Table Of Content

FISHES OF GADANARIVERIN KALAKKAD MUNDANTHURAI TIGERRESERVE 1 M.ArunachalamandA. Sankaranarayanan2 Keywords:KalakkadMundanthuraiTigerReserve,fishdiversity,WesternGhats Streams in Gadanariverbasin located inthebufferzoneofKalakkadMundanthurai Tiger ReserveweresurveyedduringJanuary 1997-98. Thirty-onespeciesbelongingto5 orders, 12familiesand22generawererecorded,ofwhichHypselobarbusdobsoni,Pseudambassis ranga, Puntiussarana orphoides and Glyptothoraxmadraspatnum were first records for this river basin. Introduction msl and flows down the eastern slopes of Western Ghats. During flooding, this river Documentation and conservation are the confluences with the Tamiraparani river. The current areas of concern for fish biologists. perennial Gadana river is drained by three Gadana river and its tributaries which flow in tributaries, viz. Pampar, Kallar and Iluppaiyar. thebufferzoneofKalakkadMundanthuraiTiger Pampar and Kallar are dammed and Iluppaiyar Reserve formasub-basinofamajorrivercalled streamjoinstheGadanariverbelowthereservoir. TamiraparaniinthesoutheasternWesternGhats. Below the confluence of Iluppaiyar, the river Tamiraparaniriverbasin constitutesthe Gadana traversesthroughplains, drainingmanyvillages river, Ramanadhi, Pachayar, Chittar, adjacent to the river for about 12 km south- ManimutharandJambunadhi.However,theriver eastwards,beforejoiningTamiraparanirivernear systems of Tamiraparani sub-basins were ThiruppudaiMarudurvillageinAmbasamudram completelyunexplored. Silas(1953) describeda taluk. The lengthofGadanariverfromits origin new species of Puntius arulius tambiraparniei to the confluence is about 33 km. from Tamiraparani river. Johnsingh and Wickram (1987) reported the freshwater fishes MaterialandMethods from Kalakkad Mundanthurai Wildlife Sanctuary. RemaDevietal. (1997)gavealistof Fishes were collected from several fishes from Tamiraparani river system. To date localities in the three tributaries using drag net, there is no record offishes from the sub-basins various mesh sizes ofgill nets and scoop nets. ofTamiraparaniriver.Thepresentsurveyispart The colour, spots ifany, andothercharacters of ofaprogramme onthe fishdiversity inWestern the fishes caught were noted andthe specimens Ghats streams. were preserved in 10% formalin. Study Area Systematic Account Among the 31 species recorded from Gadana river, with its tributaries, forms a Gadanariver,asystematicaccountof23 species sub-basin in the Tamiraparani river basin. This is given below; detailed accounts on 8 species river originates (8° 48' N lat., IT 19' E long.) (Hypselobarbus dobsoni, Puntius arulius from Alwarkurichi and Kadayam ranges of tambiraparniei, Anguilla bengalensis, Puntius m sarana orphoides Ompok bimaculatus, Western Ghats at an altitude of 1,564 above , Bhavania australis Nemacheilus triangularis , 'AcceptedMarch, 1999 and Glyptothorax madraspatnum have been 'SriParamakalyaniCentreforEnvironmentalSciences published earlier (CAMP workshop 1997; ManonmaniamSundaranarUniversity Alwarkurichi627412,TamilNadu Arunachalam and Sankaranarayanan in press). 232 JOURNAL, BOMBAYNATURAL HISTORYSOCIETY, 96(2) AUG. 1999 FISHESOFGADANARIVER 1. Labeo calbasu (Hamilton-Buchanan) J. Lit & Sci 15: 314 (type locality: tanks and 1822, Cyprinus calbasu Ham.-Buch. rivers in the neighbourhood ofChennai). Fishes ofGanges: 297, 387 pi. 2, fig. 33 (type Material: 3 examples; 48 mm to 98 mm locality: rivers and ponds ofBengal and in the fromthree localities were examined. Westernprovinces) Distribution: Gadana river. First record. mm mm Material: 2 examples; 98 to 124 Elsewhere: Cauvery and Krishna river systems fromone locality (Iluppaiyar) were examined. and Sri Lanka. Distribution: Gadana river. First record. Status: Common in Gadana river. Elsewhere: Pakistan,Nepal,Myanmar,Thailand Remarks: It attains a standard length of and Yunnan (South China). 24cm(TalwarandJhingran 1991).We recorded Status: Not common in Gadana river. a maximum standard length of98 mm. Remarks: It attains a standard length of 90 cm (Talwar and Jhingran 1991). In our 5. Puntiusfilamentosus (Valenciennes) collections, we recorded a maximum standard 1844,LeuciscusfilamentosusValenciennes length of 124 mm. Hist. nat. Poiss, 17: 96 pi. 492 (type locality: Alleppey, Kerala State) mm mm 2. Puntius amphibius (Valenciennes) Material: 3 examples; 42 to 58 CapoetaamphibiaValenciennes, 1842,Hist, from one locality (Iluppaiyar) were examined. nat.Poiss., 16: 182,pi.478(typelocality:Bombay). Distribution: Gadana river. First record. mm mm Material: 3 examples; 32 to 84 Elsewhere: Goa,KarnatakaandTamilNadu, Sri from one locality (Pampar) were examined. Lanka and Thailand. Distribution: Pampar. First record. Status: Not common in Gadana river. Elsewhere: Orissa, Madhya Pradesh, Rajasthan Remarks:Attainsatotallengthof180mm and Sri Lanka. (Jayaram, 1991). Inourcollections we recorded Status: Not common in Gadana river. a maximum total length of58 mm. Remarks: It attains a standard length of 200 mm (Talwar and Jhingran 1991). But we 6. Puntius sarana sarana (Hamilton-Buchanan) recordedamaximumstandard lengthof84 mm. 1822, CyprinussaranaHam.-Buch.Fishes of Ganges: 307, 388 (type locality: ponds and 3. Puntius bimaculatus (Bleeker) rivers ofBengal). mm mm Gnathopogon bimaculatus Bleeker, 1844, Material: 4 examples; 72 to 1 11 Verh. Nat. Holl. Maatsch. Haarlem, (2) 20: 17, from one locality (Iluppaiyar) were examined. pi. 4, fig. 1 (type locality: Ceylon) Distribution: Gadana river. First record. mm mm Material: 4 examples; 36 to 48 Elsewhere: Afghanistan, Pakistan, throughout fromtwolocalities(PamparandIluppaiyar)were India except peninsula south of Krishna river. examined. Nepal, Bangladesh and Myanmar. Distribution: Gadana river. First record. Status: Common in Gadana river. Elsewhere: Bangalore. Remarks: It attains a standard length of Status: Common in Gadana river. 31 cm (Talwar and Jhingran 1991). In our Remarks: It attains a standard length of collections we recorded a maximum size of 100 mm (Talwar and Jhingran 1991). But we 111 mm in standard length. recordedamaximumstandardlengthof48 mm. 7. Puntiussarana subnasutus (Valenciennes) 4. Puntius dorsalis (Jerdon) 1842, Barbus subnasutus Val. Hist. nat. 1849, Systomus dorsalis Jerdon, Madras Poiss., 16:16:154 (type locality: Pondicherry) JOURNAL, BOMBAYNATURAL HISTORYSOCIETY, 96(2), AUG. 1999 233 FISHESOFGADANARIVER mm mm Material: 5 examples; 74 to 11 1 from one locality (Iluppaiyar) were examined. from one locality (Iluppaiyar) were examined. Distribution: Gadana river. First record. Distribution: Gadana river. First record. Elsewhere: Eastern and Western Ghats, Elsewhere: Krishna and Cauvery river systems Maharashtra, Madhya Pradesh, Gujarat and and Kerala in Peninsular India. Myanmar. Status: Common in Gadana river. Status: Not common in Gadana river. Remarks: It attains a standard length of Remarks: Salmostoma clupeoides is not mm 250 (Talwar and Jhingran 1991). But we common in Gadana river. It attains a standard recordedamaximumstandardlengthof111 mm. lengthof15 cm(TalwarandJhingran 1991). We recorded a maximumstandard length of84 mm. 8. Puntius ticto (Hamilton-Buchanan) 1822, Cyprinus-ticto Ham.-Buch. Fishes 11. Amblypharyngodon microlepis (Bleeker) ofGanges. 314, 398 pi. 8, fig. 87 (type locality: 1853,LeuciscusmicrolepisBleekerVerh. Southeastern parts ofBengal Batav Genoot. Kunst. Wet., 25:141 (type mm mm Material: 6 examples; 48 to 52 locality: Bengal). from one locality (Iluppaiyar) were examined. Material: 5 examples; 51 mm to 76 mm Distribution: Gadana river. F rst record. from one locality (Iluppaiyar) were examined. Elsewhere: Pakistan, lower Swatriverdrainage; Distribution: Gadana river. First record. India, Nepal, Sri Lanka, Bangladesh, Myanmar Elsewhere: Uttar Pradesh, Bihar, West Bengal, and Thailand. Orissa,AndhraPradesh,TamilNaduandKerala. Status: Not common in Gadana river. Status: Not common in Gadana river. Remarks: It attains a standard length of Remarks: It attains a standard length of mm mm 100 (Talwar and Jhingran 1991). In our 100 (TalwarandJhingran 1991). Butinour collections we recorded a maximum standard collections we recorded a maximum standard length of52 mm. length of76 mm. 9. Puntius vittatus Day 12. Danio aequipinnatus (McClelland) 1865,PuntiusvittatusDayProc.Zool. Soc. 1839, Perilampus aequipinnatus Lond: 303 (type locality: Cochin, Kerala) McClelland, Asiat. Res. 19(2): 393 pi. 60, fig. 1 mm mm Material: 10 examples; 23 to 26 (type locality: Assam) mm mm from one locality (Iluppaiyar) were examined. Material: 10 examples; 48 to 84 Distribution: Gadana river. First record. from three localities were examined. Elsewhere: Goa,Karnataka,Kerala,TamilNadu, Distribution: Gadana river. First record Kutch, Bihar, Rajasthan and Sri Lanka. Elsewhere: India, Sri Lanka, Bangladesh, Status: Common in Gadana river. Myanmar and Thailand. Remarks: It attains a total length of Status: CommoninGadanariver. Inhabits 25 mm(Jayaram 1991).Werecordedamaximum hill streams up to 300 m above msl. total length of26 mm. Remarks: It attains a standard length of 150mm(TalwarandJhingran 1991).Butweonly 10. Salmostoma clupeoides (Bloch) recorded a maximum standard length of84 mm. 1782, Cyprinus clupeoides Bloch, Naturges ausland Fische 12:49 pi. 408, fig. 2 13. Esomus themacois (Valenciennes) (type locality: “Indian ocean” evidently not the 1842, Nuria thermocois Val. Hist. nat. Oceanbut a freshwaterbody). Poiss. 16: 238 pi. 472 (type locality: hot spring mm mm Material: 5 examples; 62 to 84 at Kanniya, Sri Lanka). 234 JOURNAL, BOMBAYNATURAL HISTORYSOCIETY, 96(2) AUG. 1999 FISHESOFGADANARIVER mm mm Material: 3 examples; 42 to 83 Distribution: Gadana river. First record. from one locality (Iluppaiyar) were examined. Elsewhere: Kerala, Karnataka, Maharashtra and Distribution: Gadana river. First record. Sri Lanka. Elsewhere: Sri Lanka. Status: Common in Gadana river. Status: Not common in Gadana river. Remarks: Lepidocephalus thermalis Remarks: It attains a standard length of attains a standard length of80 mm (Talwar and 120 mm (Talwar and Jhingran 1991). But we Jhingran 1991). We recorded a standard length recordedamaximumstandardlengthof83 mm. of42 mm. 14. Parluciosoma daniconius 17. Mystusarmatus (Day) (Hamilton-Buchanan) 1865, Hypselobagrus armatus Day, Proc. 1822, Cyprinus daniconius Ham.-Buch. Zool. Soc. Lond: 289 (type locality: Cochin) mm mm Fishes ofGanges: 327, 391 pi. 15, fig. 89 (type Material: 3 examples; 42 to 68 locality: rivers ofSouthern Bengal) fromtwo localities(PamparandThoniyar) were mm mm Material: 10examples;42 to 103 examined. from Pampar and Thoniyar were examined. Distribution: Gadana river: First record. Distribution: Pakistan, India, Sri Lanka, Elsewhere: Wynaad hills, Western Ghats and Bangladesh, Myanmar and Thailand, Mekong Nagaland; probably also lower Myanmar. river basin. Status: Common in Gadana river. Status: Common in Gadana river, prefers Remarks: Mystus armatus attains a mm running water. standardlengthof145 (TalwarandJhingran Remarks: It attains a standard length of 1991). But we recorded a maximum standard 100 mm (Talwar and Jhingran 1991). We lengthof68 mmonly. recordedamaximumstandardlengthof103mm. 18. Aplochelius lineatus (Valenciennes) 15. Garra mullya (Sykes) 1846, Panchax lineatum Val. Hist. nat. 1841, Chondrostomamullya, Trans. Zool. Poiss., 18: 381 (type-locality: Peninsula, Soc. Lond. 2: 359, pi 62, fig. 3 (type locality: India) mm mm Bheema river at Daunde, near Pune). Material: 9 examples; 33 to 60 mm mm Material: 10examples;42 to 122 fromtwolocalities(PamparandIluppaiyar)were fromthree localities were examined. examined. Distribution: Gadana river. First record. Distribution: Gadana river. First record. Elsewhere:IndiaexceptAssamandtheHimalaya. Elsewhere: Western and Southeastern regions. Status: Common in Gadana river. Status: Common in Gadana river. mm mm Remarks: It grows upto 170 in Remarks: It attains a length of 100 standardlength(TalwarandJhingran 1991). We (Day 1878). But we recorded a maximum recordedamaximumstandardlengthof122mm. standard length of60 mm only. 16. Lepidocephalus thermalis 19. Pseudambassis ranga (Valenciennes) (Hamilton-Buchanan) 1846, Cobitis thermalis Val. Hist. nat. 1822, Chanda ranga Ham.-Buch. Fishes Poiss., 18: 78 (type locality: Malabar) ofGanges 113, 371 pi. 16, fig 38 (type locality: mm mm Material: 2 examples; 35 to 42 freshwaters ofall Gangetic provinces). mm mm fromtwo localities(PamparandThoniyar)were Material: 3 examples; 32 to 38 examined. fromone locality (Iluppaiyar) were examined. JOURNAL, BOMBAYNATURAL HISTORYSOCIETY, 96(2), AUG. 1999 235 FISHESOFGADANARIVER Distribution: Gadana river. First record. Remarks: It attains a standard length of Elsewhere: Pakistan, India, Bangladesh, 75 mm (Talwar and Jhingran 1991). But we Myanmar, Thailand and Malaysia. recordeda maximum standard length of32 mm. Status: Not common in Gadana river Remarks: It attains a standard length of 23. Mastacembelus armatus (Lacepede) mm 70 (Talwar and Jhingran 1991). But we 1800, Macrognathus armatus Lacepede, recordedamaximumstandard lengthof38 mm. Hist.nat.Poiss,2: 286(type-locality:notknown) mm mm Material: 2examples; 140 to510 20. Etroplus maculatus (Bloch) from three localities were examined. Chaetodon maculatus Bloch, 1785, Syst. Distribution: Gadana river. First record. Ichth. PI. 427, fig. 2 (type locality: India) Elsewhere: Pakistan, India, Sri Lanka, Nepal, Material: 5 examples; 30 mm to 44 mm Myanmar through Thailand and Malaya to from one locality (Iluppaiyar) were examined. Southern China. Distribution: Gadana river. First record. Status: Common in Gadana river. Elsewhere:India: Orissa,AndhraPradesh,Tamil Remarks: It attains a standard length of Nadu, Kerala and Sri Lanka. 61 mm (Talwar & Jhingran 1991). In our Status: Not common in Gadana river. collections we recorded a maximum size of Remarks: It attains a standard length of 510 mm. mm 80 (Talwar and Jhingran 1991). But in our collections we recorded a maximum standard Resultsand Discussion length of44 mm. From our study, it is evident that Gadana 21. Oreochromis mossambica (Peters) river drainage system is rich in fish diversity. A 1852, Chromis (Tilapia) mossambicus total of 32 species belonging to 21 genera were Peters, Montab, Akad, Wiss., Berlin: 681 (type recordedfromeachlocalityofthethreetributaries locality: Mozambique) (Table 1). Of all these three localities, Pampar mm mm Material: 6 examples; 42 to 240 and Iluppaiyar are less disturbed. The banks of fromone locality (Iluppaiyar) were examined. theIluppaiyarstreamareunstableduetofarming Distribution: Gadana river. First record. activities. Even though Iluppaiyar stream is Elsewhere: East Africa; introduced in India, disturbed beyond the riparian zone, the fish Pakistan and Sri Lanka. species richness is high compared to the other Status: Common in Gadana river. two streams. This maybe due to the presence of Remarks: In our collections we recorded rich riparian strips. a maximum standard length of240 mm. Puntiussaranaorphoides, Pseudambassis ranga Glyptothorax madraspatnum are , 22. Macropodus cupanus (Valenciennes) comparatively rare and Hypselobarbus dobsoni 1831, Polyacanthus cupanus, Hist. nat. occurs in great abundance in Gadana river, Poiss., 7: 357 (type locality: Ariancoupon river but they are not recorded in other river systems at Pondicherry) constituting the Tamiraparani river basin. mm mm Material: 2 examples; 28 to 32 Puntius sarana orphoides was originally from one locality (Iluppaiyar) were examined. described by Valenciennes from Java. Menon Distribution: Gadana river. First record. (1963) recorded this subspecies from Manipur. Elsewhere: Eastern India, Sri Lanka, Western This species was first reported from Western Myanmar, Malay peninsula and Sumatra. Ghats of South India. Menon (1992) recorded Status: Not common in Gadana river. Hypselobarbus dobsoni from the Krishna river 236 JOURNAL, BOMBAYNATURAL HISTORYSOCIETY, 96(2) AUG. 1999 * FISHESOFGADANARIVER Table 1 FISH SPECIESRECORDEDINGADANA RIVER Species Pampar Kallar Iluppaiyar Species Pampar Kallar Iluppaiyar I Order:Anguilliformes iv) Family:Cobitidae i) Family:Anguillidae Genus:Lepidocephalus Genus:Anguilla 22. Lepidocephalusthermalis 1 1 2 1. Anguillabengalensis bengalensis l l 1 III) Order:Siluriformes v) Family:Bagridae II Order:Cypriniformes Genus:Mystus ii)Family:Cyprinidae 23. Mystusarmatus 1 2 2 Genus:Hypselobarbus 2. Hypselobarbusdobsoni* 4 4 4 vi) Family:Siluridae Genus:Labeo Genus:Ompok 3. Labeocalbasu - - 2 24. Ompokbimaculatus - - 2 Genus:Puntius 4. Puntiusamphibius 1 - 2 vii) Family:Sisoridae 5. Puntiusbimaculatus 2 3 4 Genus:Glyptothorax 6. Puntiusdorsalis 2 2 2 25. Glyptothorax 7. Puntiusarulius madraspatnum 1 - 2 tambiraparniei 2 2 4 8. Puntiusfilamentosus - - 3 IV) Order:Cyprinodontiformes 9. Puntiussaranaorphoides** - - 4 viii)Family:Aplocheilidae 10. Puntiussaranasarana 2 1 3 Genus:Aplocheilus 11. Puntiussarana 26. Aplocheiluslineatus 2 5 6 subnasutus 2 5 6 12. Puntiusticto - - 10 V) Order:Perciformes 13. Puntiusvittatus 3 2 10 ix) Family:Ambassidae Genus:Salmostoma Genus:Pseudambassis 14. Salmostomaclupeoides - - 5 27. Pseudambassisranga* - - 3 Genus:Amblypharyngodon 15. Amblypharyngodon x)Family:Cichilidae microlepis - 5 Genus:Etroplus Genus:Danio 28. Etroplusmaculatus - 5 16. Danioaequipinnatus 3 3 4 Genus:Orechromis Genus:Esomus 29. Orechromismossambica 2 2 6 17. Esomusthermoicos - - 3 Genus:Parluciosoma xi) Family:Belontiidae 18. Parluciosomadaniconius 6 4 2 Genus:Macropodus Genus:Garra 30. Macropoduscupanus - - 2 19. Garramullya 4 3 3 iii) Family:Balitoridae xii) Family:Mastacembelidae Genus:Bhavania Genus:Mastacembelus 20. Bhavaniaaustralis 2 - - 31. Mastacembelusarmatus 1 1 1 Genus:Nemacheilus *FirstrecordforTamirabaraniriverbasin. 21. Nemacheilustriangularis 2 2 5 **FirstrecordforWesternGhats. drainage. Itextends itsrangetoTamilNadupart it for the first time in Tamiraparani river basin. of Western Ghats as a dense population. Pseudambassis ranga was originally described Glyptothorax madraspatnum has so far been intheGangeticProvincesbyHamilton-Buchanan reportedfromAralamWildlifeSanctuary,Kerala (1822). Later Ajithkumar and Vijayan (1988) byShajietal. (1995).Thepresentsurveyreports recordedthisspeciesfromKeoladeoNationalPark, JOURNAL, BOMBAYNATURAL HISTORYSOCIETY, 96(2), AUG. 1999 237 1 FISHESOFGADANARIVER We Bharatpur, Rajasthan. are reporting it from Reserve) andMr. Sornappan (Asst. Conservator Gadana river for the first time. of Forest, Kadayam range) for their co-opera- We ; tion. also thank Mr. A. Vanarajan and Acknowledgements Mr. K. Sankar (Project Assistants - DBT) for their assistance, Dr. K. Rema Devi (Scientist, One of the authors (M.A.) thanks the ZSI, Chennai) for confirming the identifica- Department of Biotechnology, Ministry tions of fish specimens and Dr. P.T. Cherian ofScienceandTechnology,GovernmentofIndia (Officer-in-charge),ZoologicalSurveyof India, We for financial assistance. thank Dr. Melkani SouthernRegionalStation, Chennaiforresearch (Field Director, Kalakkad Mundanthurai Tiger facilities. Refer :nces Ajithkumar,C.R.&V.S.Vijayan(1988):Onthefishfauna Mundanthurai Wildlife Sanctuary, Tamil Nadu. ofKeoladeo National Park, Bharatpur (Rajasthan), J. Bombaynat. Hist. Soc. 84(3): 526-533. J. Bombaynat. Hist. Soc. 85(1):45-49. Menon,A.G.K.(1963):Spoliazeylanica, 36(1):p.69. Arunachalam, M. & A. Sankaranarayanan (in press): Menon,A.G.K.(1992):Conservationoffreshwaterfishes Some economically important/cultivable fishes in ofPeninsularIndia.Unpublishedreport(GrantNo. 14/ Gadana river. Western Ghats ofsouth Tamil Nadu. 24/87-MAB/REdt. 12.8.88)MinistryofEnvironment Proc. Workshopon Germplasm inventoryandGene &Forests,Govt,of India,p. 136 bankinginfreshwaterfishes. NationalBureauon Fish Molur,Sanjay&SallyWalker(eds.)(1997): Reportof GeneticResources,Lucknow. the workshop “Conservation Assessment and Arunachalam, M. &A. Sankaranarayanan (inpress): Management Plan (CAMP) for freshwater fishes of OrnamentalstreamfishPuntiusaruliustambraparniei India”ZOOOutreachOrganisation,Coimbatore,India, endemictotheTamiraparaniriver.WesternGhatsof p. 156 southTamilNadu.Ibid. RemaDevi,K.,T.J. Indra,M.B.Raghunathan,M. Mary Arunachalam, M. & A. Sankaranarayanan (inpress): Bai, & M.S. Ravichandran (1997): Ichthyofaunaof EconomicallyimportantcultivablefishHypselobarbus theTamiraparaniriversystem,TamilNadu.Zoosprint. dobsoniendemic to Gadanariver, WesternGhatsof Vol.12(7): 1,2. southTamilNadu.Ibid. Shaji, C.P., P.S. Easa & S. Chand Basha (1995): Day(1878):TheFishesofIndia,4thedn.JagmanderBook FreshwaterfishdiversityinAralamWildlifeSanctuary, Agency,NewDelhi,p.778. Kerala,SouthIndia.J. Bombaynat.Hist.Soc. 92:360- Hamilton-Buchanan,F.(1822):Anaccountofthefishes 364. foundintheriverGangesanditsbranches.Edinburgh Silas,E.G.(1953):NewfishesoftheWesternGhatswith andLondon,pp.vii+405,39pis. notesonPuntius arulius (Jerdon). Rec. Indian Mus. Jayaram, K.C. (1991): Revision ofthe genus Puntius 57:27-38. HamiltonfromtheIndianregion.Rec.Zool.Surv.India. Talwar, P.K. & A.G. Jhingran(1991): Inland fishes of Occ.PaperNo. 135: 178. India and adjacent countries, Oxford & IBH, New Johnsingh, A.J.T. & D. Vickram (1987): Fishes of Delhi.Vol. I&II,p. 1158 238 JOURNAL. BOMBAYNATURAL HISTORYSOCIETY. 96(2) AUG. 1999