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Descriptions of four new species of Chrysotimus Loew from Tibet (Diptera: Dolichopodidae) PDF

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©EntomologicaFennica.16June2006 Descriptions of four new species of Chrysotimus Loew from Tibet (Diptera: Dolichopodidae) MengqingWang&DingYang* Wang,M.&Yang,D.2006:DescriptionsoffournewspeciesofChrysotimus LoewfromTibet(Diptera:Dolichopodidae).—Entomol.Fennica17:98–104. TheoccurrenceofthegenusChrysotimusLoew,1857inTibetisconfirmedfor thefirsttime.Thefollowingfourspeciesaredescribedasnewtoscience:Chryso- timusbifurcatussp.n.,C.grandissp.n.,C.liisp.n.andC.linzhiensissp.n.Akey isgiventodistinguishtheChinesespeciesofthegenus. M. Wang and D. Yang (correspondent author), Department of Entomology, ChinaAgriculturalUniversity,Beijing100094,China Received26January2005,accepted15July2005 1.Introduction Inthispaper,wedescribefourspeciesnewto science,allfoundfromTibet,anarealyinginthe The genus Chrysotimus Loew, 1857 belongs to SouthwestofChina,onthenorthernsideofHi- thesubfamilyPeloropeodinae.Thegenusisgen- malayas, and bordered with Pakistan, India, erally characterized by the yellow or yellowish Burma,Nepal,andBhutan.Theareabelongsto hairs and bristles, small first flagellomere, hind thePalaearcticandOrientalRealms.Perhapsthe tarsomere1inmostmalesbearingseveralshort high-altitude climate is responsible for the re- erect black ventral bristles at base, and mid strictedoccurrenceofthesefourspecies.Wealso tarsomere1inmaleatleastaslongastarsomeres provideakeytoChrysotimusspeciesmalesfrom 2–4.60specieswereknowntooccurintheHol- China. arctic and Oriental Realms. Previously, there were 8 species known from the Palaearctic (Negrobov1991)and2speciesfromtheOriental 2.Materialandmethods Realms(Becker1922,Dyte1975).Sofar25spe- cieshavebeenknownfromChina(Parent1944, The specimens for this study were collected by Wang et al. 2005, Yang 2001, Yang & Saigusa Fasheng Li (Beijing) from Tibet in 1978 by 2001a,2001b,2005,Zhangetal.2003). sweepnetting,andaredepositedintheEntomo- Guzeriplia Negrobov, 1968 embodies the logicalMuseumofChinaAgriculturalUniversity charactersofChrysotimusLoewintheheadand (CAU),Beijing.Allmaterialispreservedin75% thoraxwithyellowhairsandbristlesandbiseriate alcohol. acr.However,inGuzeriplia,thehypopygiumis Abbreviations are as follows: acr = acro- large (almost as long as abdomen), bearing the stichal,ad=anterodorsal,av=anteroventral,dc= long surstylus and cercus. Bickel (2004) noted dorsocentral,LI=foreleg,LII=midleg,LIII= thatitshouldbewithintherangeofvariationof hindleg,oc=ocellar,pd=postero-dorsal,ph= thelatter,probablyasasynonymy. posthumeral,pv=posteroventral,v=ventral. ENTOMOL.FENNICAVol.17 (cid:127) FournewspeciesofChrysotimusfromTibet 99 3.KeytospeciesofChrysotimusLoew Hairs and bristles on thorax brownish or fromChina(males) brown 13 12.Foretarsomere1withrowofabout10v;hind 1. Hind tarsomere 1 at most with sparse black tarsomere1with22shortblackventralbris- bristlesatbase 2 tlesonbasal1/4;surstylusbasallywithoutin- Hindtarsomere1withbundle(s)ofblackven- nerprocess.Gansu tralbristlesatbase 8 chikuniWang,Yang&Grootaert,2005 2. Hindtarsomere1withoutblackventralbris- Fore tarsomere 1 without row of v; hind tlesatbase(notknowningrandis) 3 tarsomere 1 with less than 20 black ventral Hindtarsomere1withblackventralbristlesat bristlesonbasal1/4;surstylusbasallywithin- base 5 nerprocess.Hubei,Shaanxi,Yunnan 3. Cercusdividedintoouterandinnerlobes 4 shennongjianusYang&Saigusa,2001 Cercussingle.China. 13.Hind tarsomere 1 with about 12 short black sinensisParent,1944 ventral bristles; surstylus not furcated api- 4. Acrabsent;midtarsomere1shorterthanthe cally.Yunnan tarsomeres 2–5; epandrium with nearly bispinusYang&Saigusa,2001 quadrate lateral process. Guangxi Hindtarsomere1with15–16shortblackven- guangxiensisYang&Saigusa,2001 tral bristles; surstylus furcated apically. Six to seven irregularly paired acr; mid Guangxi tarsomere 1 longer than the tarsomeres 2–5; xuaeWang,Yang&Grootaert,2005 epandrium with long hook-like lateral pro- 14.Acr2–4pairs 15 cess.Tibet. grandissp.n. Acrmorethan5pairs 22 5. Antenna with scape and pedicel yellow. 15.Hind tarsomere 1 with group of 8–12 black Zhejiang basiflavusYang,2001 basal bristles at base (which are somewhat Antennawhollyblack. 6 sparse),butwithoutdistinctv. 16 6. Fivedc;acrabsent.Zhejiang Hind tarsomere 1 with 1 (or 2) bundles of apicicurvatusYang,2001 blackbasalventralbristles,androwof7–8pv Sixdc;acrpresent 7 18 7. Four to five irregularly paired acr; hind 16.CuAx ratio about 0.2; lateral process on tarsomere1with6sparseblackventralbris- epandriumnotconcavenearmiddle 17 tlesonbasal1/4.Guangdong CuAx ratio 0.35; lateral process on acutatusWang,Yang&Grootaert,2005 epandrium concave near middle. Yunnan Nine to ten irregularly paired acr; hind yunlonganusYang&Saigusa,2001 tarsomere1with2sparseblackventralbris- 17.Firstflagellomereaslongaswide;hindtibia tlesatbase.Ningxia withoutdistinctv.Tibet liisp.n. ningxianusWang,Yang&Grootaert,2005 Firstflagellomereabout1.5timeswiderthan 8. Fourorfivedc;acrabsent 9 long;hindtibiawith2pv.Tibet Sixdc;acrpresent 14 linzhiensissp.n. 9. Aristadorsal;hypandriumwithbroadlateral 18.R4+5andMparallelapically;hindtarsomere process.Zhejiang 1withbundleof4–5blackventralbristlesat dorsalisYang,2001 base 19 Aristasub-apical;hypandriumwiththinfin- R4+5 and M slightly convergent apically; ger-likelateralprocess 10 hindtarsomere1with2bundlesof3–4black 10.Hindtarsomere1with3–4shortblackspine- ventralbristlesatbase.Shaanxi like ventral bristles at base. Beijing bifasciaYang&Saigusa,2005 songshanusWang,Yang 19.Hairs and bristles on thorax yellow or pale; &Grootaert,2005 surstylusonepandriumnotfurcatedapically Hind tarsomere 1 with about (or more than) 20 10blackventralbristlesatbase 11 Hairsandbristlesonthoraxbrown;surstylus 11.Hairsandbristlesonthoraxyellow 12 on epandrium furcated apically. Guangxi 100 Wang&Yang (cid:127) ENTOMOL.FENNICAVol.17 sanjiangyuanusWang,Yang tarsomere1with15–16blackventralbristles. &Grootaert,2005 Yunnan pingbianusYang&Saigusa,2001 20.Fore tarsomere 1 without rowed v; hind 28.Hindtarsomere1with12shortblackventral tarsomere1with4–5blackventralbristlesat bristles;8irregularlypairedacr;cercuswith base 21 round apex; palpus dark brown. Yunnan. Fore tarsomere 1 with row of 5–6 v; hind incisusYang&Saigusa,2001 tarsomere 1 with 8 black ventral bristles at Hindtarsomere1with7–8shortblackventral base.Guangdong bristles;6irregularlypairedacr;cercuswith guangdongensisWang,Yang acute apex; palpus yellow. Yunnan &Grootaert,2005 lijianganusYang&Saigusa,2001 21.Hind femur with row of ad and pd; cercus longandnarrow.Beijing xiaolongmensisZhang,Yang 4.Descriptionsofnewspecies &Grootaert,2003 Hind femur without distinctly d; cercus Chrysotimusbifurcatussp.n.(Fig.1) round.Shaanxi unifasciaYang&Saigusa,2005 Material examined. Holotype #: China: Tibet, 22.Hindtarsomere1withrowofabout10pv. 23 Bomi,Jieda,3050ma.s.l.,16.VII.1978,leg.F.Li Hindtarsomere1withoutdistinctv 26 (Beijing). 23.Hind tarsomere 1 with 10–12 black ventral Diagnosis.Allcoxaeyellow.6–7irregularly basalbristles;midtibiawithoutdistinctv 24 pairedacrshortandhair-like. Hindtarsomere1 Hindtarsomere1withabout20blackventral withgroupof10–12shortblackventralbristles basalbristles;midtibiawith1pv. 25 on basal 1/5, and row of 8–9 pv. Cercus bifur- 24.Firstflagellomeresomewhatround,2.0times cated. wider than long; cercus not furcated, dorsal Description.Male.Headmetallicgreenwith lobeonsurstylusthickandstraight.Shaanxi qinlingensisYang&Saigusa,2005 Firstflagellomeresubtriangular,aboutaslong aswide;cercusbifurcated,dorsallobeonsur- stylusthinandcurved.Tibet bifurcatussp.n. 25.Foreandmidtarsomere1withoutdistinctv; epandrium basally with short process. Shaanxi setosusYang&Saigusa,2005 Foreandmidtarsomere1eachwithrowof5– 6v;epandriumbasallywithlongbroadpro- cess.Guangdong xiaohuangshanusWang,Yang &Grootaert,2005 26.Hindtarsomere1with1bundleofshortblack ventral bristles; surstylus rather wide and thick;hypandriumwithsmallapicalincision 27 Hindtarsomere1with2–3bundlesof14–15 blackventralbristlesonbasal1/6(whichcon- tains1–2smalltightbundlesand1largeloose bundle);surstylusnarrowfinger-like;hypan- drium without apical incision. Yunnan digitatusYang&Saigusa,2001 Fig.1.Chrysotimusbifurcatussp.n.,male.a.First 27.Hairsandbristlesonthoraxpaleoryellow 28 flagellomere,lateralview.b.Wing.c.Genitalia,lateral Hairsandbristlesonthoraxdarkbrown;hind view. ENTOMOL.FENNICAVol.17 (cid:127) FournewspeciesofChrysotimusfromTibet 101 palegraypollen.Hairsandbristlesonheadyel- about 10 black basal ventral bristles on hind low.Ocellartubercleweak,with2verylongoc tarsomere1,butcanbeseparatedfromthelatter and2veryshortposteriorhairs.Antennabrown; bytheepandriumwithslendersurstylusandbi- pedicel with circle of brown apical hairs; first furcated cercus. In qinlingensis, the surstylus is flagellomere subtriangular (Fig. 1a), about 0.6 thickandstraight,andthecercusisnotfurcated timesaslongaswide;aristadorsal,brown,with (Yang&Saigusa2005). veryshortpubescence.Proboscisblackish,with Distribution.KnownonlyfromTibet. brownishhairs;palpusyellow,withyellowhairs and2yellowapicalbristles. Thoraxmetallicgreenwithpalegraypollen, Chrysotimusgrandissp.n.(Fig.2) mesonotum and scutellum brilliant. Hairs and bristlesonthoraxyellow;6strongdc,6–7irregu- Material examined. Holotype #: Tibet: Bomi, larly paired acr, short and hair-like; scutellum Jieda,3700ma.s.l.,12.VIII.1978,leg.F.Li(Bei- with2pairsofbristles.Propleuronwith1yellow- jing). ishhairand1yellowishbristleonlowerportion. Diagnosis.Palpusblackish.6strongdc,6–7 Legsyellow,all5thtarsomeresbrown.Hairs irregularly paired acr short and hair-like. Mid andbristlesonlegsyellow;coxaewithyellowish tarsomere1longerthanthetotallengthofcorre- hairsandbristles;forecoxawith6–8yellowbris- spondingtarsomeres2–5.Surstylusofepandrium tles, mid coxa with 2–4 anterior bristles, hind withlargeswollenapex. coxawith1yellowouterbristlenearmiddle.Fore Description.Male.Headmetallicgreenwith femur with 2 av apically, mid and hind femora palegraypollen.Hairsandbristlesonheadyel- eachwith1avand1pvapically.Firsttibiawith rowofshortpd,apicallywith3shortbristles;mid tibiawith2adand2–3pd,apicallywith4bristles; hindtibiawith1adand2pd,apicallywith3bris- tles.Midtarsomere1with2–3v,hindtarsomere1 withgroupof10–12shortblackventralbristles onbasal1/5,androwof8–9pv.Relativelengths oftibiaand5tarsomeresoflegsLI2.8:1.5:0.6: 0.5:0.3:0.4;LII3.5:1.8:0.7:0.6:?:?(the questionmarksmeanbrokensegments);LIII3.8: 1.4:0.9:0.6:0.3:0.4. Winghyaline(Fig.1b);veinsbrownish,R4+5 andMparallelapically;CuAxratio(lengthofm- cu/lengthofdistalportionofCuA)0.24.Squama brownishyellowwithpalehairs.Halteryellow. Abdomenmetallicgreenwithpalegraypol- len,dorsumbrilliant.Hairsandbristlesonabdo- menyellow. Malegenitalia(Fig.1c).Epandriumdistinctly longerthanwide,withroundleaf-likelateralpro- cess; surstylus slender, with acute apex; cercus somewhatround,bifurcatedapically,onebranch shortandround,anotherbranchlongfinger-like; hypandriumshortandbroad. Size.Body1.3mm,wing1.7mm. Female.Unknown. Remarks. Chrysotimus bifurcatus sp. n. is Fig.2.Chrysotimusgrandissp.n.,male.a.Apicalhalf similar to Chrysotimus qinlingensis Yang & ofwing.b.Genitalia,lateralview.c.Genitalia,ventral Saigusa from Qinling in having row of pv and view. 102 Wang&Yang (cid:127) ENTOMOL.FENNICAVol.17 Fig.3.Chrysotimusliisp.n.,male.a.Firstflagellomere(notincludingapicalsegmentofarista), lateralview.b.Wing.c.Genitalia,lateralview. low.Ocellartubercleweak,with2longocand2 Female.Unknown. shortposteriorhairs.Antennabroken.Proboscis Remarks.Chrysotimusgrandissp.n.issome- blackish, with brownish hairs; palpus blackish, what similar to Chrysotimus ningxianus Wang, withyellowhairs. Yang & Grootaert from Ningxia in having the Thoraxmetallicgreenwithpalegraypollen, longmidtarsomere1,butcanbeseparatedfrom mesonotum and scutellum brilliant. Hairs and the latter by the following features: 6–7 irregu- bristlesonthoraxyellow;6strongdc,6–7irregu- larly paired acr, mid tarsomere 1 longer than larlypairedacrshortandhair-like;scutellumwith tarsomeres2–5,andsurstyluslargewithswollen 2 pairs of bristles. Propleuron with 1 yellowish apex;inningxianus,9–10irregularlypairedacr, hairand1yellowishbristleonlowerportion. midtarsomere1nearlyaslongastarsomeres2–5, Legs [fore tarsus, hind tibia and tarsus bro- andsurstylusslender(Wangetal.2005). ken]yellow;midtarsomere5brown. Hairsand Distribution.KnownonlyfromTibet. bristles on legs yellow; coxae with yellowish hairsandbristles;forecoxawith4–6yellowbris- tles, mid coxa with 2–4 anterior bristles, hind Chrysotimusliisp.n.(Fig.3) coxawith1yellowouterbristlenearmiddle.Mid femurwith1avand1pvatapex.Midtibiawith2 Material examined. Holotype #: Tibet: Bomi, ad and 3 pd, apically with 4 bristles. Mid Jieda, 3050m a.s.l., 16.VII.1978, leg. F. Li. tarsomere1longerthanthetotallengthofcorre- Paratypes 2 ##: Tibet: Linzhi, 1.–3.VI.1978, sponding tarsomeres 2–5. Relative lengths of leg.F.Li(Beijing). tibiaand5tarsomeresoflegsLI3.2:?:?:?:?:?; Diagnosis. First flagellomere subtriangular, LII4.0:2.7:1.0:0.6:0.3:0.5. aslongaswide.Hindtarsomere1withgroupof Winghyaline(Fig.2a),veinsbrownish,R4+5 10–11shortblackventralbristlesonbasal1/6. andMparallelapically;CuAxratio0.2.Squama Description.Male.Headmetallicgreenwith brownishyellowwithpalehairs.Halteryellow. palegraypollen.Hairsandbristlesonheadyel- Abdomenmetallicgreenwithpalegraypol- low.Ocellartubercleweak,with2longocand2 len,dorsumbrilliant.Hairsandbristlesonabdo- short posterior hairs. Antenna brown; pedicel menyellow. with circle of brownish apical hairs; first Male genitalia (Figs 2b–c). Epandrium dis- flagellomere subtriangular (Fig. 3a), as long as tinctlylongerthanwide,somewhatsquare,with wide; arista [apical segment broken] dorsal, long hook-like lateral process; surstylus with brownish,withveryshortpubescence.Proboscis largedorsallobe,thickbaseanddistinctlyswol- blackish, with brownish hairs; palpus pale yel- len apex, and long curved ventral lobe; cercus low,withyellowhairsand2brownishapicalbris- longandbifurcated,outerlobewithapexwider tles. thanbase,withseveral(about7–8)palelongsoft Thoraxmetallicgreenwithpalegraypollen, apical hairs, inner lobe long and acute; mesonotum and scutellum brilliant, pleuron hypandriumthinandshort,symmetricalinven- brownwithpalepollen.Hairsandbristlesontho- tralview. raxyellow;6strongdc,3–4irregularlypairedacr Size.Body1.3mm,wing1.6mm. shortandhair-like;scutellumwith2pairsofbris- ENTOMOL.FENNICAVol.17 (cid:127) FournewspeciesofChrysotimusfromTibet 103 tles.Propleuronwith1yellowishhairand1yel- lowishbristleonlowerportion. Legsyellow,all5thtarsomeresbrown.Hairs andbristlesonlegsyellow;coxaewithyellowish hairsandbristles;forecoxawith2yellow bris- tles, mid coxa with 2–4 anterior bristles, hind coxa with 1 yellow outer bristle at middle. Mid andhindfemoraeachwith1avand1pvatapex. Midtibiawith2adand2pd;hindtibiawith2ad. Hindtarsomere1withgroupof10–11shortblack ventralbristlesonbasal1/6.Relativelengthsof tibiaand5tarsomeresoflegsLI3.7:2.0:0.9:0.6 :0.4:0.5;LII4.7:2.6:1.0:0.7:0.4:0.4;LIII5.0 :2.3:1.1:0.8:0.5:0.5. Winghyaline(Fig.3b);veinsbrownish,R4+5 andMparallelapically;CuAxratio0.22.Squama brownishyellowwithpalehairs.Halteryellow. Fig.4.Chrysotimuslinzhiensissp.n.,male.a.Wing. b.Firstflagellomere,lateralview.c.Genitalia,lateral Abdomenmetallicgreenwithpalegraypol- view. len,dorsumbrilliant.Hairsandbristlesonabdo- menyellow. Malegenitalia(Fig.3c).Epandriumdistinctly low.Ocellartubercleweak,with2verylongoc longer than wide, with wide lateral process; and2veryshortposteriorhairs.Antennabrown; surstyluswithroundapex,with3longapicalbris- pedicel with circle of brown apical hairs; first tles; cercus long and rather narrow apically; flagellomere rather wide (Fig. 4b), about 1.5 hypandriumshortandbroadwithroundapex. timeswiderthanlong;aristadorsal,brown,with Size.Body1.6mm,wing2.5mm. short pubescence. Proboscis blackish, with Female.Unknown. brownishhairs;palpuspaleyellow,withyellow Remarks. Chrysotimus lii sp. n. is similar to hairsand2yellowapicalbristles. Chrysotimus bifascia Yang & Saigusa from Thoraxmetallicgreenwithpalegraypollen, Qinling in having the pale yellow palpus and mesonotum and scutellum brilliant, pleuron brown5thtarsomere,butcanbeseparatedfrom brownwithpalepollen.Hairsandbristlesontho- the latter by the hind tarsomere 1 with 10–11 raxyellow;6strongdc(thefirstdcshorter),4ir- black ventral bristles at base and the first regularly paired acr, short and hair-like; flagellomereaslongaswide.Inbifascia,thehind scutellumwith2pairsofbristles.Propleuronwith tarsomere 1 just has 3–4 black ventral bristles, 1yellowishhairand1yellowishbristleonlower and the first flagellomere is about 0.6 times as portion. longaswide(Yang&Saigusa2005). Legsyellow,all5thtarsomeresbrown.Hairs Distribution.KnownonlyfromTibet. andbristlesonlegsyellow;coxaewithyellowish hairsandbristles;forecoxawith4yellow bris- tles, mid coxa with 2–4 anterior bristles, hind Chrysotimuslinzhiensissp.n.(Fig.4) coxa with 1 yellow outer bristle at middle. Mid andhindfemoraeachwith1avand1pvapically. Material examined. Holotype #: Tibet: Linzhi, Midtibiawith2adand1pd;hindtibiawith1–2 3050ma.s.l.,3.VI.1978,leg,F.Li(Beijing). ad,1pdand2pv.Hindtarsomere1withgroupof Diagnosis. Palpus pale yellow. Legs yellow 13shortblackventralbristlesonbasal1/6.Rela- with 5th tarsomeres brown. Hind tarsomere 1 tivelengthsoftibiaand5tarsomeresoflegsLI with group of 13 short black ventral bristles on 4.0:2.1:1.0:0.6:0.4:0.5;LII4.7:2.7:1.1:0.8 basal1/6. :0.4:0.4;LIII4.8:2.1:?:?:?:?. Description.Male.Headmetallicgreenwith Winghyaline(Fig.4a),veinsbrownish,R4+5 palegraypollen.Hairsandbristlesonheadyel- andMparallelapically;CuAxratio0.25.Squama 104 Wang&Yang (cid:127) ENTOMOL.FENNICAVol.17 brownishyellowwithpalehairs.Halteryellow. Bickel,D.J.2004:Alishania,anewgenuswithremarkable Abdomenmetallicgreenwithpalegreypol- female terminalia from Taiwan, with notes on Chrysotimus Loew (Diptera: Dolichopodidae). — len,dorsumbrilliant.Hairsandbristlesonabdo- BishopMus.Bull.Ent.12:27–34. menyellow. Dyte,C.E.1975:FamilyDolichopodidae.—In:Delfi- Male genitalia (Figs. 4c). Epandrium dis- nado,M.D.&Hardy,D.E.(eds.),Acatalogofthe tinctlylongerthanwidewithwidelateralprocess; Diptera of the Oriental region, 2: 212–258. Hawaii surstyluscurveddownward,roundapically,with UniversityPress,Honolulu.618pp. 3apicalbristles;cercusroundwithyellowmar- Loew,H.1857:NeueBeitrezurKenntnissderDipteren.F fter Beitrag. — Programm K. Realschule Meserit, ginal bristles and hairs; hypandrium short and 1857:1–6. broad. Negrobov, O. P. 1991: Family Dolichopodidae. — In: Size.Body1.8mm,wing2.6mm. Soós,Á.&Papp,L.(eds.),CatalogueofPalaearctic Female.Unknown. Diptera7:11–139.AkadiaiKiadBudapest.291pp. Remarks. Chrysotimus linzhiensis sp. n. is Parent, O. 1944: Diptères Dolichopodidae recueillis en similar to Chrysotimus yunlonganus Yang & ChinedeNord,enMongolieetenMandchourieparle P.P.E.Licent.—Rev.Fr.Ent.10(4):121–131. Saigusa from Yunnan in having the short first Wang,M.,Yang,D.&Grootaert,P.2005:Chrysotimus flagellomereand3–4pairedacr,butcanbesepa- Loew from China (Diptera: Dolichopodidae). — ratedfromthelatterbythehindtarsomere1with Zootaxa1003:1–32. 13 black ventral bristles at base and curved Yang,D.2001:Diptera:Dolichopodidae.—In:Wu,H.& surstylus. In yunlonganus, the hind tarsomere 1 Pan,C.(eds.),InsectsoftheTianmushanMountain: just has 8 sparse black ventral bristles, and the 428–441.SciencePress,Beijing.764pp.[InChinese.] Yang,D.&Saigusa,T.2001a:Newandlittleknownspe- surstylusisnotcurved(Yang&Saigusa2001b). ciesofDolichopodidiae(Diptera)fromChina(VIII). Distribution.KnownonlyfromTibet. —Bull.Inst.RoyalSci.Nat.Belg.Ent.71:155–164. Yang,D.&Saigusa,T.2001b:Newandlittleknownspe- Acknowledgements. The authors wish to thank Mr. ciesofDolichopodidiae(Diptera)fromChina(IX).— FashengLi(Beijing)forcollectingspecimensandhelpin Bull.Inst.RoyalSci.Nat.Belg.Ent.71:165–188. manyways.ThisstudywassupportedbytheNationalNat- Yang,D.&Saigusa,T.2005:Diptera:Dolichopodidae.— uralScienceFoundationofChina(No.30225009). In: Yang, X. (ed.), Insect Fauna of Middle-West QinlingRangeandSouthMountainsofGansuProv- ince:740–765.SciencePress,Beijing.1055pp.[In References Chinese.] Zhang,L.,Yang,D.&Grootaert,P.2003:Newspeciesof Becker,T.1922:DipterologischeStudien.Dolichopodidae ChrysotimusandHercostomusfromBeijing(Diptera: derIndo-AustralischeRegion.—CapitaZoologica1 Dolichopodidae).—Bull.Inst.RoyalSci.Nat.Belg. (4):1–247. Ent.73:189–194.

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