Fishery Data Series No. 03-03 Estimates of Chinook Salmon Abundance in the Kenai River Using Split-beam Sonar, 2001 by James D. Miller, Debby L. Burwen, and Steve J. Fleischman February 2003 Alaska Department of Fish and Game Division of Sport Fish Symbols and Abbreviations The following symbols and abbreviations, and others approved for the Système International d'Unités (SI), are used in Division of Sport Fish Fishery Manuscripts, Fishery Data Series Reports, Fishery Management Reports, and Special Publications without definition. All others must be defined in the text at first mention, as well as in the titles or footnotes of tables and in figures or figure captions. Weights and measures (metric) General Mathematics, statistics, fisheries centimeter cm All commonly accepted e.g., Mr., Mrs., alternate hypothesis H A deciliter dL abbreviations. a.m., p.m., etc. base of natural e gram g All commonly accepted e.g., Dr., Ph.D., logarithm hectare ha professional titles. R.N., etc. catch per unit effort CPUE kilogram kg and & coefficient of variation CV kilometer km at @ common test statistics F, t, c 2, etc. liter L Compass directions: confidence interval C.I. meter m east E correlation coefficient R (multiple) metric ton mt north N correlation coefficient r (simple) milliliter ml south S covariance cov millimeter mm west W degree (angular or ° Copyright ª temperature) Corporate suffixes: degrees of freedom df Weights and measures (English) Company Co. divided by ÷ or / (in cubic feet per second ft3/s Corporation Corp. equations) foot ft Incorporated Inc. equals = gallon gal Limited Ltd. expected value E inch in et alii (and other et al. fork length FL mile mi people) greater than > ounce oz et cetera (and so forth) etc. greater than or equal to ‡ pound lb exempli gratia (for e.g., harvest per unit effort HPUE quart qt example) less than < yard yd id est (that is) i.e., less than or equal to £ Spell out acre and ton. latitude or longitude lat. or long. logarithm (natural) ln monetary symbols $, ¢ logarithm (base 10) log Time and temperature (U.S.) logarithm (specify base) log etc. 2, day d months (tables and Jan,...,Dec mideye-to-fork MEF degrees Celsius °C figures): first three minute (angular) ' letters degrees Fahrenheit °F number (before a # (e.g., #10) multiplied by x hour (spell out for 24-hour clock) h number) not significant NS minute min pounds (after a number) # (e.g., 10#) null hypothesis HO second s registered trademark (cid:226) percent % Spell out year, month, and week. trademark (cid:228) probability P United States U.S. probability of a type I a Physics and chemistry (adjective) error (rejection of the all atomic symbols United States of USA null hypothesis when alternating current AC America (noun) true) ampere A U.S. state and District use two-letter probability of a type II b calorie cal of Columbia abbreviations error (acceptance of direct current DC abbreviations (e.g., AK, DC) twhhee nnu flal lhsyep) othesis hertz Hz second (angular) " horsepower hp standard deviation SD hydrogen ion activity pH standard error SE parts per million ppm standard length SL parts per thousand ppt, ‰ total length TL volts V variance Var watts W FISHERY DATA SERIES NO. 03-03 ESTIMATES OF CHINOOK SALMON ABUNDANCE IN THE KENAI RIVER USING SPLIT-BEAM SONAR, 2001 by James D. Miller and Debby L. Burwen Division of Sport Fish, Anchorage and Steve J. Fleischman Division of Sport Fish, Research and Technical Services, Anchorage Alaska Department of Fish and Game Division of Sport Fish, Research and Technical Services 333 Raspberry Road, Anchorage, Alaska, 99518-1599 February 2003 This investigation was partially financed by the Federal Aid in Sport Fish Restoration Act (16 U.S.C. 777-777K) under project F-10-17, Job No. S-2-5b. The Fishery Data Series was established in 1987 for the publication of technically-oriented results for a single project or group of closely related projects. Fishery Data Series reports are intended for fishery and other technical professionals. Fishery Data Series reports are available through the Alaska State Library and on the Internet: http://www.sf.adfg.state.ak.us/statewide/divreports/html/intersearch.cfm This publication has undergone editorial and peer review. James D. Miller, Debby L. Burwen, Alaska Department of Fish and Game, Division of Sport Fish and Steve J. Fleischman Alaska Department of Fish and Game, Division of Sport Fish, Research and Technical Services 333 Raspberry Road, Anchorage, Alaska 99518-1599, USA This document should be cited as: Miller, J. D., D. L. Burwen, and S. J. Fleischman. 2003. Estimates of chinook salmon abundance in the Kenai River using split-beam sonar, 2001. Alaska Department of Fish and Game, Fishery Data Series No. 03-03, Anchorage. The Alaska Department of Fish and Game administers all programs and activities free from discrimination based on race, color, national origin, age, sex, religion, marital status, pregnancy, parenthood, or disability. The department administers all programs and activities in compliance with Title VI of the Civil Rights Act of 1964, Section 504 of the Rehabilitation Act of 1973, Title II of the Americans with Disabilities Act of 1990, the Age Discrimination Act of 1975, and Title IX of the Education Amendments of 1972. If you believe you have been discriminated against in any program, activity, or facility, or if you desire further information please write to ADF&G, P.O. Box 25526, Juneau, AK 99802-5526; U.S. Fish and Wildlife Service, 4040 N. Fairfax Drive, Suite 300 Webb, Arlington, VA 22203 or O.E.O., U.S. Department of the Interior, Washington DC 20240. For information on alternative formats for this and other department publications, please contact the department ADA Coordinator at (voice) 907-465-4120, (TDD) 907-465-3646, or (FAX) 907-465-2440. TABLE OF CONTENTS Page LIST OF TABLES..........................................................................................................................................................................iii LIST OF FIGURES........................................................................................................................................................................iv LIST OF APPENDICES...............................................................................................................................................................vi ABSTRACT ......................................................................................................................................................................................1 INTRODUCTION............................................................................................................................................................................1 METHODS........................................................................................................................................................................................5 Study Area..........................................................................................................................................................................................5 Site Description.................................................................................................................................................................................5 Acoustic Sampling............................................................................................................................................................................5 Sonar System Configuration......................................................................................................................................................7 Bottom Mapping and Beam Coverage...................................................................................................................................10 System Calibration....................................................................................................................................................................11 Sampling Procedure..................................................................................................................................................................11 Echo Sounder Settings..............................................................................................................................................................11 Data Acquisition........................................................................................................................................................................11 Fish Tracking and Echo Counting...............................................................................................................................................14 Data Analyses..................................................................................................................................................................................14 Tidal and Temporal Distribution............................................................................................................................................14 Spatial Distribution...................................................................................................................................................................15 Target Strength Distribution....................................................................................................................................................16 Species Discrimination.............................................................................................................................................................16 Passage Estimates......................................................................................................................................................................16 Net-apportioned Estimates............................................................................................................................................................19 RESULTS........................................................................................................................................................................................20 System Calibration.........................................................................................................................................................................20 Beam Coverage...............................................................................................................................................................................20 Target Tracking...............................................................................................................................................................................20 Tidal and Temporal Distribution..................................................................................................................................................20 Spatial Distribution.........................................................................................................................................................................21 Vertical Distribution.................................................................................................................................................................21 Range Distribution....................................................................................................................................................................22 Target Strength...........................................................................................................................................................................26 Passage Estimates......................................................................................................................................................................30 DISCUSSION.................................................................................................................................................................................33 Spatial Distribution.........................................................................................................................................................................33 Bank Preference.........................................................................................................................................................................33 Vertical Distribution.................................................................................................................................................................39 Range Distribution....................................................................................................................................................................39 Target Strength................................................................................................................................................................................40 Direction of Travel.........................................................................................................................................................................41 Comparison of Passage Estimates with Other Indices.............................................................................................................41 Early Run....................................................................................................................................................................................42 Late Run......................................................................................................................................................................................45 Outlook.............................................................................................................................................................................................48 i TABLE OF CONTENTS (Continued) Page Inriver Netting............................................................................................................................................................................51 Pulse Width................................................................................................................................................................................51 Imaging Sonar............................................................................................................................................................................52 ACKNOWLEDGMENTS.............................................................................................................................................................52 LITERATURE CITED..................................................................................................................................................................53 APPENDIX A. TARGET STRENGTH ESTIMATION........................................................................................................57 APPENDIX B. SYSTEM PARAMETERS..............................................................................................................................59 APPENDIX C. DATA FLOW....................................................................................................................................................67 APPENDIX D. DAILY PROPORTIONS OF UPSTREAM AND DOWNSTREAM FISH FOR THE 2001 EARLY AND LATE KENAI RIVER CHINOOK SALMON RUNS..................................................................................69 APPENDIX E. AVERAGE VERTICAL ANGLE BY TIDE STAGE, RUN, BANK, AND FISH ORIENTATION (UPSTREAM OR DOWNSTREAM) FOR THE 2001 KENAI RIVER CHINOOK SALMON RUNS................................................................................................................................................................................................73 APPENDIX F. HISTORIC ESTIMATES OF INRIVER RETURN BY YEAR AND DATE (1987–2001)...............77 APPENDIX G. FILTERED (CONVENTIONAL), UNFILTERED, AND NET-APPORTIONED CHINOOK PASSAGE ESTIMATES, KENAI RIVER SONAR, EARLY AND LATE RUNS, 2001...............................................81 ii LIST OF TABLES Table Page 1. Principal components of the split-beam sonar system used in 2001........................................................................7 2. HTI model 244 digital echo sounder settings used in 2001....................................................................................11 3. Echo acceptance criteria for digital echo processing, 2001.....................................................................................13 4. Results of 2001 in situ calibration verifications using a 38.1 mm tungsten carbide standard sphere..............21 5. Estimates of chinook salmon passage by tide stage and direction of travel for the 2001 early run (16 May to 30 June).......................................................................................................................................................22 6. Estimates of chinook salmon passage by tide stage and direction of travel for the 2001 late run (1 July to 10 August)...................................................................................................................................................................22 7. Estimates of early-run fish passage by direction of travel, 2001............................................................................32 8. Estimates of late-run fish passage by direction of travel, 2001..............................................................................32 9. Mean target strength (dB) for upstream and downstream targets by bank (chinook only) during the early (16 May-30 June) and late (1 July-10 August) runs, 2001............................................................................32 10. Estimated daily upstream passage of chinook salmon, Kenai River sonar, early run, 2001.............................35 11. Estimated daily upstream passage of chinook salmon, Kenai River sonar, late run, 2001................................36 iii LIST OF FIGURES Figure Page 1. Cook Inlet showing location of the Kenai River.........................................................................................................2 2. The Kenai River showing location of chinook salmon sonar site, 2001.................................................................6 3. Cross-sectional (top) and aerial (bottom) views of sonar site showing insonified portions of the Kenai River, 2001.........................................................................................................................................................................8 4. Daily right- and left-bank transducer locations and counting ranges relative to bipod tower located on the right bank, Kenai River, 2001..................................................................................................................................9 5. Left- and right-bank bottom profiles at the Kenai River chinook sonar site with approximate transducer locations and sonar beam coverage for 16 May 2001..............................................................................................10 6. Schematic diagram of 2001 split-beam sonar system configuration and data flow............................................12 7. Distribution of upstream and downstream fish by tide stage during the early and late runs, Kenai River, 2001...................................................................................................................................................................................23 8. Vertical distributions of early-run upstream and downstream fish on the left and right banks, Kenai River, 2001.......................................................................................................................................................................24 9. Vertical distributions of early-run upstream fish during falling, low, and rising tide stages on the left and right banks, Kenai River, 2001.............................................................................................................................25 10. Vertical distributions of late-run upstream and downstream fish on the left and right banks, Kenai River, 2001.......................................................................................................................................................................26 11. Vertical distributions of late-run upstream fish during falling, low, and rising tide stages on the left and right banks, Kenai River, 2001.....................................................................................................................................27 12. Range distributions of early-run upstream and downstream fish on the left and right banks, Kenai River, 2001.......................................................................................................................................................................28 13. Range distributions of early-run upstream fish during falling, low, and rising tide stages on the left and right banks, Kenai River, 2001.....................................................................................................................................29 14. Range distributions of late-run upstream and downstream fish on the left and right banks, Kenai River, 2001...................................................................................................................................................................................30 15. Range distributions of late-run upstream fish during falling, low, and rising tide stages on the left and right banks, Kenai River, 2001.....................................................................................................................................31 16. Early-run target strength distributions for all upstream and downstream targets on the left and right banks, Kenai River, 2001..............................................................................................................................................33 17. Late-run target strength distributions for all upstream and downstream targets on the left and right banks, Kenai River, 2001..............................................................................................................................................34 18. Daily sonar estimates of passage by bank (top panel), total passage (center panel), and historical cumulative proportions for the early run of chinook salmon returning to the Kenai River, 2001....................37 19. Daily sonar estimates of passage by bank (top panel), total passage (center panel), and historical cumulative proportions for the late run of chinook salmon returning to the Kenai River, 2001......................38 20. Daily CPUE of chinook and sockeye salmon from inriver gillnetting, 16 May-10 August, 2001...................42 21. Range distribution of all (unfiltered) upstream targets on the left and right banks during the early run, 16-30 May, 31 May-15 June, and 16-30 June, Kenai River, 2001........................................................................43 22. Daily discharge rates at the Soldotna Bridge and Secchi depth readings in front of the sonar site, Kenai River, early run (16 May–30 June), 2001...................................................................................................................44 23. Daily sonar estimates and inriver gillnet CPUE of chinook salmon during the early run (16 May–30 June), Kenai River, 2001...............................................................................................................................................45 24. Filtered (conventional), unfiltered, and net-apportioned estimates of chinook salmon passage, for the early and late runs, Kenai River sonar, 2001.............................................................................................................46 25. Daily chinook salmon sonar estimates with chinook salmon sport fish CPUE (open triangles represent days on which only unguided anglers were allowed to fish), early run (16 May-30 June), 2001....................47 26. Daily chinook sonar estimates and river mile-19 sockeye sonar estimates lagged one day, late run (1 July–10 August), 2001...................................................................................................................................................48 27. Range distribution of all (unfiltered) upstream targets on the left and right banks during the late run, 1- 13 July, 14-31 July, and 1-10 August, Kenai River, 2001.......................................................................................49 28. Daily discharge rates at the Soldotna Bridge and Secchi depth readings in front of the sonar site, Kenai River, late run (1 July–10 August), 2001...................................................................................................................50 iv LIST OF FIGURES (Continued) Figure Page 29. Daily sonar estimates and inriver gillnet CPUE for chinook salmon during the late run (1 July–10 August), 2001..................................................................................................................................................................50 30. Cumulative sonar estimates with cumulative inriver net CPUE, late-run chinook salmon (1 July–10 August), 2001..................................................................................................................................................................51 31. Daily chinook salmon sonar estimates with chinook salmon sport fish CPUE (open triangles represent days on which only unguided anglers were allowed to fish), late run (1 July-31 July), 2001..........................52 v LIST OF APPENDICES Appendix Page A1. Using the sonar equation to estimate target strength with dual- and split-beam applications...........................58 B1. Example of system parameters used for data collection on the right bank (transducer 733)............................60 B2. Example of system parameters used for data collection on the left bank (transducer 738)...............................63 C1. Inseason data flow diagram for the Kenai River chinook salmon sonar project, 2001......................................68 D1. Daily proportions of upstream and downstream fish for the 2001 Kenai River early chinook run..................70 D2. Daily proportions of upstream and downstream fish for the 2001 Kenai River late chinook run....................71 E1. Average vertical angle by tide stage and orientation for the 2001 Kenai River early chinook run, 2001.......74 E2. Average vertical angle by tide stage and orientation for the 2001 Kenai River late chinook run....................75 F1. Kenai River early-run chinook salmon sonar estimates of inriver return, 1987-2001........................................78 F2. Kenai River late-run chinook salmon sonar estimates of inriver return, 1987-2001..........................................79 G1. Filtered (conventional), unfiltered, and net-apportioned chinook passage estimates, Kenai River sonar, early run, 2001.................................................................................................................................................................82 G2. Filtered (conventional), unfiltered, and net-apportioned chinook passage estimates, Kenai River sonar, late run, 2001...................................................................................................................................................................83 vi
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