Descriptions and distributional records of American Mecoptera. 4

29 Pages·1996·13.2 MB·English

MCZ
LIBRARY
:•^^^x•^^^^x•^x•^x<•xx•^x•^x•:x•:•^^:•^^^^^:•^:•^^x•^x•^^:•^x•^x•^:•x•^:•:•^xc«•^x•^x•^x•^^
THE UNIVERSITY OF KANSA&rvard
^^
SCIENCE BULLETM^^^
Vol. 55, No. 14, pp. 519-547 March 4, 1996
Descriptions and Distributional Records of American Mecoptera. IV. '
George W. Byers
TABLE OFCONTENTS
Abstract 519
Introduction 520
Acknowledgments 520
520
Panorpa capillala
The Panorpa involulaspeciesgroup 522
(Panorpa involuta, 522; contoiia, 523; ramosa, 526; attnuiata. 528;
redusa, 530; mucronala, 530; key tospecies, 532)
Panorpa himacula 532
Panorpaserta 533
Panorpa terminata 535
Bittacusdistemum 535
Bittacussylvaticus 537
Bittacusspatulatus 538
Bittacuspeninsularis 540
Bittacuspignalellii 541
Bittacus maculosus 542
Kalohitlacus maniculatus 542
Kalobillarus mornatus 543
Kalohitlacus demissus 545
Geographic distribution 545
Literature Cited 547
ABSTRACT
Sixteen new species are described and illustrated. These are Panorpn rapilhila (southeastern LT.S.A.); P. involuta
(Wracriiz), F. ccmtorta (San Luis Potosi and Hidalgo), P. ramosa, P. mucronalaand P. nrlusa (Hidalgo), P. attenuata (San
Luis Potosi), P. bimacula (Oaxaca), P. serta (Michoacan), Bittacus sylvaticus (Veracruz) and B. peninsularis (Baja Califor-
aninadSBuirt)tafcruosmspMaetxuliactou;s K(aCloosbtiattRaicucsamaanndicNuilcaatrusag(uGau)a.teAmalleac)to,tAy',pedninsidsseussi,gnK.atiendorfnoartuPsaanonrdpaBitttearcmuisnadtiasKtleumgu,ma(nCdostthaeRimcaal)e,
is ilhtstrated. First known males ofBittacuspignalelliiNaviis, from Panama, and Bittacus maculosusB\ers, from Trinidad,
are illustrated and briefly described. Some new distributional data forvarious species are provided.
' C'ONTRIBL'TION NLMB1:R.'5176FROM THE SnOVVENTOMOMK.KAr MCSKLM AM) DEPARTMENT OF EnTIl\l(U(K.\, L'MXERSrTVOF RWSAS, LAWRENCE, KANSAS
60045-2119.
520 The UNRTRsm'of K.\nsas Science Bulletin
INTRODUCTION
patienceinenteringthepaperintocomputerfiommy typedand
re\ised copy.
Students and associated with the Snow Entomo-
facult)-
logical Museum at the University' ofKiuisas have in recent new
yearsdoneextensivecollectingofinsectsinMexicoandCen- Pauoi-pa capillata species
tral America. Some results ofthis collecting include new Descriptionbasedon37males, 13femalespinned,and5inales,
AsmpdaedtciiidoeonsnoaafbloMnuetecwotphsetpeegcreiaoegsdraeasnpcdhridicibseotdcrcihuebrureur,oennaacslewdeoalftlaoatashreenrebawspseiecndifeoosrn.- o5mcoeflHrelemtia:aedlml:eosscDtoiorlniromarsiluccymohehaluonllngo.eewv)ie,snhltebyrrymoeilwnlnaolwtiossehagmbmbreeonrwtnboerfxomcwaenxpit(lppliiaconebnapsballayprsopioaspmtid--
collections received for identification from other sources. cally to wholly brown. .Aiuennal scape yellowish brown, pedicel
CoDmesstcorcipkt-iNoenesdohfamwisnvgstveem.natPairotnicfuollarllovwwtihteh rweifdeerleyncuesetdo d8abrlTkaheocrrk,asxfe:ltaaPgerelaollnuoomntgduaamrntkdeulbrliroo\rwelnml,aoxrwvgiiitshhng3aia3n\d't,o2t3rna7enasf\rlearpgsoeeslltl\eormrieuorgreossm.ea.rg\viint.h
theBittacidae, in the taxonomyofwhichwingvenation has Mesonotum\ellowishgra\ withdiffusegra\-brownspotatbaseof
longbeen utilized,someadditionalabbreviadonshavebeen eachforewingandatanterioredgeoneachside,spotsconnected
usedbyEsben-Petersen,othersbvme. Theseare: FM-first in some individuals. Metanotum sordid yellowish brown to yel-
sfeocrtkoro;rObrMan-cohriingginofofthtehememdeidai;aFfRrso-mfiCrUst]fo(rakppolfietsheherradeiatlo bflaorcwoeiwssnhofwgciroat\xh.aseh.oPrlFtee,umrsaoplraars,suertfipabacileaese,haacniordsx,taaemrsoiaspntadleevmiyederelanltouwoinnsehvaebnnrtloeyrwinao:rmcbsouenrr--
Bittacidaeonly);ORs-originoftheradialsectorfromR (or spicuousblack setae on tibiae and tarsi.
Rj)N;oSckvey-ssuabrceoisntacllucdreosdsf-voerimn,osftroomftShcetsoevReir.al Mexican or graWyiisnhgsbrtoiwnng.edNwoitchompaplleetyeelblaonwdiss:hmbolsotwnc,rosspsa-ivseeilnyssinpodtitsteadlwtiwtoh-
CreenmtarianlAtomebreicdaenscsrpiebceidestdheastcarikbeeyd hceorueldbebceaumsieslseoamdianngv; wsthpiionrtgdslmaaorrfggewirinntgihnadnsarokomhteh'eimrnsad.rigviiFdniureasdlt:s,aoneusdtpeesrceiracalodlniyadflebcmaeasllaleslsd.sapProtktesernopesrdteisngeemnaatrl;
however, a tentative key to the Panorpa involuta species humeral spotsabsent.
group in Mexico is included. Abdomenofmale: Unevenlysordidyellowishbrownthroughout,
with short, pale pubescence. No posterior horn on sixth seg-
ment. Notal organ a slightlyextended, broadlv rounded caudal
ACKNOWLEDGMENTS marginontergum3,withlong,down-cuivedsetae,andamedian,
setoseprocessonanteiiortergiun4. Hvpovalvesofsternum9 (Fig.
molFoogristthsewlhooasnespcoelcliemcetniso,nsIaaremiinnddiecbatteeddhteorteliineaffotlelro\b\vintgheenitnoi-- pt1ah)liwecikydeeenlnleeodwdiasbnhedysoedtnaadrekmaeildno-enldgenmtgeotshba,rltomhwaenrngininanrssroeomxwceeedpmtatlotewhsao.rsdeTanpeeearxrg,uawmpietxh9
tialsin parentheses: bilobed,with subrectangularspace between lobes. Inner, apical
Mr. Wes Bicha - private collection in Oliver .Spiings. Tennessee (postero-venual) marginofeachbasistvlebearinggroupof(>9black
(WB). setaeofvaiyingthickness,theiracuminatetipsextendingintobasal
Dr. Da\idK.Faulkner-NaturalHistoiy Mu.seum,BalboaPark,San cup ofrespective dististvle. Outer margins ol dististvfes (ventral
Diego, California (SDM). aspect) nearlystraight,exceptapical portionstronglycuivedand
Dr. Eric M. Fisher-California Department of.\griculture, Sacra- darklysclerotized;basalcupshallowlyconcave,ventromesalin po-
mento, C:alifornia (CDA). sition,vWth stionglvsclerotized. amber-colored bordercomplete
DDrr..ofODlaNivavtieudrrSGa..lFHFliiunsrttt.ohJrry.-,-MWSuamssihetihunsmgotnooifna,CnohDmi.psCta.irtai(itUliiSovNne.MNZ)ao.toiloongayl,MHuaisveaurmd gceoxrvcoeeorpvitendbgasnvaeelnaltyrr.amlVieda-nnltedrnagmltoipsiat,roawfimtelhartedestaar(lkFsigua.rmf3ba)ecterwoi"fsbtaaeprdibcasa"nld(hstatrloafunotsfvseeeratscaeehl),y
University, Cambridge, Massachusetts (now at the USNM) exceptforglabrous,acuteapex:marginalbarbsofbasalhalfpale,
(MCZ). short. Dorsalparameres (Fig.2)yellowish,dorso-ventialh llattened,
Dr. WilfordJ. Hanson - Department ofBiologv. Utah State Uni- blade-like, short, usualh concealed byventral parameres in ven-
versity, Logan, Utah (USU). tralaspect. Ventralanddorsalvalvesofaedeagusshort,concealed
Dr.Jean-Michel.Maes-MuseoEntomologico,SIL\..\.P.527,Leon, in ventral aspect. Base of aedeagus withom distinct hamulus
Nicaragua (MELN). (Byers, 1993:59),lateralwallsofprojection formingcup-likestruc-
Dr.MarkF.O'Brien-MuseumofZoology,UniversityofMichigan, ture concave caudallv (Fig. 5).
.Ajin.Aibor, Michigan (UMMZ). Abdomenoffemale: Terga2-6darkyellowishbrcjwiitobrown,more
Di.EdwardG. Riley-DepartmentofEntomology,TexasAand M tenninalseginenrsunevenlyyellowishbrown:correspondingsterna
University, College Station.Texas (TAM). paler, yellowish brown. .Subgenital plate of stennim 8 (Fig.8)
Dr. Roy R. Snelling - Natural Histon- Museum of Los .\ngeles mosth flattened\entrall\,lateralmarginscuivedupward,apexnar-
County, Los.Angeles, California (UACM). rowly rounded,pale (sligluh projectinginafemalefromRiinsville,
TheSnowEntomological Museum isabbre\iated SEM. Alabama, Fig. 8), major marginal setae sparse along sides, more
numeroirs (severalsubmarginal) nearapex. Genital plates (Figs.
I pardcularly thank my colleague Wes Bicha for making so 6,7) withprolongationsofapicalplateslender,acute,bowedout-
manyspecimensandcollection records available for my use and ward: basal plate loughlv U-shaped: axial portion compressed
forhis helpful commentson this paper. (deeperthanwide),extendingbeyondpostei(_)-medianmatginof
TwospeciesofB///«a«describedhere {B. liistmiiimandB. spal- apicalplate:anteiiorapodemesslenderinventralaspect,strongly
ulatus) wereincluded, in 1985, in an impublishedmaster'sthesis divergent.
byMr.RicardoRoggero.thenastudentattheUniversiu-ofKansas. Measurements: Bod\ length (basedon|)iunedspecimens),male,
I had illustiated and tentatively named them, and he described 9.8-11.3 mm. (holotvpe 10.4 mm.): female, 10.0-11.2 mm. (allo-
them, which I acknowledgewith thanks. The descriptions here, tvpe 10.0mm.). Length offrontwing, male,9.2-12.0 mm. (holo-
however,arenothisandarebasedon man\ additionalspecimens npe 11.0mm.):female,9.9-13.1 mm. (allot^pe 12.1 mm.). .Anteimal
collectedin more recentyears. length, male holotype, about 7.3 mm.; female allotype about 7.4
Iwish tothankSharonHopkinsandJudyWigleswoith l()rtheir mm.
American Mecopter.\ 521
mm
ol. _jt
0.5mm
b L. I
5 6 7 8
Figs. 1-3, 5-8. Fnnorpa capiUala n. sp. 1. genital bulb, male holot\pe, ventral aspect (most hairs omitted). 2. right dorsal paia-
mere, male paratvpe,ventral aspect. 3, leftventral paramere ofmale,ventralaspect. Fig.4, leftventral paramere, toponpic male of
Panorpasigmoi/iesCarpenter,ventralaspect. 5, lowerportion ofaedeagusofmale,ventral aspect. 6,genital platesotfemale parat)pe,
ventral aspect. 7,same, right lateral aspect. 8, subgenital pl.ite offemale. Scale a. figs. 1, li-8; scale b, figs. 2-5.
o.fMaFTyaypyiee9st7:t8eH,,oGlF.oaytWev.tpteBe,yeCmioa-.lsea,(nGadWnTCdi.1Wfmi.eallYdeocupanatgraa.lt.o^Agpluel,eot1ny0op.mei,1.)f.(emAKiallakebm,am)(ianm.oarl2teh4 f1(en9mo8a.2l,e1W)s,;ePsSuEBsMik)cuh;saTL(ra4ikgemg,alC1eo2s..;,8W8kBmM,aSNvEEM1)9o.f82OMx(If2SoSmrIadSl,SeIs2;P3PWIMB:a)yLaa1f9na7dv8e,3t1Gt.eMCaWo.y.
ancl 3 female pararvpes, Tombigbee State Park southeast of Tu- BversandC.W.Young (4males,3females;SEM);LeeCo.,Tombig-
pelo, l.ee Co. (no. 2), Mississippi, 24 May 1978, G. W. Byers and bee State Park, 18 Mav 1993, G. W. Bxeis (4 males; SEM); 5 May
C.W.Young. Additional paratvpes: ALABAMA: BibbCo.,6Mav 1982 (1 male) and 30 Ma\ 1982. Wes Bicha (5 males; WB, SEM)
I11c99i98,022,,3WWAeeprssilBBii1cc9h9h4aa,((W.2temmsaallBeeissc;,haW1'(Bf2)e;mmaaCllheeso;;cSWtEaBMw));C;oC.Dh,eoKOci.-ti^SalbwmiCC.oo..W,,.1Do1efASpBorutitol- WT1BEf)Ne.mNaElHSeoSlpEoaEur;apte^,pMeacslMliointnytnpheeCaUon.Sd,NmM2o0;stM9apmvaarla1et9s8\(pi5en.scWioenllstehcBetiiScoEhnaMo;f(1W1Bmm.aallee,;
Slate Park, 20 Mav 1993, G. W^ Bvers (3 males; SEM): same but Thetypelocality isaForestryReseaixhStation ofAuburn
RCR1.ao9i.-nB.2sr3vMoioMlwnlanetv,e&13S9JDa9i.0nm,oPeoTSl.t1la9Sot7c,ce"hkiPi,ae(Grf1.ke.mr\a\1(l\21e:BMmvaaSelvEresM:1)9(S;84E8DM.fe)eDKm;.iailWsle.asbm;BCeroSz.bEo,uMst)3k.;a128(M-k22am4mdaiMNlsaeo\son,,f vlUaenircviteeordussfitrcyoo,mmpohonosniUet.yesSs.uacHnkildgehpw(oLaioysnoi4cn3e.ria\)yP,aiwnniottrhhpeabrscaaampmibellleahtsaabi(wtRaaustb,ucsoa)ll,-l
2 females; SEM); Maiion Co., 30 Mav 1982, Wes Bicha (1 male; shaded by an almost pure stand of tulip trees (or yellow
WB); Winston Co. (no. 2), 8.9km NEofDouble Springs, 19 Mav poplar, Liriodendron lulipifem). planted in 1953. The iin-
C1(.9no9W.3,.1)GY,.o1Wu4.n.4gBkve(m2rsNfe(1moafmlaPelesen;;ciSSlEEMBMl))u;f.f,M.o2\1nRKtMAgNaoSym.Ae1.i9S7y:8,CMoGo..nWt(.gnoo.Bmve2e)ir,sTGaCanodp. edecrosttoonreysisbteytpwiecaelnhawboiotdaltafnodrsthaisndspeocpieesn, fuiseuladls.ly Aftouontdheirn
Creek,2.1 kinSEofJoplin,OuachitaNationalForest,22Mav1978, placeswhere P. rapillalawascollected, the habitatwashon-
G.W.BversandC.W.Yoimg (1 male;SEM);PolkCo.,RichMoiui- eysuckle, brambles, poison iv\', in some places greenbriar
tKaEinN.TpiUtfCaKllYt:rapL,i(vii)nughs-tolnONCoov.,.31199M5,ayH.1W9.82R,oWbeissoBni(c2hama(l2esm;aSleEsM,).2 andalwaysvariousbroad-leavedherbaceousplants, but the
522 The Universit*'ofKansas Science Bulletin
treesprovidingshadewereofmanyspecies. WesBichahas havingmuch thesamemoiphologyofthemale'sgenitalbulb
found capillata (and its more northern counterpart, P. sig- asin capillataand sigmoides. Particularlywith regard to the
moides) also in deep shade alongsmall streams. P. capillata ventral parameres and the apical setae on the basist\'les,
isoften foundtogetherwith thesimilarandprobablyclosely Panorpakirisimaensislssikiisveiylike capillata, and P. kamiko-
related P. nchulosaWeshvood. liensis Issiki is veiy like sigmoides (Issiki, 1929: 184-187).
Forsome time, I had tentativelyidentifiedthisspeciesas Thesejapane.sespeciesoccin*on the islandsofKyushuand
an aberrantform ofPanorpasignioidesCarpenter; butitbe- Honshu, respectively. Unlike capillata or sigmoides, they
cameapparentthatsigmoides\\-Aiamorenorthern range (In- have a horn-like projection from the sixth abdominal seg-
diana, Illinois, Ohio, Wisconsin, Minnesota), while all ment in males, complete pterostigmal band on the wings,
specimensofthe rap/llnUifomiarefrom thesouthernstates. and other differences.
It will be interesting to see what future collecting in Ken-
tucky andTennesseewill reveal about the relationships of The Panorpa involutaspecies group
t(hCeaPsraepneotnrvtvpoearf,rofrl1/m«9s73/.1rt:/«b2e2l1)onagnsdtointhmeanneybudleotsaailgsroofupbootfhspmeaclieess P.Trhameosfao,llP.owmiuncgrosinxatsap,ecPi.esre-clPuasanmo'ripdaP.inait'otleuntua.atPa.-choent\ooilnag,
batcraanaprldielplfbaaeitrmauaammhlaeae\ryseebsrseeh(soeFeraitmsgisbbl.lyaer1dsb,i3sfs)fai.eglrnoeiTnonhigtedietsashteeeadanirndfenresonrlembeeunbdldogoeetsr,ah.aibnnyMdntaehllbeauerlsvgoesenoal.-yf waticoaogra,rdofgtrritooopmutptahhpaeeppyravoarirtxceiifnmifratootymuenolofydthtOiehrneritzlMhaaeetbxiaeti.uacdsaetInetorfsnspCeeimecuoimdiesasmdipoafrMioPanbansantobeorlfpesaoM.utethxAahs-t-
Ttttrhhhaeelenbpaanapraribcrasarmlaoerowreneeeld-o(ttFnhoiggiea.rrnd.3)aoIcfsnuehtcaoeacwpthsiilpvlt,eahitnenatratsapihligencpivaoaelinrdpteaorsmraaeltnriadeosnpcisaeopcefitxltlpohaafteana,pd\aaeerndnad,-- aht5hd0oad0lti-otkPtiiayolnpnooeamrlepftsarepormemecbxiaieOncsradiinznoaaftbBhmaaio,snukgbnsretol(au1oip9nn1gsw3si)il,lntdkobinectoahdtiwiesnsdc.ioonnvIvlenoyrlfuefatdciato,wgmiittrihiotssiulnmpiaktelholeefy
mere covered by dense, slender, recinnbent barbs, except species. Allsixhave,in themale,averycomplicatedaedea-
oatlrapnatplart=phoaexhraieiamixmaytet)rre.eleOmysenotnhttieahpt-.etghdiIiotvrreisdssalttthhhseeeurwhsfiapaiedcrcteyi,heatsophfieptrsteehaneriasapmanaecgrleaa(moLbeafrrtoteiuh.nsezcvTaoephninlee-- AgnttinaicsdmeaiebamlllgleiihkvieanevnttehhtaewothptsehleeneemnoaanlilinevroeietnaghciedloreuncMdioeedfdxeitdschlpeayecnagirbePseadenoanomnrdcepoanltsoo,art,nhhdmeeontgschroteorunatpxho..e-
sthaemveensttrraulctsuurrefaicne,siwgintiohidbeosr(dFeirgi.n4g)biasrbbsr.oaIdnlycagplialblartoaumsoosnt WpiToine-tfhlieerismtohkrinshopawisng.smoemnte iostdheerrivseodiufcreom(e.tgh.e, dmieettaboofltihce)sicsorat-
setae along the mesal margins of the hypovalves of the preAsent
ninth sternum are slender and yellowish, biU in signioides keyto thespeciesofthe involutagroup (and including
they are dark brown to black. And in capillata the apical, P. mexicana) follows the species descriptions.
ventro-mesal setae of the basistyles are thick and black;
these are rarely thick or dark in sigmoides. Panorpa involutanewspecies
ltahtaIttnertch(oeCrmaepraaprreeintnsegorm,cea1pi9elr3lr1ao:tras2a5inn0d-t2hs5ei1g,onirofiiiggdi.ens3,a7l)it.desshTcohrueilpdht\ibpoenovnoafoltvtehedes 17DHfeeesamcdar:liepsDtioionrnsabluacmsoehuodnloe.xneC5no2hl'omrallsiegbsha,ts4be0drfooewnmnalpteiosn,hnrpeoidxnvnisiep;de,cbilamancedkns3a.rmaoluensd,
are not particularly long and do not reach the ends ofthe ocelli. Rostrum and frons below ocelli yellowish brown to pale
Aboafscitoshtmeyplgoeessn;iitnteealidtbrhuaelrwbiinscgtohuoelfdg\eepnnoitstrsaaillblbayunlrdbesaeunllttoenirgnoa-ttvheeen.atsnerdaflasulalestnpydeeicrmt.s- dbbtaarriroonkswwnnggrreteexoeecnnbeilpspathcikgrg,morsematnayrxttiuolimnlaadmliamavroknspevtarlibpbnlsrdaiobcvwakisndoaualnathlsts)bei:drremomsiwonn(sa,atlpgmesroxeaugodtmfihenlpngoatsr.ttthrsrSnocdmuaargprehek-
pressions. andpedicelyellowishbrown,flagellumdarkbrownwith35-37fla-
Differentiatingfemalesofcapillatafrom thoseofsigmoides gellomeres, basal one longer than second and third together.
aat(h1nn)eddpan3o)ersbteuielnmrontsiaahoenristmpdoaiosfrffigittciiuhnoletn.ospfrIotonfhxgteiehmnfeeaorblrareleaw,dinntgcgheh,eerosiefnoaafcrnaetpihfalepoliuapcrttaasel;prtobohtasesntsaidegl,moaarn(leg2 vauraenelrtclTesyulrewimdrofabaorrelxkmnd:eastrnPsgibrlilnionaggn.chaokttlMtsyseneitdsmdaaeoesrbnkrl(oeIoojrtinw:tugtnemw.s5wi-tiit7mthaehttohxuesictilcndkeph,e\a'satbtbtalerenaodr\dcn\pk)iooi,isn,weliptsiatacolhennetsnsaetutolfmmleeeuladamrcit:ohaerul1ahss-l'3isdtgemerrraaealoncldsn,t--
band, and (4) the second basal spot (actuallynearbut not black setae above each wing base. Metanotum brown without
doloensna.stehTlehyempaigreggnmiietnna)lt.epdlaTtihenesssoiefgmcmoaaipdrielklsiaatnangasdreacarmpeoilrlleaatrasgleterhnadanenrdininmnvoeerbnue-- cctmoeaoxrrxaekaedie.npagasnFl,deewmmhioaetrihrraasn,xudelmalorelnkrogwoyeieussltshlaosbwhnriodosrwhtmnbbo,urstwtoiwntdnohentlgosorenargyeoidssndehtiaastenhi.tnebgPirelioeoiwurnnra,ssluowrsmifutearhcfseaspcsceeascot,-f-
tralaspectthanthoseofnebulosaandlessslenderthan those imens:tibiaeandbasitarsidarker,gradingintodarkreddishbrown
of.ugmoides (compare myfigure 6with figures 65and 66 in on distal tarsomeres.
CarApnenitnetre'resst1i9n3g1criervciusmisotn)a.nce,possiblyofbiogeographical bmeafrWokirinenggessn(dbFirogof.wR9.n9)att)oompsolaidlgeehitg-lrayatvbeielshyhiibniriddoeewsnncd.enotAf,pMil.ic.gahlbtlubytatpniidnggfmerdeonwmtitsdhliifggfrhuatsyle,y
significance, is that there exists inJapan a pair ofspecies and clearareas included; in males, apical band often reduced to
American Mecoptera 523
clouding over outer radial and medial cross-veins and narrow AttlietimeIfir.stcollectedthisspecies,thehabitatwascloud
darkc'iiiui;alongapicalwingmargin. Pterostigmalbandcovering forest of numerous kinds of tropical broad-leaved trees,
mbaosralehsallefnOdferstciognmtai,neuxatteinodnionfgbtaonMd|atchreosnsnfaursrtowcellylcMoinnteocetneddotof sometree-ferns,shrubs,manyvines,ferns,mosses,abundant
Mi; in some males band interrupted from Mi to M>j. Narrow bromeliadsandotherepiphyteson treebranches,andape-
trans\'ersespotfromM3toendofCui,anotherfrom Ri acrossfirst culiargroundcoverofSelaginellaabout 10-15cm. deep. Re-
forkofRstoM] (mavbeweakbetween R2+:',andRi+r,);widerspot turningto thesitethreeyearslater, I found thatwhilesome
fsfiprrosottmfa(otlriokirigotifonMC.ouf2Rsa,bofurtommiSdcwaacyrobsesnRvieetonRcsu.biWthalitcirsohss-l\he\irnisd:iismmialalt otfhethleowheirghterseetstarenedsnhaatdivbeesehnrulbefsthtaociprboeveindecusthaadned,bmuorsntedo,f
Abdomen ofmale: Terga 2-5from evenh- mediiuii brown tosor- andcoffeeshrubsplantedintheii place. Thereweieextensive
did darker brown, corresponding sterna paler brown; segments areas of bare or nearly bare soil with only patches of
s(Dte>iot9oarnbesf;lurosmowmmanol.pflo,ssNetsocegtlrmaeileronorottrmizga6e,ardngi,niwfnleoaaontkfvehrata'reldrdlageysuvpmeperlc3oto,jpweeicctdoth:nisfnnpegaiwprcdoruoioonwwut,nslcosyunhnocrteotendrc,pgarbvuoljemaecci4kn-. pSeelcWailgaiilnleeynlliaanllisevftetr,.ePaNamenvsoeirrdfteuhileeilaiefmsyos,l,uhlPea.rdbiinasvcpoelloauyutsasawvaebsgreistgtiahlttlipgorrnee.seetn!t(,leesss-
]30steriorhalf Hypovalves (Fig.9) shortbutexceedingventralcon- oftenyellowishgreen) color,particularlynoticeableon the
nection between basistyles, borne on gradually narrowing pro- thoracic andabdominal pleura, rostrum and legsbut even
longation ofninth sternum nearly twice length of hypovalves, in in some well sclerotized structures, for example, the geni-
voefncterracli;aasppeecxt.shTaellrogwulyme9ma(rFgigi.na1t0e).nBaarsrisotwveldcsgsreapdauraaltlevdplaes.ststlehvaenl tal bulb ofmales. Eventually this color largely disappears
halftheirlength,inventralaspect. Inner,postero-ventralmargin fromspecimenspinnedanddried,butitleachesoutrapidly
of each basistvle bearing about 12 mesally directed setae (Fig. from those preseiTed in alcohol.
11); other setae on mesal surface ofbasistvle. Outer margins of Panorpairwolulasomewhat resemblesP. mexicaiiaBanks,
dttiiessrtti^iolsret)velsleusornfsgalaictgeh;etdlryvoewcnotonrfcoal-vomneegsbahelafhior,rsseheaxmlitldeo-nwldleivnngcgtohnf;craob\maesbaoalnsaclluopcwueoprfp(daaisnst-- anesaprecOireiszsatbilal,kVneorawcnruozn,lyonflryomabiotsutma3l5ekhmolfortoypmet,hceoltlyepcetleod-
mid-length of dististyle; inner margin of dististyle with sharp, calit)'ofP. itwolutn. hi the male ofinvohtla. the hypovalves
stronglysclerotized,downwardlycnnedpointconcealedbybasal extend slightly past the edge oftheventral connection be-
csp(aFruliopgjsbei.rcna2tv2nie,cnngh2te3rc)saol,nevsalepsnoiptncergucaatoltueb(s,Frliaygen.afccr1hh2oe)ms.beegVelaenorninittgnraaglatlebp,upelwanibrdtauihlmnoaelpuraisetce,sersatmlwcearomso-psbbesrrciatan;ngncdoahounersdd- sstehwtoeareetnootfnhethtbihaseseipsdtogyselt.eesr,Towhvheeirnlteeraainrlemsneuxurifmcaeacrneaooutfhsetdyhaecrlkbe,aasrcilsoytnytsleepr,imciiunnoauitnse-
inner appendage. Dorsal paramereselongate, not darklysclero- voluta, butin mexiranathereareonlymoreslender, shorter
li/ed. with two acute, triangular "teeth"onventro-lateral surface and paler setae. The ninth tergum in involuta is shallowly
\odbalfalerevkeabscebhtsuhwtaoernnetdon,tnbtdvlieiposmnttso,iefnldyyla"orstwkceeleraetrnhbo"dtriandzneeecandhr,seeeerslvybeoansfsclevly.eenrcotVutrenianzcltedrdp,.aalrtaivpmaselvrueessusa;sllldeyondrvesisrai,l- eprrmooauxrnigdmieandlat(tesoeaettnheaonppletyxei,forwsohtmiilgtemhaetlhsaibtdae)no.fdTmihneextirhceeaanfraoeraeopnwplieynagtrswsootsfoptobhtees
Abdomen ofjemale: Terga 2-4 or 2-5 iinevcnh' bi"own, sterna tspe ofmexicanabut four in malesofinvoluta. I did notex-
paler;moreposteriorsegmentsgravishbrown,sometimeswithfaint amine the aedeagusof mexicanain detail, but in\entral as-
ibnedhiicnadtimoind-olfegnrgetehn,.poSsutbegrieonritmaalrgpilantebro(aFicgl.ly1r.3o)usnldieghdthwiwthidaebnoeudt pect (cf Byers, 1962: 303,fig.8) itisevidentlyveiydifferent
20setaelongerthanothersonplate. Genital plates (Fig. 24) nar- from that ofinvoluta.
row, with onh small, nearh transparent lateral extensions repre-
senting apical plate, "basal plate" semimembranous and poorly
defined, attached to adjacent tissues anteriorly; axial portion Panorpa contortanewspecies
es8rmm1.etna0m(lr.ldi)aMo5e-sten;1,imga1fvohs.meaef'u0l)bmryasom.eacppumllamoteeLlerndee8ot.e(n.t1smh71ig:I.ozetnme,lhsB3doml-ot1oabc,d2tyfu.ueypt8frfneoalaemelrenm1edntama0egws.lsrtpi3elieh(niocagmagrtl(,hmb,lbteo)omalnstuay;stdyltervpsedfuee8,ocen.oftm1t51nu2ars0r.lm.alpe2eid7mig,-.nnhn.1net18nanh..er)64ddl.-imy1.\2Assme.tnpr0retga(ceemiihnngmomthneltaano,lstp(a)yaonl,lpdtelemeneoargtm1ltiy1eheop.s,,re4 ofmitcsaerehxmulHDilabmeeillrsa.lesocdalwrR:riinioygnps.hDtttpaloarilSylicorcpoundsashna.puboraelamk.nsAeypdeyreiedlfcallrtoaloolonwsnwiishids3eashb5slhe,fbmlbrwaooroilfwoetwshtwno.ecn,ter,lr1mpla2iedicdnaefmiareocolkmsefatlsbglleerbrygseroym,eoweenwnplnnlint,otnewonoafieflbrsdahlp.agabapec4lrlekpolxmswauanamrlb,neoldsdaura.conrkisdkn--1
km)Typneos:rtHhoelaosttypoef,Cmoaslceo,mcaotlelpeectce,dVienrcalcoruudz,forMeesxt,ic4o.,8 meilleevsat(i7o.n7 btoroswencotnodbalnadcktwhiitrhd t3o3g-e3t5hefrl.agellomeres, firstsubequal in length
about 1420 m, on 8 August 1969, by G. W. Bvers (field cat. Ver- Thorax: Pronotimiyellowish brownindepressedareas,slightly
ac.PVm1•aoeca9\rhrr7luaoau1getlucz.svs,rtnNpuisoe.nz1.sa,9Da.m7.l11Ce7c1,oP)o5d.shGeamoc.ntAalonWal,lmyl.eaa.ossts(tB,5ayehympop1emeel3era,ocldsftea,faeystenmp,makdaeal;m3alRes.efs12,ae,6T9lmhs,mpaloaailor1lmennn6eshnpes.iee.,ld;d\Sla,uF1s.tg:1aa3u8Msmf)bmtmee;uaamtlls1aleeo9la9ssc9ema,..3sleA,,i95tul\pfWgfo'ieu'cemsbenamatlsunailtetle(Bdvesn2i,,so3cb.huS1Jiat2nE1u8Mnfa3)(el;e1.7-- ledomltivaoonebnernrignkegaaeleaeesyrcrytihelbsmalierseglntiohovdadtwweeeinmnbosoolrhonfsyonabtwmnryneedtoe,lseelweornlwnsinsoai,ootwetretiuhodssam.snhulahartoaslbrgrncharittun,noetws.erwi\dlitenalohMrtirsmekwteisiis.usnrtoregihfntPedaagolsoceeetpefsadmsugroimroasrrlfeeeafe,scconunotlrdpxedvfsaadiam;lescbee.i7eba-tslc8Fs,aeekebncitwlmnaooaaoextrfc;raedukaem;mssoeaaenruttnnvaanaddo;e-e
Lmina'ltMehsMe.ZS3EafMne;dmapslaeorsma,etWyTop5te)hse.hrHaovcolelolbteeycepteni,osnasel.nltottoyptehaenUd.SmNoMs.tptahreaMtxCpZe,satrhee sieritWdaieesncbgelsnatcw,ki;twhitaotrhustibalrimoghwtcnobmsrptoliwgenmt.aebaannddsm,artkiinnggeds.widNiayrerlolwowdisahrkbernoiwnng,
theThmiaslespe(lc.iaelsintaiknveosliiittsan=acmoempflreoxm,tihnetrciocamtpel)i.cated aedeagusof artadeiadlgecelolfswfirnogmianboouutteRr:>,rtaodisallighcetlllys;belhriannsd\eRr5soerstpootMia.crPotsesroosuttiegr-
524 The U\r\-ERSiTi'of Kansas Science Bllletin
10
12
14
m m
1
16
Figs.9-13. Panorpii involulaii. sp. 9.genital bulb, male holoU'pe.ventralaspect (mosthairsomitted). 10, ninth abdominal terginn,
male parat\pe, dorsal aspect. 11. right dististvle, male parat\pe, posteroventral aspect. 12, setaeofveiitrome.sal siufaceofbasist\le. 13.
subgenital plate, female paratope. Figs. 14-17. Ptinorpa tontortan. sp. 14, genital bulb, male paratvpe,ventral aspect. 13, ninth abdom-
inal tergum, male parat>pe, dorsal aspect. 16, terminal abdominal segments and cerci, female allot\pe, ventral aspect. 17, subgenital
plate,femaleallotype,ventralaspect. Figs. 18-21. Panorpa ramosan. sp. 18, ninthabdominal tergum, male holotype, dorsalaspect. 19,
genital bulb, male holotvpe, ventral aspect. 20, right dististyle, holotype, posteroventral aspect. 21. subgenital plate, female allot\pe,
ventral aspect. Scale, figs. 9-11. 13-21.
American Mecopter.\ 525
24
Figs. 22-24. Patuirjxi itwoliiUi n. sp. 22, waedeagiis, male paratvpe, left lateral aspect: dp - dorsal paiamere, dv - dorsal valve, vp =
ventral paramere (both branches labelled), - ventral valve. 23. same, ventral aspect. 24, genital plates of feiriale, ventral a.specl.
Figs. 25-28. Paiwijxi (otitoria n. sp. 2.5, apex ofmale aedeagiis, left lateral aspect. 26, left halfofaedeagns, ventral aspect. 27, genital
platesol li-male, ventral aspect. 28, same, right lateral aspect. Scale, all figures.
526 The UNrvERSiTYofKansas Sciencie Bulletin
inal band repit-senlc-d by wedge-shaped spot from basal halfof where a road turns north to .S;ilto de Agua (known locally as El
stiirma to outer nvgma, slightlvbehind Rj (interrupted between Salto). Holotype, allotvpe and 4 male, 4female paratypes in the
R_l and R-,oroffsetnearforkofRj+-, in some individuals). Small SEM; 18 males, 2 females in collection of Wes Bicha; 6 male
spotsin firstcell Ri.at origin ofradialsector,between lAand 2A paratypes in collection ofTAM.
abouthalhvavbetweencross-veins,andatendofCiii. Otherspots Panorpa(ontortnistheonlyMexicaiispeciesknown toocctir
uinnmsaormkeesdpeincismoemnes.maHleisn.dCwrionsgs-svweiintshilnigohutteerrmraardkiianlg-sme,dviiarltuaarlelay significantlybelow5000feet (approximately 1525 meters),
ucisaulahl'livnnfoetmabloers,demroesdtionfwmahliecsh,hoarvemamyorbeewsilingghtshpobtsortdhearnedin,measlpees- wPahniocrhpmaakynorewlnateintoMetxhiecfoa.ct thatit isalsothe northernmost
(e.g.,fromendofCui toMs,Cui toCu-jatle\elofORs,oratends At the tvpe localitv in San Ltiis Potosi, the general habi-
oopafflbAlerbAsodaeaotndamldee;y2ncArooo)]ni.memasdlepe:odn,Tdesirlnigggahts2t-me6erundnaieasvlneingphlrtyohvkpe)alnllgeoarw.tiisNohontbaorlfowoternrgwgaiintihmfos3rhaomrnetdd, trbarotoamwdealssioaaundtsho,waemksotsfosorefessttahneindsfluaiimvimelriyvtvh)oirg,tisiwiomtnohttamhbteualinmandrsagenr(t2t.5re(epieispk.hmyUtnbi-cy
wlosDamuuoaritrlidslfs.eoutprmaHmeevg,opwfwoitiodvptotaahshlctvsecethrsohairmoroormpfuohgtndahiilaponftutctoehi,rnsg^metsenleodiirrgtncehaitelmlopivbfhuad(slliFebivagged.wmrgeho1enen4nt)nta,6snbltredeexoenrptudireegoernrh,sdtstioeefnfdruglg,nliyetwmaoiifrtoolh4ryr.-- wudhaeensrrnbgafarmocoueewnodttd.hi,sTithpnolcegalenduttedhsceeid.r(diwwetiodotloh1ydmagyrnseohhetirnghtihiesbroshsnuPp.pwechctioioencts2hoortPfaaiPhhiaieognrlhojdi,rapiataclsoswanoislonlyrgoetswta
slightly beyond edge of ventral connection between basistyles roof-like above the body, at rest, occasionally raising them
(mav be bent to sides in apical halfdue to diying), separated at in a "fanning" motion.
bmaetdaegssleeaesvl.belveTdowgefierdcstgehrulcomion,fg9saeharharlyaclpnhooedwvdlamyldovcere)mreaosiarngdumiimonnerabetra,eosuawalsinthdthahlbfva,ienllaoltbobhrweouidspsethaltavhlaanoipnaresgrxraool(uwFotienegdgr. =coimBnpytlriiitccsaatgter,eedecnosmtcrpculjcletoxru,rientwoliiffsett,heedit)sm,waPil.negcaomnetadoceruatlagaustshioo(wnL,satacilnnodsceobnystiotmrhite-a
15). Inner,subapical surfaceofeachbasistylebearingnumerous larityto P. involutaofVeracruz. Itdiffersfrom invobdaand
mesallv-directed, thick black setae. Oiuer margins ofdistisrsles other related species in details of the aedeagus and con-
isllnoicngrghetaalssyiicntsognglryceasavctleeesrntoetwaii/rd.temdhi.,dw4Bieatnshagltpahcl,ue,pstcoruforndbgielsdtyisscteiytulaeTeeaadblosoiunbUgapl1i.o5cwaetlrilmeyedasgnead,s bfiolrombaetdiosnubogfentihteahlvppolvataelvaesn,dicnhatrhaectmearlies,tiacngdentihtealbrploaatdelsy,
sparsesetaeonoutercunature. Stronglysclerotizedpointonmesal in the female.
marginofdististvleconcealedbvbasalcupinventralaspect. Both
wbirtahncshmeasllofmveemntbrraalnpoaurasmearpepsenthdiacgk,ewoelnlsscolmeerowthiazetdt(iFviigsst.ed25d,or2s6a)l, Panorpa ramosanewspecies
branch. Dorsal paramereswith single spinousprojection at apex Description based on one male, two females, presened in al-
and heel-like s\velling on dorsal sinface near base. Dorsal valves cohol.
short,rounded,inconspicuous;ventralvaheselongate,notstrongly Head: Dorsimi paleyellowishbrcjwntotanwithappressedblack
sclerotized. hairs; black aroinid ocelli; rostrimi pale vellowish brown, un-
Abdomenoffemale: Terga 2-6 imevenlydarkyellowish brown to marked,with nimierouserect, blackhairs. Moiuhpartsyellowish
brown; corresponding sterna pale vellowish brown; more termi- brown; apical halfofterminal segment ofmaxillaiy palp and all
nal segments sordid brown or grayish brown. Basal segment of ofterminal segment oflabial palp dark brown. Antennal scape
cercusbrown,short(Fig. 16);apicalsegmentblack,morethantwice pale tan,pedicelbrown, flagelkmi blackwith 37 (holotype) or38
lengthofbasal segment;cerealbasesiniusualh deeph separated. flagellomeres.
Subgenital plate (Fig. 17) broad, apicallvbilobed with short me- Thorax: Pronotumlighttanvariegatedwithbrown,almostwhite
dian projection in most specimens. Genital plates (Figs. 27, 28) at lateral edges; six (male) to eight (female) conspicucjus black
withstoutaxialportionandonlvdistalplatewellsclerotized;basal setaeandnumeroussmalleronesateachsidealonganteriormar-
platelargelymembranous, itslimitsindistinct;distal platecurved gin;shortblacksetaeon posteriormargin. Dor.simiofcenixlight
ventradatsides. Anteriorapodemesthick,moderatmehm'divergent. browninfemales,grayin male. Mesonotimiandmetanotum pale
Measuremerils: BocK lengtmh,mmale,about9.3-11.6 (hc)lotype tanwithirregularlightbrown shadingalongeachside; nimierous
10.2mm);female,9.2-m11m.0 (allotype9.6mm). Lengthoffore reciunbenlblacksetaeonscutimiandscutellum,directedcaudad.
wmimng,(amlalloet,yp1e0.131-.163.m0m). A(nhtoelnotnyaplel1e0n.g5thm,mm)a;lef,emaabloeu,t181..(S5-m1m1.,9 Pplaelienhaalirssu,rfbauctesh,aicrosxdaaerkaenrdonmearnaterpailoerstuarnfawcietshosfccatotxearee.d,Fmeomsotrlay
female about9.9 mm. andtibiaepaleyellowishgraywithblacksetae;tarsislighth'darker,
Types: Holotype male, allot\pe female, 4 male and 3 female apicaltarsfjmeredarkbrowntoblack,setaeblack;basitarsusnearly
paratvpescollected 14.9miles(23.8km) bvroadwestofElNaranjo, aslongas tarsomeres 2-5 together.
SanLuisPotosi, Mexico,26Augtist 1972,'elevation 4000ft. (1220 Wings hvaline, veiy lightly tinged with gray-brown, iridescent,
m),bvG.VV.BversandR.Thornhill (GWBfieldcatalogueno. 15). with dark brown markings; apical spot (or band) approximately
Additionalparat\pes: SanLuisPotosi, 16mileswestofElNaranjo, from end ofR'j toendofMi; pterostigmal band roughlvtriangu-
819.9S2e.ptWeemsberan1d992F,reWdesBiBcichhaa((195mamlaelse,s,3f4emfaelmeasl)e;ss);amTeabmuatul9i.pSaepst,. alapr,exitsatsohrorntebaarsMeja;loountgercohstaal,focfovsetriignmgaplirgohxtibmraolwnha.lfofstigma,
RanchcjdelCielo,nearGomezFarias,3July1989,R.Jones (5males, Abdomen ofmale: Terga 2-5 pale yellowish gray medially, irreg-
one withoiu terminal segments); same, about 7 miles (11.2 km) ularlv darker grav at sides: corresponding sterna extremely pale
westofCiomez Farias, 6-7 Jiilv 1986, Jones and Kovarik (1 male); gra\, nearlywhite; pleurawhite (but see note on color in discus-
Hidalgo, 22 miles (35.2 km) bv roaci northeastofJacala, 5300 ft. sion); segments (i-9 vellowish brown. Notal organ weakly devel-
(1615m),27August1972,G.W.ByersandR.Thornhill (1 female; oped, with slight, wide backward extension oftergum 3 bearing
G1V4NkTimnno.or1t6h)e.asTtlioeftGyipuedlaocdaldietlyiMnaiSza.nELluiNsaProatnojsofdisoaepsprnooxtiampapteealry d4.ow(niecnuintealclb,ulbbla(cFikg.set1a8e);eslmoanllg,atmeeadnidannaerlrevoawt.edHvppoionvtalovnestesrhogrutm,
on most mapsofMexico. It ison Highwav80. the roadfromAit- slender,deeph'separated. Tergum9naiTowedbeforele\elofcerci,
tiguoMorelos (onHighway85,theInteramericanHighwav) toGiu- apicallybilobed; lobesbroad, rounded,slightlvdarkenedatapex
daddelMaiz,andisapproximatelv29kmwestof.AntiguoMorelos, (Fig. 19). Basistvlesseparated\'entrallvlessthan halftlieirlength.
American Mecoptera 527
Figs. 29-31. Panorpa ramosan. sp. 29, aedeagus ofmale holotype, left lateral aspect. 30, genital plates, female, ventral aspect.
?i\.genital plates, female,anteriorapodemesomitted, rightlateral aspect. Figs. 32-35. Panorpaatterjuatan. sp. 32, aedeagusofmale
holotype, left lateral aspect. 33, dorsal branch ofleftventral paramere,ventral aspect. 34, genital plates, female,ventral aspect. 35,
same,rightlateralaspect. Fig.36. Panorparechisan. sp.,aedeagusofmaleholotpe,ventralparameres (bothbranches) removedfrom
leftornearside. Scale, all figures.
eachwithgroupofblacksetaeon inner,postero-ventral margin. mere,withblunUvroundedlobeprojectingdorsadandnearlyuans-
Outer margins ofdististvles slightly concave before mid-length, parent, pendulous branch directed basad. Dorsal parameres
stronglvcin"vedindistalone-thirdtodarklysclerotizedtips;basal elongate, paler than ventral parameres, eachwith two subapical
cupofdistisrsleelongatedventro-mesally,shallowlyconcave (Figs. spines, or "teeth", on ventral sinface, basal one larger; nvo tiny
g1i8,n2o0f)d,iwsittishtvfloem-belaornign,gblaaccukteseptoaienotnnocarumdaalllysicnofnaccee;almeedsailnmvaern-- veennetdratloswpairndesacnienaiirinbaatsee.tipVse;ndtorraslalvavlavlevseselsohnograt,ter,osilmednedde,r,dadrarkkl-v
tralaspect. Ventralparameres (Fig. 29) biramous,lowerbranches sclerotized.
cunedmesad,theirtipscrossing;dorsalbranchessinuous,acutely Abdomen offemale: Tergayellowish brown tobrown, sterna 2-7
lipped, each extendingcaudadalongside respectivedorsal para- palegray,nearlvwhite. Irregulardarkspotfromeachsideofter-
528 TuF, UniversityofKansas Science Bulletin
gum 1 downoverplcurontometathoracicmeron. Siibgenitalplate spondingsternapaler;segments6-9yellowishbrown. Notalorgan
g(iFinga.t2i1on)wbiidoeasdtlnyeaUr-smhida-pleeidi.gtlGi,enbiitlaolbepdlaatteaspe(Fxigwsi.th30m,ed31i)animeumsanr-- aapsehoxratnpdronjeucmteironoufsrodmopwonstceuinioerdmaharigris,naonfdtetrrgiuanmg3u,lwairtphergoounndteerd-
allylongforgenus, axial portion projecting bevond apical plate, gum 4 with scleroUzed apex curved cephalad. Posterior halfof
sidesofbasal plate stronglycuned i:i lateral a.spect (Fig. 31); an- dorsumofsixth segmentconcave in lateral aspect,withoutsetae.
teriorapodemesabout ten timesaslong aswidth ofportion pos- Hypovalves (Fig. 37) short, notreaching edgeofventral connec-
terior toseparation, onlv slightlvdivergent. tion benveen basistyies, slender and attenuate toward tips, sepa-
malMeeaaslulroetmyepnets1:3.8Bomdmy,lfenegmtahl,empaalreathyopleot9y.p9em,mabo(ubtiu1p4o.s2simbml,ydfee-- raabtdeodmibnasaallltverbgymmno(rFieg.t3h8a)ntbraismaclatwiedotrhveonfshaalhlvopwolvaylevmea.rgNiinnatthe
formed bydamage when collected); fore wing, male, 12.7 mm. atapex,slighd\'darkenedatcorners,withsparsedarksetaeondor-
female 12.7 (allotype) to 12.8mm. Antennallength, male,aboiu simi,pale,yellowishsetaeonmargins;cercidarkgrayatapex. Ba-
11.3 mm, female, about 11.6 mm. sist)les separated slightly more than halftheir length, in ventral
allTcyoplelse:cHteodlonte\aprehmiaglhe,waayllo1t0\5p,e4f.e4maklmeanonrdtohenaestfeomfalTelapnaclriaittyipoel a1s4petchtic(kF,igb.l3a7c)k;isnenteaerpdoisrteecrtoe-vdenmtersaladmaarngidnboefneeaacthhbetahreisengaabcoount-
(about98kmNNEofPachuca), Hidalgo, Mexico,on 6July 1992, spicuous,roundedprominenceon mesalsurfaceofbasisrvie. Dis-
byJ. S. Ashe (his collection no. 8). Elevation at this site is 1420 tistvieswith6-7longsetaeabovebasalcup (Fig. 37);outermargin
m; the habitat is cloud forest with large tree ferns, dense moses slightlyconca\enearmid-length;shaip,sclerotizedpointonmesal
on trees; the scorpion-flieswere on low foliage. M\ three speci- margin, concealed by basal cup in ventral aspect. Ventral para-
mensare in the SEM. meres (Fig.32) two-branched,ventralbranchesacuteatapex,with
Pannrpa ramosa is one ofseveral species (most of them small subapical tuft of short setae, their tips overlapping, pro-
ipcroevtihoautslwyhnenndeaslciiveibaerde)doisfttihnecthliygghrleaenndissho,fetahsatt-cceoiillorralbMeeixng- jpDeoecrntsdiaunllgopsualsir,gahmtmleeyrmefbsrroeamlnogoneugnasitteaa,lppsbieunlnubdo;augse,eacfohlnadtmotreessnaaeldlbasrunardfnacechxepb(aeFniadgr.eind3g3)i.na
particularly evident in the pleural areas ofthorax and ab- apical one-third, with small, subapical point and larger, more
domen. In callow individuals, even sclerotized areas may proximalpointonventralmargin (Fig.32). \'entralvalvesyellowish
abthenadgtrlteehesensi"soohfr,tiefgnoirntaehlxeachmoelpaoldre.,oftDhbreo.gdAeysnihwteaa'slsbafuisletblrdiokfniomntagellesismc,eotmhgermeleeengnst", dc(aoarrpAkrpb,eedsawopirmtoedhnnadrriokonfeufgrenmsdiatneledevr:enlnaoTtrbpeaaerllgaeaatrsep2ia-ne5ccmthuo)nss,eitlvdoeefnenogihfn'aavlnesednost;rsrdslaieeldngvdymaeeelrlvn;letosds.woir6ss-ha9lbdrvaaorlwkvnee,rs
which "faded in alcohol within a few hours." Puiwrpa mex- yellowish brown throughout. Subgenital plate widest near mid-
icana, described by Banks (1913) as "pale yellowish," was length,unusuallydeepin lateralaspect,with threeposteriormar-
pserrovbeadb;ltyhoenoenloyfkthneoswenthsaptewcaismegrneoefniitsihsparmiaorletofbreoimng"Oprriez-- oagifnndeallsoolnomgbaeetwse,haca.texnidatlaerrpkolerontbeiedo.nmoaGr:eiendibtlraatolearpdalllalytyersroo(iiFmimgddse.edd34t,ahpa3in5c)allactaoemnrapdlrbioasnseeadsl
aba, Mexico." plates (notdistincdyseparated);anteriorapodemesdivergent,but
inteiTeningregion sclerotized,cur\edslightlyventradanteriorly.
broDHweensa;cdr:diapDrtkoiorbnsruobPmawasnsneoadorrrdcopjiinadmoydanetoeltcleemolnwaluiila.seth,aRbsonirsxeotfwwrenumsmap(laehenocslid,oetfpsyriopnnens)edbt.eolmowedocieullmi mll1aee1cnl.MTtge0eyet,padhmes,1smbu0:e.rm;sea1Hi-mlf1odeee0eln,.rotn6Hatsai:lbymegpo,mheBu,wto1(ad1m7ay.ya.l38ll8-loe51emt,2n\m.(gpa6It,elnhlm1,tfo0ete.mmry1maapalieml(neeaeamilh)nilao.docbltaoo\Lfnutpiet\eyHnepig1e1fgt0,1ehh..mw0a3aoabmflymoe)mfumo,t.)pra3e.1r21wa.ikt,6noAmgmpn,etsmmseo;aniclnuofeahle,l---
dark\ellowishbiown (holot\pe) tounevenh'brown,rostrumdark west ofTamazunchale (labels sav 20 mi. west), San Luis Potosi,
amber-brown at sides, palest apicallv; niouthparts light brown to Mexico, 19 September 1974, byW.J. Hanson and G. E. Bohart.
fobbafioanermtlsdwobudTaiweslshmv;nroli,o-rdetsnabunhofxaredrrl:leeodasneriwgPeden(soral,nerryolotleauntafylpmeoolihmetbocnoc^lawulejlaioabumcstalrskhlydaeippcasibgoekrhnrms)kotse.bleew\nertr'nSntoaei,gcwleootalrnfwhaoptiwmmetittaaoesohaxrahboignscllubidehaltrnccapo.skordeiwynawddnMdireip,etckaoahesedlnnlrapo3dsrn7yamk-edontae3ltahtsr8leirtkuorrkfiwmoidlenionagsntarghsoentem;gdrlbdaealrtnrinomohngusmeewivemmtrenrneoa.re;rsnsstt--e,ao AcohatlrtfheaeeLPrtc'idashotfelenliapoloehspoedrueScspgttitaaagehttbedeeaesodtbtcvUtooiienetlnoinhlntvuteheahPcoertetafsainhSDiooisrnEnrgu.Mphp,s,eawWLitra.oaefayJg;nin..acdhdnieF,aBtllihfLatle'ontytasupareonhauend.rs,s,aieIttntfmyhwrsbpeaoeelsmscheoosaahlnlriisodetsot-syigDfgnprireee.otolwhdgHaternanhacdnopoftsloaheolrlsieenl,cscoattwttliIyholopaknneyt>
ousshort,recumbentsetaeonscutum,slightlylongeratsides;few nearest neighbor species, P. contotia (from Hidalgo), and
longersetae on scutellum. Pleural surfaces, coxae and meraun- inwing markingssomewhatreseinblesotherspecies in the
evenh'yellowishbrown^\ithscattered\ellowhairs,lotigestandmost involutaspeciesgroup. Inseveraldetails,however,itdiffers
denseonanteriorsmfacesofcoxae. Femoraandtibiaeyellowish fiom all these. Males ofattenuatacan be recognized read-
bdarrokwenr,tlhaarngetribitaieb;iatlasrestoameerbelsac4k-;5gtraardsionmgerfersom1-g3rayyeltloobwliaschkgnreaayr, ily bv the short, attenuate hypovalves (Fig. .37), for which
claws; trace ofgreenish pigment in tarsi ofsome individuals. the species is named (Latin attenuata= drawn out, thin, ta-
Wingsmoderatelyiridescent, lighth- tingedwith brown, mark- pered). The rounded protuberance on the mesal margin
ings brown. Apical band represented by faint, narrow marking ofeach basist\le and the unusually long setae on the ven-
aPinctreoorusotssetoriugRtm3earclerlbalasd:inamdlocaronemdpnlmeeeardtlieyablcuotcmenplllaserwtrieotwhapalinocdnalgiibrtrauedngidunlaianlr,ocfnorenonfmeecmptarilooexn.- MtrealxiscurafnacPeaonfortphaesd.istDiisstysleecstairoenbroetvheaulsnitqhueeeaqmuaolnhguknnioqwune
imalendofstigmatoendofM4 (interruptedbetween Mi andMa aedeagus (Fig. 32),unlikethatofP. contortabutshowingaffin-
finroomneR]fetmoanlye)g.maTrbaentswveeernseR4s+p5otanfdroMm1+M2:jwtiodetirpsopfotCufir,omanCoutjhetro fiteymwailtehstohfePi.nvaotltuetnauastpaecrieessegmrbolueps.thTahteofsuPb.gceonnittoarltapilnatheaov-f
piousmteartioFrM.margin atend oflA; small spot at FRs. Wiidsh thyrid- ingtheposteiiormarginsomewhattliree-lobed;buttheplate
Abdomen oj male: Terga 2-5 unevenly yellowish brown, corre- iswidestnearitsposteriorendandhasbroadlyroundedlat-