Section Heterodontus Bunge
21
Subgenus Calycophysa Bunge
21
Section Alopecias (Steven) Bunge
21
Subgenus Cercidothrix Bunge
21
Section Euodmus Bunge
2,
Section Craccina (Steven) Bunge
:8
Section Onobrychium Bunge
:2
Section Ammodytes (Steven) Bunge
:2
Section Helmia Bunge
:+
Section Cystium Bunge
:1
Section Tanythrix Bunge