Descriptions and records of the genus Diplurodes Warren (Geometridae, Ennominae) from the Philippines

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TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan
uetma 7)'ansiopiaSloa Japan se(4)2:67-280,September 1999

Descriptions and records of the genus Diplurodes Warren (Geometridae,
Ennominae) firomthePhilippines'

RikioSATo

2-27-29Shindori-nishNii,igata.950-2036Japan

Abstract Twelve species of the genus Diplarodas are recorded from the Philippines , Nine of
them are new spocies and the others are newly recorded,
Key words Diplarodes, Boarmiini, Geometridae, new species, new records, Philippines.
Dipfurocie sis a monobasic genus describe dby Warren (1896 )for the single species n vastha
from the Khasia Hills in India. Later some new species were added to Diplarodti sfrom the
Indo-Malayan subregion, and some species from various genera were transferred to Dipinrodes
on the basis ofcharacteristics in the abdominal segments and the genitali ain the male. But
there had been some confusion in the systematic treatment of Diplarode sand it salli egdenera
until Holloway (199 3ga)ve a strict definiti oonf the genus ,Dipfurodes and the other three
known genera ,Myrioblophara Warren, Allaayop aWalker and Ectmpidia Warren, In addi-
tion he established three new genera ,thi:pychAzph aSrkeat,obkephar Mag,ribkphara, to receive
from Dipinrocles.
species excluded
Aocording to Holloway (1993 t)h,e genus Dipinrode sis define dby the fbllowin gcharacteris-
tic osf the male: "strong pair spf coremata between segments 4 and 5, and 7 and 8; a setose,
tongue-like proces sarising as a sort ofpseudouncus from the tegumen; gnathus with a broad,
tongucrlike dista lponion densel ycoyered with pale weak spines; a weak costa to the valve,
the lamina of which is often erinkled; a sing!e basal saccular process ,digitat eb,earing a
robust spine at it sapex; a narrew juxt tahat fuses with the vinculum and extends beyond it
as a slender rod". In panicular the gnathos and coremata feature sare characteristic of
Diphtrodes and effectively used to distingui siht from the other related genera . The genus
Dipfurodes contains extremely variable and similar species and it is not easy to identify each
species only by size, colour and maculation. The most usefu1 and reliable diagnostic
characteristios are found in the male abdomen and the genitali aof both sexes,
From the Philippine st,wo species were describe das Diplaroci ebsy me (Sat o1,990) :one is D.
kai:shot frriom Palawan and the other D haonensi sfror nLuzon. Both species are closely
related to parvularia (Leec hf)rom Japan, and have already been transferred to S2xtobkphara
along with the type species nparvulan'a by Holloway (199 3i)n his original des¢ription.
Therefbre ,we have no rocords of Dipinrode sfrom the Philippine sto date.
In thi spaper twelve species of Dipinlodbs will be treated . Nine ofthem are new species and
the others are newly recorded from the Philippine s.Some species remain undescribed,
because of lack of male spccimens. A taxonemic study on the related three genera,
Myriobfiephat uE,ctngpicf ianad AiigriZ)Zbph afrrao,m the Philippine sis under preparatio nby
me. The materials in ZMC were supplied by the Danish Noona Dan Expeditio nin 1961
i This study is supported in part by the Grants-i nAid for Oversea sScientif Riecsearch Nos 60041078
and 61043074 from the Ministry of Education, Scienc eand Cultur eJ,apan.
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Figs 1-24. Diplitrocla sspp, 1-2. D. inundeta Prout. Negros. 3-6. D, schintbneisten" sp, nov,
(3-4 ,Helotype, dn, Mindanao,ZFMK. 5-6. Paratype ,\, Mindanao,NIAES). 7-8.
n vennoleni sp, nov, Holotype, cl, Mindanao, NIAES. 9-12, D. spinivalva sp, nov.
(9-1 0H.olotype, cl, Mindanae, NIAES, ll-12 ,Paratype ,\, Mindanao, NIAES).
13-14, D. negrosana sp, nev. Holotype, tii ,Negros, NIAES. 15-16. a vetvnoleni sp.
nov. Paratype ,4, Mindanao, NIAES, 17-18 .D. lativaf usap. noy. Paratype, cin,
Mindanao, ZFMK. 19-22, D. tanakai sp, nov, (19-20 ,Paratype ,din ,Mindanao,
NIAES. 21-22 .Paratype , \, Mindanao, ZFMK), 23-24 ,D, semichzrularis Hol-
loway. dii ,Palawan,
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The Genus Drpinrodes fremthe Philipptnes 269

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Figs25-44 Drplarodes spp 25-28 D sinecoremata Holloway (25-2 6csa, Mmdanao 27-
28 \, Mindanao) 29-32 D mmdana sp nov (29-30 Holotype, dl, Mmdanao,
NIAES 31-32 Paratype, \, Mindanao, NIAES) 33-36 D subdlecursana sp nov
(33-34Holotype, Luzon,NIAES 35-36 Paratype,9,Luzon,NIAES) 37-44
[iT,
D tttzoni casp nov (37-3 8Holotype, cln, Luzon, ZFMK 39-44 Paratypes, 4,
Luzon, NSMT)
and 1962, whose detail wsere grven by Petersen (1966 )w,tth special reference to entomology,
and those m ZFMK were taken by Dr Schmtlmeiste rand/or his colleagues during the
collectrng expeditions to the Phihppmes m 1988 and 1993 ,of which accounts were given by
Schintlmeiste(r1993)

All the specimens mcluding the type material recorded m this paper will be deposite din the
Nanonal Ifisutu toef Agro-Environmental Science s(NIAES) T,sukuba Japan, except where
indicated to the contrary
The fbllowin gabbreviations are used to indicate the locatio nofspocimens BMH Bishop
Museum,Honolulu NSMT NationalScieneeMuseum,Tokyo NIAES NationalInsti-

tute of Agro-Environmental Sciences ,Tsukuba, Japan ZFMK Zoologrsches Forschungs-
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institut Museum AiexanderKoenig,Bonn. ZMC: ZoologicalMuseum, Copenhagen,
und
Denmark.
DipturodesWarren

DiplurodesinundataProut(Fig1s-2)

Diplarodes inundotaProut, 1929:74.

This species was deseribe dfrom Peninsula rMalaysia by Prout (1929 a)nd, was re-described
inciudin gthe gharacteri sotfi cthse male genital ibaased on Bornean material (Holloway,
1993) . The female genitali ahave not hithert obeen iilustrat edI.t is newly recorded from
the Philippines.

Male genital i(aFig.4 5C)h,aracterize dby two long straight, slender saccular processes.
Also shown by Holloway (199 f3i,g ,555).
Female genitali a(Fig 6,3) . Ostium bursae widely and unifbrmly sclerotized, longer than
wide; bursa copulatrix sclerotized in U-shape posteriorl wyit,h a 1arg ebowl-lik esignum near
posterio srclerotization.
Materia lexamined, Luzon. 3 cl, Ifugao ,Jacmal Bunhian 800-l,OO Om, 24 km E Mayoyao,
11-13, 1967(H.M. TorrevillasB)M,H; 1cl, Ifugao,Banawe 3,550feetCa, 1. 1957,
v. v.
ZMC: 1\,Ifugao,Banaue 1,2oo 20km N Lagawe,22.ix-16. 1988(K.Cerny A.
m, x, and
SchintlmeisterZ)F,MK, 1 2 \, dtrto,1988-i,1989(T. F,Vermolen);14,Nueva
ciZ x. and
Vizcaya, Dalton Pal}, Santa Fe 8oo m,17. x. 1988 (K. Cerny and A. Schintlmeister) ,ZFMK.
Negros.12(1,Mt Canlaon, 1995,6c72\, ditto,viii.1995. Mindanao. 2di1?,
x.
Bukidnon,40km NW Maramag, Dalongdong,Talakag800 1-3. 1988(K.Cerny A.
m, x, and
Schintlmeister1)\,, ditto3,1.xii.1991-2.i,1992(K.Cerny),ZFMK; 4 Mt Busa,10-17.
cl'i,
1997(nativq 3 S,Cotabato,Mt Matutum, 1996(native 1?,
vi, collector); c3Z, v. collector);
Bukidnen,Mt Kitanglad,iv.1997(native 1 MisamisOr,,Hindangon 20km S
collector); cl ,
of Gingoog, 6oo-700m, 20-24, iv .1960 (H.MTorrevill aBsM)H,. Palawan. 1?,
Mantalingajan,Tagembung 1,150 18.ix.1961,Noona Dan Exp.61-62,ZMC.
m,
Geographica lrange, Luzon, Negros, Mindanao, Patawan; Peninsula rMalaysia ,Borneo,
Sumatra,Java.

Dipturodes (Fig3s-6)
schintimeisteri sp. noy.
Length offbrewing 13-14 mm. Simila rto inundata . Both wings with les scontrast between
dark and pale areas, while in inundota "the basal half of the hindwing and the medial zone
ef the fbrewing predominantly white" (Hollowa y19,93) ;dorsa ldark areas of both wings
brownish grey ,not irrorate wdith white; postmedia liine more strongly angled beyond cell on
fbrewing and les swavy on hindwing than in inundata; longitudin albar centrally placed
between the postmedial and submarginal line salways lacking ,while in inundota varying
from well developed to vanished; underside ofwings without any reliable differenc ebsetween
the two species,
Male genital i(aFi g4.7) , Simila rto those of inunclat a.Tongue-like proces sfrom tegumen
in inundutadeveloped;
absent, while costa more strongly sclerotized; sacculus wider and
longer with a single sinuous process a,pically more darkly selerotized (i ninuntlat asaoculus
with two long straight s'pine-like processes).
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Female (Fig.64)O.stium bursae than length bursa
genitali a wider with paralle lsides;
copulatrix more swollen than in inunduld ,posterior quarter sclerotized and ribbed ,with a
1argebowl-like
signurn at middle.
Holotype. di,"Philippinen Mindanao Bukidnon, Mt,Kitanglad S-SeiteP,rimtirwald1,700

m VIII .1993 ,8e07'N Breit e124055' ELa n'ge, leg .V, Sinjae vex Coll .Schintlm. ",ZFMK.
Paratypes .Luzon. 1 IP, Ifugao ,Banaway 1,2o om, x. 1988-i ,1989 (T .and F. Vermolen); 3
9, duto, 11-12. v. 1986 (nati cvoellector); Mountain Prov. ,Mt Puguis 1,90 0m, 18. vii. 1985
(M.Owada) N,SMT; 1 4, duto, 9-13, ii ,1988 (K.Cern yand A.Schintlmeiste rZ)F,MK.
Negros.1 Mt Canlaon,850 29,iv.1997(B,Tanaka);4 39, duto, 1995(native
cl, m, cl x.
collector). Mindanao. 1 dT, Davao, Upper Baracatan ,Apo Range, Mt Talomo 1,1o om, 3-
6, viii. 1985 (M. Owada), NSMT; 1 4, Bukidnon, Dalongdong, Talakag 8(X )m, 1-3. x. 1988,
3 cl 4 \, ditt o3,1. xi. 1991-2 .i .1992 (K, Cerny and A. Schintlmeister) ,ZFMK; 1 cl 1 9,
Bukidnon,Mt Kalatungan,Mt Bagonsilang1,450 30. 1991,2 ditto1,250 29.
m, xii. (in, m, xii.
I991 (K. Cerny) ,ZFMK; 9 6Z 3 4, Bukidnon, Mt Kitang!ad 1,7o om, viii 1.993 (V ,Sirlj'aev),
ZFMK; I dn 1 \, Bukidnon, Mt Kitanglad ,data unknown (nati covlleector); t c7i 1 \, S.
Cotabato,Mt Matutum, 1995(native 4 9,duto, 1996(native
x. collector); v. collector).
Geographical range. Luzon, Negros, Mindanao.
Diplurodes venvoleni sp. noy. (Fig 7s-8, 15-16)
Length offbrewing 12-14mm. Sjmila rto the previou stwo species. Variablei ncolour and
maculation. In most specimens, medial area of forewing and basal area of hindwing paler
than the rest, as in inundota ;pestmedia lline weakly angled as in inunciat aon forewing,
almost even as in schintimeisten' on hindwing; medial line developed in medial pale area on
forewing ;discocellul saprot of hindwing not roundish but short streak,
Male genitali (aFi g4.6). Very similar to those of schintimeisten', but distinguish efdrom it
by weak tongue-like process from tegumen, shorter sacculus with more sinuous process.
Also similar to those of vestita, which have two saccular prooesses ,one a sinuous process and
the other a short triangula rone from .basal part of the fbrmer.
Female genital ia(Fig.6 5)S.imila rto those of schintbneisteri. Ostium bursae tapered
heavily laterally,
anteriorly, sclerotized with a pair of roundish sclerotizations posteriorly;
bursa copulatrix swollen as in schintbneisten', posterio rsclerotized area not ribbed and
bearing
smaller, a much smaller signum posteriorly,
Holotype, 8i. Mindanao, Mt Busa, x. 1996 (nati vcoellector). Paratypes. Luzon. 2\,
Ifugao ,20 km N. Lagawe, Banaway 1,2oo m, x. 1988-i .1989 (T .and F, Vermolen). Min-
danao, 1 cSi, Bukidnon, 45 km NE Maramag, Mt Binansilan g1,2o om, 2. x, 1988 (K. Cerny
and A, Schintlmeiste rZF)M,K; 1 (fn, Bukidnon, 15 kmNW Maramag, Mt Kalatungan 1,250
m, 29. xii. 1991 (K ,Cerny) ,ZFMK; 1 \, Mt Busa x. 1996 (nati cvolelector).
Geographical Luzon,Mindanao.
range.
Dipinrodes spinivatv asp. noy. (Fi g9s-l2)
Length offorewing 15-17mm. The 1arges stpecies ofDiplarodes fromthePhilippi neTsh.e
contrast between dark and pale areas developed ;dark area unifom)ly grey with whitish
submarginal line ,pale area with medial line and grey irroratio np;ostmedia lline on forewing
beyond between CuAi CuA2.
angled cell and veins and
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as:ta'
v
as
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Figs45-52 . Male genital iaof Dipinrodlas spp, 45, D, inunclata Prout . Mindanao. RS-
5441. 46. D. vetmoleni sp. nov. Mindanao, RS-4237, 47. D. schintbneisteri sp.
nov, Negros. RS-S267. 48. D. spinivalva sp. noy. Leyte. RS-5269. 49, D.
lativat vasp, nov. Mindanao, RS-5271, 50. D. negrasana sp. noy. Negros. RS-
5266. 51. D, tanakai sp. nov. Negros, RS-5323. 52. D. semicircularis Holloway.
Palawan, RS-5309.

Male genitali a(Fi g4.8) . Simila rto those of the previous two new species, but can be easily
distinguish ebdy some strong spines from dista lmargin of valva. In additio nva,lva rhom-
boid in shape rather than lenticul aars in the previou stwo species; saccular proces slonger
and slender, not sinuous,
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Female genitali a(Fi g6s7-69) . Ostium bursa elonge rthan wide, both sides with heavily
sclerotized latera mlargins. Bursa copulatrix more slender than in the previou stwo new
specios, varying geographically in the whole shape, especially in the development ofposterior
sclerotized and ribbed portion ;a bowl-lik esignum a litt lsemaller than in schintbneisteri,
located
anteriorly.
Holotype, Mindanao, Mt Busa, 1996(native Paratypes,Luzon, 1
c3". x. collector). cln,
Quezo nForest Nat. Park 250 m, 8-10. x, 1988 (K .Cemy and A. Schintlmeiste rZ)F,MK; 3
9, Dalton-PaB Santa Fe 8oo m, 21. ix-17 ,x, 1988 (K. Cerny and A. Schintlmeiste rZ)F,MK;
1 IfUgao,Banaue 1,2oo 8-l2.ii.1988(K.Cerny A.Schint!meisterZ)F,MK; 1\,
cl, m, and
Banaway, 11-12, v. 1986 (nati cvoellector); 1 9, Carnarine sSur, Mt Isarog 1,60 0m, 21-22,
v, 1963 (H, M. Torrevilla sB)M,H Leyte .1 (1 I \, 25-29. iii .1995 (nati vcoellector), 1 \,
30. xi. 1984 (nati vcoellector). Negros, 2\, Mt Canlaon, x, 1995 (nati vcoellector),
Mindanao. 1 cii, Bukidnon, Mt Kitanglad 1,20 0m, viii. 1993 (V .Siniaev), ZFMK; 12 cin y3
?, Mt Busa, x. 1996 (nati cvoellector); 2 cl 2 9, duto ,10-I7, vi. 1997 (nati covlleector); 3
\, S. Cotabato, Mt Matutum, data unknown (nati cvoellecter).
Geographica lrange. Luzon, Leyte ,Negros, Mindanao.
Diplurode slativatv asp. noy. (Fi g1s7-18)
Length of fbrewing 14-17 mm. Variable in intensi tQyf markings. Generall yboth wings
dark grey without distinc tcontrast between dark and pale areas, in sorne specimens showing
strong contrast between them or blackish bands fbllowin gpostmedia llines ;postmedia llines
on both wings as in schintbnetsteri; underside marked as in inundota and schintbneisteri, but
much paler ,especially dista ldark areas.
Male genitalia (Fi g4,9) . Simila rto those ofthe previou sthroe new spocies. Valva broader;
tonguelike proces sof tegumen longer ;medial spinous part of gnathos longer ;sacculus
shorter; saccular proces smore robust, apically more strongly sclerotized.
Female genitali (aFi g7.0) , Similar to those of spinivats a,Ostium bursae broader ,lateral
sclerotized margins wider; bursa copulatrix more swollen with a bowl-lik esignum at middle.
Holotype. 6n. Mindanao, Bukidnon, Mt Kitanglad, 4. i .1995 (nati vcoellector), Para-
types . 3 cl 1 \, Bukidnon, Mt Kitanglad 1,7oo m, viii. 1993, 2 (Sn 1 \, duto 2,2oo m, viii,
l993(V.Siajaev)Z,FMK; 2di1\,Bukidnon,15km NW Maramag, Mt Kalatungan,Mt

Bagonsilang 1,450 m, 30. xiL 1991 (K.Cerny )ZF,MK; 2(1 1\, Mt Kitangla d4,. i ,1995
(nati cvoellector), 5 cl, ditt od,ata unknown (nati cvoellector).
Geographic aralnge. Mindanao,
Dipinrode snegizzsana sp. noy. (Fig 1s3-l4)
Length of forewing 14-15mm, Simila rto lativa iinna markings. Both wings unifbrmly
pale brown not tinged with grey and black ;underside with narrower dista ldark areas,
Femate unknown.
Mqle genitali (aFi g5.0) ..Very similar to those of lativatv ab,ut valva much broader ,not
tapered apicad; medial part of gnathos shorter; sacculus shorter with a single more robust
process.
Holotype. di. Negros, Mt, Canlaon, data unknown (nati cvoelloctor). Paratypes N.egros.
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Figs S3-62 . Male genital ioaf Diplvroc l asspp, 53. D. sineceremata Holloway. Mindanao.
RS-5253, 54. D. mindana sp. nov, Mindanao. RS- 5 0 1 7. 55. D. decursaria
(Walke r)B.orneo. RS-5304. 5 6. Dsubclecursari a sp, no v. Negros, RS-5248,
57. D. haonie asp. nov. Luzon . RS-5434. ZFMK. 58 - 6 2 .Apical portio n of
aedeagus greatl ymagnified, 58.D. clecut:saria (Walker ) B. or n eo , RS-5304. 59. D.
subclecursaria sp, nov. Luzon. RS-5299. BMH, 60.nsinecoremata Holloway.
Mindanae. RS-5253. 61. D.min d u na sp, nov. Mind a na o . RS-5013. 62, D.
haonicasp,no v,Luzon, ZFMK. RS-S434.
4 cl, Mt Canlaon,data unknown collector).
Geographica lrango. Negros,(native
blplurode stanakai sp. nov. (Fig 1s9-22)
Length of forewing 11-13 mm, Somewhat similar to veuaoleni, but a litt lsmealler in size,
and paler in colour. The contrast between dark and pale areas ofboth wings stronger; paler
area wider, unifbrmly white, without blackish irroration.
Male genital i(aFig 5,1) . CharacteTize dby elongated saDculus with broad base . Saccular
proces sstrongly angled near base,
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Female genitali (aFig 7.1). Ostium bursae funnel-shaped with latera lsc!erotized margins,
Bursa copulatrix bulbous ,posterio rfburth sclerotized and weakly ribbed, with no signum.
Holotype. dn. Negros, Mt Canlaon 850m, 29.iv.1997 (B.Tanaka)P,aratypes.Luzon.

Ifugao ,Banaue 1,2o om, 8-12. ii ,1988 (K. Cerny and A. Schintlmeiste ZrF)M,K. Mindanao.
2 6Z, Bukidnon, 40 km NW Maramag, Dalongdong, Talakag 8oo m, 31, xii ,1991-2. i. I992
(K.Cerny),ZFMK; 1\, ditto1,-3, 1988(K.Cerny A.SchintlmeisterZ)F,MK; l\,
x. and
Bukidnon, Mt Kitanglad l,7oom ,viii. 1993 (V .Siajaev ),ZFMK; 1 cl, Mt. Busa x. 1996
(native
collector).
Geographical Luzon,Negros,Mindanao,
range.
Dipinrode ssemicircularis Holloway (Fig 2s3-24)
Dipiurocles semicireularis Holloway, 1993 :264,
This species was describe dfrom Borneo by Holloway (199 3)I,t is peculia rin having an
additional pair of coremata betwee nsegments 3 and 4 in the male, and a semicircular plate
on the valva in the male genitali a.It is new to the Philippine sa,nd seems to be cenfined
to Palawan and it snearby islands.
Male genitali a(Fi g5.2) . Holloway's descnipti on(199 3a)s fo11ows ."The genital iaare
distinguis hbeyd a broadl ysemicircular plate extending from the ventral edge of the valve
costa, The saccular sclerotisation is distal rlouynded, unspined. The tongue-like process ori
the tegumen is weak",
Female genitali (aFi g6.6) . Ostium bursae a litt lleonger than wide, with sclerotized lateral
margins. Bursa copulatrix swollen as in schintimetsteri and it sallie sb,ut smaller as a whole,
posterior fift hsclerotized and ribbed, a small signum at righ tside posteriorly.
Material examined, Palawan, 2 ci7, ,S. Vicente 400 m, 20 km NEE Roxas, 12. i-17 .i. 1988
(K. Cerny and A. Schintlmeister) ,ZFMK; 1 \, Mantalingajan, Pinigisa n600 m, 6. ix. I961,
Noona Dan Exp. 61-62; 1 dii M,antalingajan, Tagembung 1,15 0m, 17, ix ,1961 ,Noona Dan
Exp.61-62,ZMC; 19,Mt Salaket9io,oft,19,iii,1996(R,Q,Rodrigues),Tawi Tawi.1

\, Tarawakan, north ofBatu Batu, 15 .xi. 1961 ,Noona Dan Exp, 61-62, ZMC.
Geographica lrange, Palawan, Tawi Tawi; Borneo, Peninsula rMalaysia ,Sumatra.
Diplurode ssinecoremata Holloway (Fig 2s5-28)
Dipinrocl assineconemata Holloway, 1993: 265.
This species was describe dfrom Borneo as a member of Diplarode s,though it has no
coremata in the male abdomen, The other characteristics in the male genitalia almost agree
with those of n decursau a(Walker).
Male genitali (aFig 5s3, 60). Holloway's descripti o(n1993 )'`T:he gnathus is typical of the
genus . The uncus tapers gentl yrather than ending in a `bird's head' as in decumari a.The
vinculum is distal lbyilobed as in ciectii:sana, The valve is regular in shape, only slightly
sclerotised, and without distincti vorenamentation. The aedeagus apex is unspined but the
reflexed vesica bears two directe dspines near it sbase".
Female genital i(aFi g7.2) , Ostium bursae glass-shape dsc,lerotized posterio qruater fbrming
colliculum. Bursa copulatrix not so swollen, posterio rhalf lightl ysclerotized and ribbed,
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TThhee LLeepipdiopdteorpoltoegircaollSoocgieitycal Society ooff JJaapapnan
276 Rikio SATo
without signum.
Materia lexamined. Luzon,2 6i, Banaway, 10. iv .1986; 1 c{n 1 4, dtrto 2,8, i .1986 ;8 ciT l4
4, duto ,l1-12. v, 1986; 2 \, Ifugao ,Banaue 1,20 0m, vic .20 km N,Lagawe, x. 1988-i .1989
(T .and F.Vermolen); 1 (Jn 1 \, cfitto ,9-12. ii .1988, (K.Cern yand A.Schintlmeister),
ZFMK; 1 cJi, Ifugao ,Mayoyao, 1,2oo-1,30 m0, 12. vii, l966 (H .M. Torrevilla s2) ;g, ditto,
5-9. l966(H.M.Torrevillas)2;9, duto,10.viii.1966(H.M,Torrevi11as),1 4, duto,
viii,
13-15 .viii. 1966 (H .M. Torrevillas) ;2 cl, Ifugao ,Liwo, 8 km E Mayoyao, 1,OOO-1,3o mo,
29-30 .y. 1967 (H. M. Torrevi11a s)1 ;di, Camarines Sur, Mt Isarog ,Pil i8oo m, 29. iv .1965
(H.M. Torrevi11asB)M,H. Negros.2 2 \, Mt Canlaon. Mindanao, 1 S.Cotabato,
cf c7,
Mt Matutum; 24, Bukidnon, Mt Kitanglad ,iv ,1997 ;7di, Mt Busa ,x. 1996 ;1di 19,
Bukidnon, 40 km NW Meramag, Dalongdong, Talakag 8oo m, 31. xii .1991-2 ,i .1992 (K.
Cerny),ZFMK; 18, Bukidnon, 15km NW Maramag, Mt KalatunganlMt Bagonsilang

1,250 (K.Cemy), ZFMK; l Davao,Upper Baracatan,Apo Range,Mt Talomo 1,1oo
m c{i,
m, 3-6. viii. 1985 (M.Owada )1c;7, duto ,17-19 .vii i1,985 (M Owada), NSMT. 1\,
Misamis Or. ,Mt Kibungol, 20 km SE of Gingoog ,7oo-8oo m, 9-18, iv .1960 (H. M. Torrevil-
las) ,BMH. Palawan. 1 cl, Mantalingajan ,Pinigisan 600 m, 6. ix .1961 ,Noona Dan Exp,
61-62; 1 6i 1 \, Mantalingajan ,Tagembung 1,150 m, 18, ix .I961 ,Noona Dan Exp, 61-62,
ZMC; 1cji ,10 ,v, 1986; 1\, 30, i .1986.
Geographical range. Luzon, Negros, Mindanao, Palawan; Borneo, Sumatra,
Diplarodes mindana sp. nov. (Fi g2s9-32)
Length of forewin g13-14mm. Simila rto sinecoftemata, b'u tcan be easily distinguished
from it in the fbllowing .Both wings much paler ,almost unifbrmly white, while in sineco-
dorsal dark in female;fbrewing
emata area generally grey, strongly marked, especially
narrower with postmedia lline more dentate ;hjndwjng with discocellul saprot les sdeveloped;
underside much paler ,marked as in sinecoremata on forewing ,maculation lacking or very
hindwing. In lacking in
reduoed on male abdomen coremata as sinecoremata.
Male genitalia (Fig 5s4, 61) . Very similar to those of sinecoremata. Medial part ofgnathos
more slender; valva narrower, dorsa lmargin smooth, while in sinecoremata gentl yexcurved;
basal narrowly sclerotized band from costa to sacculus almest straight, while in sineconemata
gently curved near saeculus; aedeagus les sslender.
Female genitalia (Fi g7.3) , Simila rto those of siheeatemata in the shape of ostium bursae.
Ostium bursae bursa
shorter; copulatrix more slender without sclerotization nor ribbing;
lacking.
signum
Holotype, cin. Mindanao, Bukidnon, Mt Kitanglad, 4. i ,1995 (nati vceollector). Para-
types . Mindanao. 3 [1, Bukidnon, Mt Kitanglad ,4. i ,1995 (nati cvoleloctor); 1 cjZ, Bukid-
non, 45 km NW Maramag, Mt Binansilan g1,200 m, 2, x. 1988 ,1 \, ditt o1,8 .x. 1988 (K.
Cerny and A, Schjntlmeiste rZ)F,MK; 10 cl, Mt Kitanglad 1,7oom, viii. 1993 (V .Siajaev),
ZFMK; 2 \,Bukidnon, 15km NW Maramag, Mt Kalatungan,Mt Bagonsilang, 30.xii.

1991(K.Cerny),ZFMK; Bukidnon,40km NW Maramag, Dalongdong,Talakag800 31.
m,
xii,1991-2.i.1992(K.Cerny),ZFMK.

Geographical range. Mindanao,
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