Five New Species and a New Subspecies of the Genus Phaedis (Coleoptera, Tenebrionidae) from Sulawesi, with a Key to the Sulawesian Species

2007·15.4 MB·English

by  Ando, K

Elytra,Tokyo,35(1):163-176,May30,2007
FiveNewSpeciesandaNewSubspeciesoftheGenusPhaedis
(Coleoptera, Tenebrionidae) fromSulawesi,
witha Key to theSulawesianSpecies
Kiyoshi AND0
Koffldai5-3-5,Toyono-cho,Toyono-gun,Osaka,563-0104Japan
Abstract Five new speciesand a new subspeciesof the genusPhaedis are
describedfromSulawesi. ThegenusMlcrogauromaiaPfc,1921isregardedasajunior
synonymofthegenusPhaedis,andMicrogauromaiaminutaPIc,1921istransferredto
thegenusPhaedis. Totallysevenspeciesandonesubspeciesofthegenusarerecorded
fromSulawesi,sixspeciesandonesubspeciesofthembeingbrachypterous. Akey to
theSulawesianspeciesisgiven. NewtaxadescribedhereinarePhaedismasataka1,P
altilis, P. ohbaJ,ashii, P nemorivagus, P. celebensisspp nov., and P. ovalipennis
inornatussubsp nov.
First ofall,I wouldliketodedicatethisshortpapertothememoryoftheexcellent
coleopterologist,thelateDr.MasatakaSAT0,inrecallingapleasanttimespentwithhim
andsomeother coleopteristsin therain forest ofSulawesi.
ThegenusPhaedlsbelongingtothetribeCnoda1onini isaconspicuousgenushaving
awelldeveloped frontalhalfofhead,broadened fronsanddistinctlyarmedprofemora.
ThisgenuswaserectedbyPAscoEin1866forPhaediselysiusfromBorneo. Sincethen,
fifty-ninespecieshavebeenknownfromawiderangeofSoutheastAsia. InSulawesi
Phaediswas representedbyonlyasinglespecies,P. ovalipennls KRAATz, 1880,which
belongstothebrachypterousgroupofthegenus. Inthecourseofrevisingthegenus,five
speciesandonenewsubspeciesarerecognized fromthisisland, fourspeciesanda
new
subspeciesof thembeingalsobrachypterous,andclosely related tooneanother. They
musthavedivergedintheislandinratheralateera. Ontheotherhand,myexamination
of PIc's typeof Microgauromaiaminuta PIc, 1921 fromCelebes deposited in the
Museumnationald'Histoirenature11e,Paris,provedthatthegenusMicrogauromalawas
ajuniorsynonymofthegenusPhaedis. Iwilldescribehereinfivenewspecies,onenew
subspeciesandonenewsynonym,withakeytotheSulawesianspeciesof thegenus.
Theabbreviationsand terminologyemployedhereinareas follows: IE- width of
dorsal interspacebetweeneyes; TD-transversediameter of aneye,measured from
dorsalaspect;PL- lengthofpronotum,measuredalongmedianline;PW-widthatthe
widest levelofpronotum;EL-medianlengthofelytra;EW-greatest widthofelytra.
Types and examined species used for the present paper are deposited in the
followinginstitutions:CKA0-Co11ection Kiyoshi ANDo,Osaka; CKMT-Co11ection
164 KiyoshiANDo
Dr. Kimio MAsUMoTo, Tokyo; CRGT-Co11ectionDr. RolandGRIMM, Tiibingen;
EUMJ -TheFaculty of Agriculture, EhimeUniversity; MNHN-Museum national
d'Histoire nature11e, Paris; SMNS-Staatliches Museum ftir Naturkunde, Stuttgart;
ZSM -Zoo1ogischeStaatssammlungMiinchen.
GenusPhaedisPAscoE, 1866
PhaedisPAscoE,1866,474.
PseudeumolpusKRAATz,1880,111.
Microgauro,naiaPIc,1921,23.Sy11nov.
Key totheSulawesianPhaedisSpeciesBasedon theMaleCharacteristics
Elytra with metallic coloured fasciae; hind wingssoundandelytrawithdistinct
shoulders.Bodylength:8.8-9.1mm P masataka1sp nov
Elytradevoidof fasciaeandhumeri; brachypterousspecies. 2
2 Presternal process horizontal or weakly sloping posteriad behind coxae never
adunc
Presternal processdistinctlyaduncbehindcoxae
Elytra fused witheach other; elytra1intervalsevenly convex; inner ocular sulci
distinct. Body length: 9.5mm P minutus (PIc)
Elytrafree;elytra1intervalsflatorfeeblyconvexinpart;innerocularsulciobscure
4
4 Pronotumwithanteriormarginthicklybordered;puncturesonheadandpronotum
verydenseanddistinct;bodymoreoval; innermarginsofmeso-andmetatibiae
stronglycurvedatbase;elytradevoidofstripes.Bodylength:7.5-9.5mm. __..
P. (P.)altiiissp nov
Pronotumwithanteriormarginnarrowlyandfinelybordered;puncturesonhead
andpronotumsparserandfiner;bodymoreelongate; innermarginsofmeso-and
metatibiaeweakly curvedatbase;eachelytronsometimeswithstripe. Length:
8.6-11.0mm
Elytra with stripes, humeral corners not angulate; punctures onpronotumand
elytra1intervalsfiner;headlongerinfrontalhalf;pronotumwithlateralmargins
distinctlysinuousbeforebase;parameressuddenlynarrowedinmedianthirdand
convergent again in apical fourth. Length: 9.l-11.0 mm
P. (P.)ovalipennisovalipennis (KRAATz)
Elytrawithoutstripes,humeralcornersobtuselyangulate;puncturesonpronotum
andelytra1intervalscoarser; headshorter infrontalhalf;pronotumwith lateral
margins hardly sinuousbefore base; parameres gently narrowed from baseto
apical fourth,thenstronglyconvergent toapex.Length:8.6-11.0mm
pl (P.)ovalipennislnornatussubsp
no v
6. Presternalprocessflabellateatapex;pronotumwithanteriormarginstraight; inner
New P/1aedls from Sulawesi 165
ocular sulci shallow; postgenae convex, with punctures coarse; body small
Length: 7.3-7.5mm. P. (P.)ohbayashiisp nov
Presternalprocessgentlyconvergentapicad;pronotumwithanteriormarginemar-
ginate; inner ocularsulci deep;postgenaenormal,withpunctures not coarse;
bodymedium tolargeinsize 7
7 Body more oblong; elytra greenish brassy, with strong reddish purple lustre,
moderately convex above, gradually co nvex from base tomiddle, elytranot
fused; hind wings almost as long as elytra11ength, flightless because wing
venation is reductive. Body length: 12.0-12.6mm
P. (P.) nemorivagussp
no v
Body more oval; elytrametallicblue or rarely metallicgreen, lustrous, entirely
fused with each other, extremely convexabove, stronglyandabruptly convex
frombase; hind wings vestigial. Length: 6.7-9.0mm
P. (P.)celebensissp
nov
Phaedis(Phaedis)masatakaisp
no v
(Figs 3,9,16,25,32&43)
Measurements. Bodylength:8.8-9.1mm;width:3.7-4.1mm.
Typeseries. Holotype: PuncakPalopo,S. Sulawesi, Indonesia, 2-I-2000, K.
,
AND0leg. (EUMJ).Paratype:1早,samedataasfor theholotype(CKA0).
Oblong, rather weakly convex above, brilliant dorsally. Colour dark reddish
brown, anteriorhalf ofheadmetallicblue-green,posteriorhalfof headandpronotum
metallicblue (male, holotype) or blue-violet (female, paratype); elytrametallicblue,
each elytron with apair of reddish purplehumeral andsubapical fasciae which are
thicklymarginedwithbrassy-yellow; venterwithmetallic reflection.
M ale. Head hexagonal, well produced in front of eyes, distinctly depressed
behind fronto-clypea1suture, withpuncturesminuteand dense,sparser on fronsand
coarseronvertex; clypeusalmost flat, veryslightlyemarginateat themiddleof apex;
fronto-clypea1suturedeep andclear, obscure laterally; genaealmost parallel-sidedat
sidesinbasalhalf,postgenaesuddenlynarrowedposteriad;eyescomparatively large,
roundlyproducedlaterad, inner ocularsulcidistinct,deeplyengraved; fronsflatand
sloping forwards, IE/TD=2.50 (female: 2.22). Antennaereachingbehindmiddleof
pronotum,fivedistalsegmentsformingamoderateclub. Terminalsegmentofmaxillary
palpus right-angled triangular. Mentum triangular, raised along middleand deeply
excavated at sides.
Pronotumtransverselyquadrate,evenlyborderedexceptforthemiddleofanterior
margin,widestbehindmiddleandPW/PL=1.50 (female:1.55);discconvex,sloping
posteriadinbasal fourth,withpuncturesdense,irregularlyarranged,larger thanthose
on head; anterior margin gently and arcuately emarginate; anterior angles obtuse,
weaklyproduced; lateralmarginssimply narrowed forwards from thewidest portion
and weakly sinuous in basal fourth, narrowlybordered; basal margin bisinuous, nar-
166 KiyoshiANDo
rowlybordered.
Elytraoblong, gently convex,moderately roundedatsides, widest behindmiddle
andEL/EW=1.53;humeralcallidistinctlytumid;striaemoderatelyimpressed,becom-
ingdeeper laterad,stria1punctures ratherminute,spacebetween themabout 15 to7
timesaswideasapuncture;intervalsweaklyconvex,distinctlysoinapico-1atera1areas,
minutelyandmoderatelypunctate;epipleurabroadenedathumeri,thenstronglyoblique
downwards.
Prosternum rugose, slightly raised along the middle; presternal process acute
cuneiform,pointedatapex. MesosternumwithmedianridgeU-shaped,weaklyraised,
but theanterior anglesslightlyandacutelyproduced forwards. Metasternumrather
short,stronglyraisedtowardsmiddle.
Legsratherslender;profemora1teethacute,moderateinsize,posteriormarginsof
meso- and metafemora distinctlyemarginate ineachapical third; inner margin of
protibiadistinctlyemarginateat theportionbetweenbasal two-sixthsandfive-sixths,
thatofmesotibiaalsodistinctlyemarginate,withdensesetaeinapicalhalf,andthatof
metatibiastrongly depressed throughout, the emarginations of pro- and mesotibiae
somewhat twisted.
Fernale. Emarginationofprotibiaweakandsituatedbetweenbasaltwo-fifths
and four-fifths.
I)1agnosls. This new species is closely allied toP. iridipennis (GEBIEN) from
Mindanao,but isdistinguished fromthelatter bythefollowingfeatures:bodyrather
weaklyconvexabove;elytranotiridico1orous;presternalprocesshorizontal;headand
pronota1puncturesdenser, innerocularsulcideepandmoredistinct;pronotumdevoid
of fasciae,distinctlynarroweratbasethanelytra1base,1.5timesaswideaslonginstead
of almost twiceaslongin thelatter;andelytradevoidofzigzag fasciae.
Etymology. ThespecificnameofthisspeciesisdedicatedtothelateDr.Masataka
SAT0,oneofmybestfriends,whomadeagreatcontributiontotheentomologynotonly
inJapanbut alsoin Taiwan.
Figs.1-39. Phaedisspp. - 1-8,Rightlateralmarginsofpronota;9-15,mesotibiaeinmale; 16-22,
metatibiae in male; 23-24, righthumeral corners in dorsal view; 25-31, presternal processes
(right: dorsalview;left:lateralview);32-39,malegenitalia(left:dorsalview; right:lateralview).
1, 12, 19, 23, 28, 35, P. (P)ovalipennis ovalipenus (KRAATz); 2, 24, 36,P. (P)ovalipennls
inornat1lssubsp nov; 3, 9, l6, 25, 32,P. (P) masatakalsp nov ; 4, 10, 17, 26, 33, P. (P)
minltttts (PIc);5,11, 18,27, 34,P. (P)altilissp nov;6, 15, 22,31,39,P. (P)celebenslssp.
nov;7, 13,20,29,37,P. (P)ohbayashtisp nov ;8,14,21,30,38,P. (P)no,r1orivagussp nov.
Scales:0.5mmforFigs 9-22,25-39and1.0mmforFigs.1-8,23-24.
NewPhaedisfromSulawesi 167
168 Kivoshi ANDo
Phaedis(Phaedis)mmutus(PIc,1921),comb nov
(Figs 4,10,17,26,33,46&47)
McrogauromaiaminutaPIc,1921,23
Measurements. Bodylength:9.5mm;width:4.4mm.
Typeexamined. 1ex.,S.Celebes, Bua-Kraeng,5000', Febr.1896, H. FRuHs-
TORFER (MNHN).
Specimen examined. 1 , Indonesia, Pangia,Sulawesi Selatan, 27-XII-1999, N.
OHBAYASHIleg. (CKA0).
Notes. Intheexaminedtype,theprofemurisarmedwithadistinct tooth,andthe
external generic charactersare indicativeof the genusPhaedis. In thisspecies, the
medianedgeofmesosternumhasproducedanteriorangles;mentumshort linguiform,
withlateralexcavationsdeep;maxillarypalpusright-angledtriangular;innermarginof
mesotibiaecarinatealongthemiddle.
PhaedisOPhaedis)altilissp nov
(Figs 5, ll,18,27,34&45)
Measurements. Body length:7.5-9.5mm;width:3.8-4.7mm.
Typeseries. Holotype: Indonesia,PuncakPa1opo,SulawesiSelatan,2-I-2000,
,
N. 0HBAYASHI leg. (EUMJ). Paratypes: ld1', 1早,samedataas for theholotype
(CKA0); lf,1早,PuncakPa1opo,S.Sulawesi,2-I-2000,K.AND01eg.(CKA0);1早,
Po1opo,Sulawesi,7-II-1985,M.TAOleg.(CKMT);2早早,ditto,5-VI-l982,M.TAO
leg.(CKMT); ld'',ditto,6-VI-1982,M.TAOleg.(CKMT).
Oval,distinctlycalabashbottle-shaped,stronglyconvexaboveinhindbody,shiny,
shoulderseffaced. Colourdarkreddishbrown;headandpronotummetallicgreen,with
violetlustre;elytradarkmetallicgreenoraeneous;sixdistalsegmentsofantennaedark
brown;venterdarkreddishbrownordarkbrowninpart.
M ale. Headquadrate,distinctlysloping forwards, withpunctures denseand
minute,rathersparserbehindfronto-clypea1suture;clypeusdistinctlyconvex,shallowly
emarginateat apex inmedianhalf;genaelonger thanwide,weaklyroundedatsides,
eachwitharounddepressionin front ofeye;postgenaegentlyconvergentposteriad;
fronsraisedandalmost flat,IE/TD=2.86;eyessmall,hardlyproducedlaterad, inner
ocularsulciveryweak. Antennaereachingmiddleofpronotum, fivedistalsegments
forming a weak club. Terminal segment of maxillary palpus nearly right-angled
triangular. Mentumnarrowlinguiform,deeplyexcavatedatsides,withcoarsesurface.
Pronotumtransverselytrapezoidal,widestbehindmiddle,PW/PL=1.49;anterior
margin distinctly emarginate, with border thick and almost entire; anterior angles
obtusely rounded and a little produced; lateral margins weakly rounded in apical
two-thirds, and sinuous in the rest, very narrowly bordered; discconvex forwards,
denselypunctate,thepuncturesalmostsimilarinsizetooralittlelarger thanthoseon
NewP11aedls fromSulawesi 169
frons.
Elytraoval,stronglyconvex,almostdevoidofhumeralcalli,not fusedwitheach
other, widest atmiddleand EL/EW=1.43;striaefineandweakly impressed,stria1
puncturesminuteandratherdense,irregularlyarranged,slightlylarger in7thand8th
striae,spacebetween themabout one toseven timesaswideasapuncture; intervals
almost fiat, densely and minutely punctate; epipleura very broad at humeri, with
microscopical rugosities. Hindwingsvestigial,about ahalf lengthofeachelytron.
Prosternumshort in front of coxae, presternalprocessnarrowly cuneate,moder-
atelyobliquelyslopingposteriadbutnotadunc,distinctlydepressedalongmiddleand
acuteatapex. Mesosternumwithmedianridgedistinct,acutelyproducedforwardsat
thecorners. Metasternumveryshort,nearlyaslongasadiameterofmesocoxa,coarsely
punctate. Analsterniteoftenslightlyemarginateat apex.
Legs robust; profemora1teeth moderately produced laterad, rectangular; inner
margins of tibiae distinctly emarginatebasally, protibiaeemarginate in basal half,
mesotibiaeemarginateinbasalfour-ninths,andmetatibiaestronglyemarginateinbasal
half, theremaininginnermarginsofmeso-andmetatibiaedenselypubescent.
Fema1e. Antennaeshorter; legs ratherslender,emarginationsof tibiae very
weak.
Diagnosis. Thisnewspeciesissimilar top minutus (PIc) fromSulawesi,but is
easilyseparated from the latterbyhavingthebodymore tumid; elytramoredilated,
intervals hardly convex,striaemoreweakly impressed,strial punctures well visible
insteadofalmost invisibleinthelatter;mentumnarrowlinguiform; innerocularsulci
feeble; pronotummoredensely anddistinctly punctate, withanterior marginmore
thicklybordered; innermarginofmesotibiainmaledistinctlyemarginatebasallyand
devoidof carinae in middle.
Etymology. Thespecificnameisderivedfromstronglydilatedbody.
Phaedis(Phaedis)ovah'pennisovah'penm's(KRAATz, l880)
(Figs.1,12,19.23,28,35,40&41)
Pselldellmofpusovaf1pe11川sKRAATz,1880, ll3
Measuremertts. Bodylength:9.1-11.0mm;width:4.2-4.7mm.
Typee:lcamlned. 1ex.,Menado,(BATES),Haag. (ZSM).
Speclmensexammed. l ,1早,Tondano,N.Sulawesi,10-IV-1989,Y.MIYAKEleg.
(CKA0);1早, ditto,9-IV-1989,Y. MIYAKEleg. (CKA0); ld1',Sulawesi:TemOnOn,
Rurukan,Gn.Mahawu,30-XI~3-XII-1999,1,150-1,200m,1eg.A.RIEDEL(SMNS);
Indonesia,N-Sulawesi,1kmSSawangan,Flui3sta1b.RiverParkHotel,01°22'61″
l
N,,l24°56'56″Eel~03-II-2004,250-300m,1eg.A.WEIGEL,UWS/Plant. (CRGT).
Notes. Elytranot fusedwitheachother,hindwingsreduced,nearlyaston9as
eachelytron,wingvenationsvestigial.
170 KivoshiANDo
Phaedis (Phaedis)ovalipennisinornatussubsp nov
(Figs 2,24,36&42)
Measurements. Bodylength:8.6-11.0mm;width:4.3-5.0mm.
Typeseries. Holotype:d'',Sulawesi:Kotamobagu,Modoinding,Gn. Ambang,6-
XII-1999,1,100-1,450m,1eg.A.RIEDEL(SMNS).Paratypes: lf,1早,samedataasfor
theholotype (SMNS&CKA0);2 1早,Sulawesi: Temonon,Rurukan,Gn. Ma-
,
hawu,30-XI~3-XII-1999,1,150-l,200m,1eg.A. RIEDEL(SMNS).
Thisnewsubspeciesisseparablefromthenominotypica1subspeciesbythefollow-
ingpoints:bodyreddishbrown,stronglygreenishbrassyondorsum,neither reddish
purplelustrenorbluishgreensutural intervals; fourdistalsegmentsofantennae,and
femoramoreor lessdarkened; headshorter in frontal half; pronotummorecoarsely
punctate, withlateralmargins feeblyorscarcelysinuousbeforebase;elytradevoidof
stripes, with humeral corners obtusely angulate, intervals coarsely punctate;
m ore
parameresindorsalviewnotconstrictedinmedianthird,gentlyconvergent frombase
toapicalthird,andstronglysotoapicesinapicalthird.
Etymology. Thesubspecificnamereferstotheimmaculateelytra.
Phaedis(Phaedis)ohbayashiisp
no v
(Figs7,13,20,29,37&48)
Measurements. Bodylength:7.3-7.5mm; width:3.3-3.5mm.
Typeseries. Holotype: Indonesia,Bontongan,SulawesiSelatan,29-XII-1999,
,
N.0HBAYASHIleg. (EUMJ). Paratype:1早,samedataasfor theholotype(CKA0).
Oblong-oval,weaklycalabashbottle-shaped,moderatelyshiny. Colourdarkred-
dishbrown,dorsalsidedarkgreenishbrassy,sevenbasalsegmentsofantennaereddish
brown,elytrawithmoreor lesspurplishlustre.
M ale. Headsemicircular,moderatelysloping forwardsfromvertex,minutely
andsparselypunctate;clypeusflat, roundlyproduced forwards; fronto-clypea1suture
narrowandobscure,visibleinpart;genaeroundedatsides,buttheircurvatureisnot
continuoustothat ofclypeus;eyessmall,hardlyproducedlaterad, innerocularsulci
narrow,clearlyengraved; fronswithapairofovaldepressionsinfrontofsuture,wjth
Puncturesalittlelargerandsparser thanthoseonclypeus,IE/TD=3.17;postgenae
short,roundlytumidlaterad. Antennaemoderateinlength,reachingbeforemiddleof
Pronotum; four distal segments dilated, forminga small club. Maxillary palpus
triangular. Mentumlinguiform,withapairofcoarsedepressionsnearbase.
Fi9S.40-49. HabitusofPhaedisspp. - 40,P. (P.)ovatipen1usovalipennls (KRAATz),specimen
examined; 41,P. (P) oyatipennisovalipent1is(K1tAATz), type;42,P. (P)ovaljpennlsjnornatus
SubSP nov;43,P. (P)masataka1sp nov;44,P. (P)nemor、i,agussp nov;45,P. (P)altjljssp.
nov;46,P. (P)minutlis(PIc),type;47,P. (P)mintltus(PIc),specimenexamined;48,p. (p)
o ayas/1l1sp nov;49,P. (P)ce/ebensls sp nov.
NewP11aedisfromSulawesi 171
40
43
47 48
172 Kiyoshi ANDo
P「onotumtransverselyquadrate,gentlyroundedandnotsinuousatsides,moder-
atelyconvexforwards,widestatmiddle,withborderverynarrowexcept forbasalone,
PW/PL=1.48;discdenselyandminutelypunctate,thepuncturesalmostsimilarinsize
tothoseonfrons;anteriormarginnearlystraight;anterioranglesobtuse,notproduced.
Elytraoblong-oval,fusedwitheachother,slightlywiderthanpronotumatthebase,
distinctlyconvex,widest at basal two-fifths, thenstronglynarrowedtowardsapices,
EL/EW=1.43;shoulderseffaced, lateralbordersverynarrow, invisiblefromabove;
striaefine,stria1puncturesrather largeandirregular indensity,constantlybecoming
Smalleratleastinapicalthird; intervalsfeeblyconvex,denselyandminutelypunctate;
epipleurasmooth,broadenedathumeralportion. Hindwingsvestigial.
Prosternumnotraisedalongthemiddle;presternalprocessnarrowbetweencoxae,
aduncanddivergentposteriadbehindthem,roundlyconvergentapicadandobtusely
「ounded at the apex. Mesosternum with median ridge very weak and declivous
fo「Wa「dS. Metasternumshort,shorter than thediameter ofmesocoxa in themedjan
length.
Le9SShort;profemorawithanteriorteethveryweakandobtuse; innermarginof
meSOtibiadistinctlyemarginateinbasalthirdandfeeblysointherest,sparselysetous;
innermarginofmetatibiaweaklybisinuous,andhardlypubescentinapicalhalf.
Female. Innermarginsofmeso-andmetatibiaehardlysinuous.
Diagnosis. ThisnewspeciessomewhatresemblesPminor(PIc) fromTonkin,but
Si 「eadilydistinguishedfromthelatterbythemorestronglypear_shapedhindbody,the
elyt「anotpurplebutgreenishbrassy,stria1punctureslesscoarselyarranged, jntervals
denselyPunctate,feeblyconvexinsteadofdistinctlysointhelatter;thepronotumnearly
straightatapex,withlateralmarginsnotsinuousbeforebase.
Etymology. ThisnewspeciesiscordiallydedicatedtoDr.NobuoOHBAYAsHI,the
collectorofthetypeseries.
PhaedisOPhaedis)nemorivagussp
nov
(Figs8,14,21,30,38&44)
Measuremeflts. Bodylength:12.0-12.6mm;width:5.3-5.5mm.
Typeseries. Holotype:d'',PuncakPa1opo,S.Sulawesi, Indonesja,2-I-2000, K
ANDoleg. (EUMJ).Paratype:l PuncakPa1opo,Sampunaalt.1,050m,s.sulawesj,
,
23-I-2000,G.BEccEleg.(CKA0).
M ale. Oblong,moderatelycalabashbottle-shaped,largeinsjze. Colourblack,
head,P「onOtum, lateralmarginalportionandepipleuraofelytrametallicblue;major
Pa「t ofelyt「ab「aSsyorbrassy-green,withstrongpurplishrefiectjon; venterblackjsh
brown,withvioletlustreinpart.
Head heXa9ona1, almost fiat, and distinctly sloping forwards, with punctures
minute,moderateindensity;clypeusshallowlyemarginateatapexinmedjanthree_
fifths; f「onto-clypea1suturedistinct, longer in lateral parts than jn
medjan; genae
SubPa「allot-Sidedinbasal two-thirds;postgenaesimplynarrowedposteriad;eyesrather