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Fish & Wildlife: Principles of Zoology and Ecology, 3rd edition PDF

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& F W ISH ILDLIFE Principles of Zoology and Ecology Third Edition & F W ISH ILDLIFE Principles of Zoology and Ecology Third Edition L. DeVere Burton Australia • Brazil • Japan • Korea • Mexico • Singapore • Spain • United Kingdom • United States Fish & Wildlife: Principles of Zoology © 2010, 2003, 1995 Delmar, Cengage Learning and Ecology, Third Edition ALL RIGHTS RESERVED. No part of this work covered by the copyright L. DeVere Burton herein may be reproduced, transmitted, stored, or used in any form or by Vice President, Career and Professional Editorial: any means graphic, electronic, or mechanical, including but not limited to Dave Garza photocopying, recording, scanning, digitizing, taping, Web distribution, information networks, or information storage and retrieval systems, except Director of Learning Solutions: as permitted under Section 107 or 108 of the 1976 United States Copyright Matthew Kane Act, without the prior written permission of the publisher. Acquisitions Editor: Benjamin Penner For product information and technology assistance, contact us at Managing Editor: Cengage Learning Customer & Sales Support, 1-800-648-7450 Marah Bellegarde For permission to use material from this text or product, Product Manager: submit all requests online at cengage.com/permissions Christina Giff ord Further permissions questions can be emailed to Editorial Assistant: [email protected] Scott Royael Vice President, Career and Library of Congress Control Number: 2008942494 Professional Marketing: ISBN-13: 978-1-4354-1963 6 Jennifer McAvey ISBN-10: 1 4354 1963 4 Marketing Director: Debbie Yarnell Marketing Manager Delmar Erin Brennan 5 Maxwell Drive Marketing Coordinator: Clifton Park, NY 12065-2919 Jonathan Sheehan USA Production Director: Carolyn Miller Cengage Learning is a leading provider of customized learning solutions Production Manager: with offi ce locations around the globe, including Singapore, the United Andrew Crouth Kingdom, Australia, Mexico, Brazil, and Japan. Locate your local offi ce at: Content Project Manager: www.cengage.com/global. Katie Wachtl Senior Art Director: Cengage Learning products are represented in Canada by Nelson Education, Ltd. David Arsenault Technology Project Manager: Tom Smith To learn more about Delmar, visit www.cengage.com/delmar Production Technology Analyst: Visit our corporate website at www.cengage.com Thomas Stover Notice to the Reader Publisher does not warrant or guarantee any of the products described herein or perform any independent analysis in connection with any of the product information contained herein. Publisher does not assume, and expressly disclaims, any obligation to obtain and include information other than that provided to it by the manufacturer. The reader is expressly warned to consider and adopt all safety precautions that might be indicated by the activities described herein and to avoid all potential hazards. By following the instructions contained herein, the reader willingly assumes all risks in connection with such instructions. The publisher makes no representations or warranties of any kind, including but not limited to, the warranties of fi tness for particular purpose or merchantability, nor are any such representations implied with respect to the material set forth herein, and the publisher takes no responsibility with respect to such material. The publisher shall not be liable for any special, consequential, or exemplary damages resulting, in whole or part, from the readers’ use of, or reliance upon, this material. Printed in Canada 1 2 3 4 5 6 7 13 12 11 10 09 Table of Contents Preface - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -x Acknowledgments - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - xiii About the Author - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - xiv SECTION I: ZOOLOGY AND ECOLOGY BASICS - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 1 CHAPTER 1 Principles of Zoology - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 2 Classifi cation of Animals - - - - - - - - - - - - - - - - - - - - - - -3 Animal Behaviors and Habits - - - - - - - - - - - - - - - - - - - 4 Animal Structure or Anatomy - - - - - - - - - - - - - - - - - - -6 Animal Nutrition and Digestion - - - - - - - - - - - - - - - - - -8 Animal Growth - - - - - - - - - - - - - - - - - - - - - - - - - - - - -10 Animal Reproduction- - - - - - - - - - - - - - - - - - - - - - - - - 11 CHAPTER 2 Principles of Ecology - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 17 Conservation of Matter - - - - - - - - - - - - - - - - - - - - - - - 18 Laws of Energy - - - - - - - - - - - - - - - - - - - - - - - - - - - - -24 The First Law of Energy - - - - - - - - - - - - - - - - - - - - -25 The Second Law of Energy - - - - - - - - - - - - - - - - - - -26 Natural Cycles - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -27 The Carbon Cycle - - - - - - - - - - - - - - - - - - - - - - - - -27 The Nitrogen Cycle - - - - - - - - - - - - - - - - - - - - - - - -29 The Water Cycle - - - - - - - - - - - - - - - - - - - - - - - - - -30 Food Chains - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -33 CHAPTER 3 Understanding Relationships Between Wildlife and Agriculture - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 39 Fundamentals of Ecology - - - - - - - - - - - - - - - - - - - - - 40 Agricultural Impacts on Ecosystems - - - - - - - - - - - - -43 Biological Succession - - - - - - - - - - - - - - - - - - - - - - - 46 CHAPTER 4 Biomes of North America - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 53 Freshwater Biomes - - - - - - - - - - - - - - - - - - - - - - - - - -55 Marine Biome - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 60 Terrestrial Biomes - - - - - - - - - - - - - - - - - - - - - - - - - - 64 Desert Biome - - - - - - - - - - - - - - - - - - - - - - - - - - - 64 Tundra Biome - - - - - - - - - - - - - - - - - - - - - - - - - - - -66 Grassland Biome - - - - - - - - - - - - - - - - - - - - - - - - - -68 Deciduous Forest Biome - - - - - - - - - - - - - - - - - - - -69 Temperate Forest Biome - - - - - - - - - - - - - - - - - - - -70 Coniferous Forest Biome - - - - - - - - - - - - - - - - - - - -70 v CHAPTER 5 Our Wildlife Resources - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 75 A National Treasure - - - - - - - - - - - - - - - - - - - - - - - - - -76 The U.S. Endangered Species Act - - - - - - - - - - - - - -78 Controversy and Protectionism - - - - - - - - - - - - - - - - -79 Extinction and its Causes - - - - - - - - - - - - - - - - - - - - - -85 Managing Endangered and Threatened Species - - - - -88 SECTION II: ZOOLOGY AND ECOLOGY OF MAMMALS - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 93 CHAPTER 6 Gnawing Mammals - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 94 Mice, Rats, Voles, and Lemmings - - - - - - - - - - - - - - - -97 Squirrels - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 105 Marmots and Prairie Dogs - - - - - - - - - - - - - - - - - - - - 113 Rabbits, Hares, and Pikas - - - - - - - - - - - - - - - - - - - - - 117 Porcupines - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 123 Gophers and Beavers - - - - - - - - - - - - - - - - - - - - - - - - 124 CHAPTER 7 Hoofed Mammals - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 130 Pronghorn - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 133 Deer - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 135 Bison and Musk Ox - - - - - - - - - - - - - - - - - - - - - - - 142 Wild Sheep - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 145 Mountain Goats - - - - - - - - - - - - - - - - - - - - - - - - -148 Peccary - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -149 Horses and Burros - - - - - - - - - - - - - - - - - - - - - - - - 150 CHAPTER 8 Predatory Mammals - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -155 Wild Cats - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 156 Wild Dogs - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 161 Bears - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 167 Weasels - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 171 Raccoons - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 181 CHAPTER 9 Marine Mammals - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 186 Seals - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 187 Eared Seals - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 189 True Seals - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 191 Walrus- - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 193 Manatees - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -194 Whales - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 196 Baleen Whales - - - - - - - - - - - - - - - - - - - - - - - - - - - 197 Toothed Whales - - - - - - - - - - - - - - - - - - - - - - - - - 199 CHAPTER 10 Unusual Mammals - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 209 Opossums - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 210 Shrews and Moles - - - - - - - - - - - - - - - - - - - - - - - - - - 212 Bats- - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 214 Armadillo - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 218 vi SECTION III: ZOOLOGY AND ECOLOGY OF BIRDS - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 223 CHAPTER 11 Waterfowl - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 224 Ducks - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -227 Geese - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 233 Swans - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -236 CHAPTER 12 Game Birds - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 242 Quails - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 244 Partridges - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -246 Pheasants - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -247 Grouse - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -249 Turkeys - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 253 Pigeons and Doves - - - - - - - - - - - - - - - - - - - - - - - - - 255 CHAPTER 13 Birds of Prey - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 261 Hawk-like Birds - - - - - - - - - - - - - - - - - - - - - - - - - - - -262 Eagles, Hawks, and Kites - - - - - - - - - - - - - - - - - - -262 Falcons - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -270 Vultures - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 273 Osprey - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 275 Owls - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -276 Typical Owls - - - - - - - - - - - - - - - - - - - - - - - - - - - -276 Barn Owls - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 281 CHAPTER 14 Songbirds and Other Perching Birds - - - - - - - - - - - - - - - - - - - - 286 Songbirds - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -287 Crows and Jays - - - - - - - - - - - - - - - - - - - - - - - - - - - - 301 Cuckoos - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -303 Kingfi shers - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -304 Hummingbirds - - - - - - - - - - - - - - - - - - - - - - - - - - - -304 CHAPTER 15 Other Birds of North America - - - - - - - - - - - - - - - - - - - - - - - - - 309 Long-legged Wading Birds - - - - - - - - - - - - - - - - - - - - 310 Gull-like Birds - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 314 Upright Perching Waterbirds - - - - - - - - - - - - - - - - - - 316 Duck-like Birds - - - - - - - - - - - - - - - - - - - - - - - - - - - - 318 Sandpiper-like Birds- - - - - - - - - - - - - - - - - - - - - - - - - 321 Chicken-like Marsh Birds - - - - - - - - - - - - - - - - - - - - -324 Upland Ground Birds - - - - - - - - - - - - - - - - - - - - - - - - 325 Swallows and Swifts - - - - - - - - - - - - - - - - - - - - - - - - 327 Tree-Clinging Birds - - - - - - - - - - - - - - - - - - - - - - - - -329 vii SECTION IV: ZOOLOGY AND ECOLOGY OF FISH, REPTILES, AND AMPHIBIANS - - - - - - - - - - 337 CHAPTER 16 Freshwater Fish - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 338 Catfi sh - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -340 Sunfi sh - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -342 Perch - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -345 Trout, Char, and Whitefi sh - - - - - - - - - - - - - - - - - - - -347 Pike - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 352 Sturgeons - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 353 Minnows and Suckers - - - - - - - - - - - - - - - - - - - - - - -354 CHAPTER 17 Anadromous and Diadromous Fish - - - - - - - - - - - - - - - - - - - - - 359 Salmon - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 365 Steelhead - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -370 Striped Bass - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 371 Shad - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 373 Freshwater Eels - - - - - - - - - - - - - - - - - - - - - - - - - - - - 373 CHAPTER 18 Saltwater Fish and Fauna - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 378 Sharks, Skates, and Rays - - - - - - - - - - - - - - - - - - - - -379 Mackerel, Tuna, and Marlins - - - - - - - - - - - - - - - - - - 383 Flounder, Halibut, and Sole - - - - - - - - - - - - - - - - - - - 385 Herring - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -387 Codfi sh - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -389 Smelts - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -390 Sea Bass and Groupers - - - - - - - - - - - - - - - - - - - - - - 391 Mollusks - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -392 Crustaceans - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -394 CHAPTER 19 Reptiles and Amphibians - - - - - - - - - - - - - - - - - - - - - - - - - - - - -400 Reptiles - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -401 Alligators and Crocodiles - - - - - - - - - - - - - - - - - - -403 Lizards - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -405 Snakes - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -410 Turtles and Tortoises - - - - - - - - - - - - - - - - - - - - - - 416 Amphibians - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 419 Frogs and Toads - - - - - - - - - - - - - - - - - - - - - - - - 420 Salamanders and Newts - - - - - - - - - - - - - - - - - - -422 viii SECTION V: CONSERVATION AND MANAGEMENT - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -429 CHAPTER 20 Responsible Management of Wildlife Resources - - - - - - - - - - 430 Role of Private Institutions - - - - - - - - - - - - - - - - - - - 431 Farmers and Ranchers - - - - - - - - - - - - - - - - - - - - -432 Industry - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -439 Private Citizens - - - - - - - - - - - - - - - - - - - - - - - - - 440 Game Farms and Preserves - - - - - - - - - - - - - - - - -441 Role of Special Interest Groups - - - - - - - - - - - - - - - -443 Environmental Organizations - - - - - - - - - - - - - - -443 Recreational Interests - - - - - - - - - - - - - - - - - - - - 444 Role of Government - - - - - - - - - - - - - - - - - - - - - - - -445 Government Agencies - - - - - - - - - - - - - - - - - - - - 446 National Parks and Monuments- - - - - - - - - - - - - -447 National and International Law - - - - - - - - - - - - - 448 Multiple-Use Concept of Management - - - - - - - - - -450 Wildlife Careers - - - - - - - - - - - - - - - - - - - - - - - - - - - - 451 Education - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - 451 Supervised Experience Programs - - - - - - - - - - - - - 451 Employer Expectations - - - - - - - - - - - - - - - - - - - -453 Occupational Safety - - - - - - - - - - - - - - - - - - - - - -454 CHAPTER 21 Conservation of Natural Resources - - - - - - - - - - - - - - - - - - - - - 458 Conserving the Soil - - - - - - - - - - - - - - - - - - - - - - - - 460 Maintaining a Pure Water Supply- - - - - - - - - - - - - - -463 Preserving Air Quality - - - - - - - - - - - - - - - - - - - - - - -465 Preserving and Restoring Wildlife Populations and Habitats - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -467 Reclaiming Damaged or Polluted Resources - - - - - -469 CHAPTER 22 The Human Connection to Wildlife and Natural Resources - - 475 Foundation for Agriculture - - - - - - - - - - - - - - - - - - -476 Source of Materials for Medicines - - - - - - - - - - - - - -477 Mechanism for Natural Cycles - - - - - - - - - - - - - - - - -477 Recreation - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -479 Intrinsic Value - - - - - - - - - - - - - - - - - - - - - - - - - - -479 Hunting and Fishing for Sport - - - - - - - - - - - - - - -481 Game Laws and Regulations - - - - - - - - - - - - - - - -482 Using Campgrounds, Trails, Roads, and Waterways - - - - - - - - - - - - - - - - - - - - - - - - - -483 Safety Education - - - - - - - - - - - - - - - - - - - - - - - - - - -487 Basis for Commerce - - - - - - - - - - - - - - - - - - - - - - - -489 The Principle of Stewardship - - - - - - - - - - - - - - - - - 490 Ethics, Privileges, and Responsibilities - - - - - - - - - - -491 Wildlife Management - - - - - - - - - - - - - - - - - - - - - - -492 Outdoor Recreation Management - - - - - - - - - - - - - -495 Glossary - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -499 Index - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - - -508 ix

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FISH & WILDLIFE, PRINCIPLES OF ZOOLOGY AND ECOLOGY, 3rd Edition, is an introductory applied science textbook intended for use in high school programs focused on agriculture, and natural resource. The text provides a broad-spectrum overview of the wild animals of North America and the environments th
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