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Dams as new habitat in West African savannah Patrick Claffey Des observations d'oiseaux ont ete faites dans le petit lac barrage a Kakikoka dans la province des terresdusuddeBurgou province, auBenindeseptembre 1997amai 1998. Denombreuxsejoursont etefaitpendantlasaisonseche(Octobre-Avril).Soixantedixespecesonteteenregistreesaproximiter du barrage, incluant six nouvelles especes dans la liste Benin: Tachybaptus mficollis, Gallinula cbloropus, Micropari'a capensis Charadriuspecuarius Charadriusforbesi et Calidrisferruginea. . , W 'aterbirds have always been difficult to find in Geographical location and climate thedrysavannahofWestAfrica,onlyoccurringin Thedamissituated5kmsouthofthevillageofKakikoka, thesmallpatchesofsuitablehabitat,whicharescarceinthe intheGuineasavannahofcentralBenin.Thisvegetation dryseasonandofteninaccessibleduringtherains.How- coversmuchofthecountry from07°-l1°N.Theannual ever,inWestAfrica,asoccurredpreviouslyinsouthern rainfall is cl,200 mmwith,the heaviest rains inJuly- Africa,habitatisincreasingwiththebuildingofnewdams. September.Monthlymeantemperaturesarec26°C,witha Theincreaseinthewaterbirdpopulationcanbeobserved maximumof37°CinFebaiary-March. inaveryshortperiodoftime,andthedamunderstodyhere Theareaimmediatelysurroundingthedamisheavily hasalreadybeenvisitedbyseveralspeciespreviously cultivated.CropsincludeyimiDioscoivaspp. cassssiiManihot , unrecordedinnorthernBenin-4121319orthewholecoun- esculetitadeeOtyzasativa,Pennisetumamericanum,cotton , try*.Thesmall damatKakikoka (09°01'N02°l6'E) in dGousest]optbhuetvnasrpipe.tyaonfdhgarboiutantdsnpurteAsernatc—kiswboyopdolgaenad.H,orwiep—avreira,n BorgouProvinceisoneofseveralprojectsofthePeople’s vegetation,smallareasofdenseforest,andfarmland itis RepublicofChinaintheRepublicofBenin.Constaictedin stillrelativelyrichinbirdspeciesandIhaverecorded300 1995,itisc850minlengthandalittleover200mwide.It specieshereovera10-yearperiod. hasbeendesignedtomeettheneedsofthesemi-nomadic pastoralistFulaniorPeulpeoplewholeadtheirherds,often Species of particular interest on the upto500cattle,andasmallernumberofsheepandgoats, Kakikoka Dam ontranshumancethroughthenorthernsavannahsofsev- Clearlymanyofthespeciesobservedonthedamwere — eralWestAfricancountries. expectedatthistypeofwaterbody herons,bitterns, AremarkinWaterbirds promptedmetoundertakea ducks,wadersandrails.Manyothersonthelistarecom- simplestudyofthissmallwaterbodyfromSeptember monintheareaandarecoincidentaloronlydistantly 1997-May1998.Mostvisitswereduringthedryseason dependentonthewater.Afulllistofthespeciesnoted (October-April)whenthewaterislowestandthearea withintheperimeterofthedamisgivenintheappendix. aroundthedammostaccessible.Ivisitedthesite,often However,thefollowingarecertainlythemostsignificantin severaltimesamonth,tocountsomeofthespeciespresent termsofbeingrarities,newtothearea,significantPalearctic there. orintra-Africanmigrants,ornewBeninrecords.Thefirst Thewesternendofthedam,wherethefeederstream sentenceofeachspeciesaccountdescribesthespecies’ enters,isshallow,probablynomorethan1mdeep,andis overallstatusinBenin. quicklycolonisedbyNymphaeacaeruleal()duringthedry season,becomingalmosttotallycloggedbytheendofApril Little Grebe Tachybaptus ruficollis whenthewaterisatitslowest.Thereisalsoareedbedatthis NewBeninrecord.Singlemaleon12September1997. end.Theshoresofthedamarebarelateriteduringmuch Twopairsinfullbreedingplumageon19-20February ofthedryseason.However,severalspeciesofrushesand 1998.Threebirdsinbreedingplumageon8-15April1998. grassarebeginningtotakeroot,aswellasvariedsavannah Twobirdsstillpresenton15May1998.Nosignofany floraafterthefirstrains.Therearesmallareasofmuddy youngbutbreedingseemsprobablegiventhespecies’ shorewhichattractanumberofwaders,particularlyon behaviourelsewhereinWestAfrica.Thespecieswas springpassage.Thewallandtheeasternrockyshoreare apparentlyrecordedonLacNakoueinsouthBeninin favouriterestingandbathingplacesforseveralhundred March1997.Thisisasignificantexploitationofasmallnew CattleEgretswhichjoinalargeroostofc5,000birdsonthe areaofsuitablehabitat2 . OuemeRivertothesouth-westofthedam.Totheeast, behindthewall,thereisalargeareaofsavannah. DamsasnewhabitatforWestAfricansavannah: Claffey BullABC Vol6No2- 117 Long-tailed Cormorant Phalacrocoraxafricanus beenrecordedto06°38’NinTogo3.Newrecordforthis Commonresidentthroughout.Upto20onthedamatany areaandasignificantexploitationofsuitablehabitat. time. Vulturine Fish Eagle Gypohieraxangolensis iMeBittern^o^rvc^nsm/nito Uncommonthroughout.ApairatthedaminNovember Singleinthereedsatthestreamendofthedamon20 1997. February1998.Fewrecords,althoughitisprobablynot EgyptianVulture Neophronpercnopterus uncommonandhasbeenpreviouslyrecordedbothinthe Rareintra-Africanmigrantorvagrant. Anadulton 20 north,ontheTerouRiver13,andinthesouth,atCotonou December1998.Nopreviouslypublishedrecords9 but , (pers.obs.). thereisarecordfromthenorth-eastpartofPendjariPark Squacco HeronArdeola ralloides inthelatedryseasonof1977,andrecordsfromParkW, CommonresidentandPalearcticmigrantrecordedthrough- neartheChutesdeKoudou,inDecember1977(A.Green out.Evidenceofmigrationwithacountof20+on11April pers.comm.).ItwasalsorecordedbySzaniawskyski18in 1998andinthefewdaysfollowing,upfrom1-6birdson theBeninsectorofParkW.UncommoninNigeria10and earliercounts,anddowntotwobirdson9May1998. consideredavagranttobothGhana13andTogo3 . Cattle EgretBubulcus ibis Marsh Harrier Circusaeruginosus Abundantintra-AfricanmigrantinOctober-June.Large Uncommoninthisarea.Widelyrecordedinthesouth. numbersroostonthewallsandstonyshoresduringthe SinglebirdonthedaminDecember-March. afternoon. Allen’s Gallinule Poiphyrio alleni Green Heron Butoridesstriatus Uncommonintra-Africanmigrant.Somepreviousevi- Commonyear-roundresidentonallsuitablewaters.Prob- denceofbreeding".Immaturebirdon12September1997. ablybreedshere2’11. Maybreedhere. Little EgretEgrettagarzetta Common Moorhen Gallinula chloropus Commonresidentandmaybreedinthearea211 Several Notuncommonresidentthroughout,breedingprobableas . presentatalltimesonthedam. attemptednestinghasbeenobserved.Severalprevious GreatEgret Egretta alba records3butnotlistedbyDowsett&Forbes-Watson9.Six NotuncommonresidentandPalearcticvisitor,especially birdspresentonthedamon23February1998,andasingle in the south. One ortwo occurrences onthe dam in on9May1998.Probablybreedshere. February-March. AfricanJacanaActophilornisafricanus Purple HeronArdeapurpurea Commonbreedingresidentthroughout.Breedingprevi- NotuncommonPalearcticmigrantwithsomepresent ouslyrecordedintheareaandalmostcertainlybreedson the dam where up to 30 including juveniles present year-round.SeveraloccurrencesinOctober-April. GreyHeronArdea cinerea at all times4. CommonPalearcticmigrantwithsomeyear-round.Sev- LesserJacanaMicroparra capensis eralbirdspresentatalltimesonthedaminOctober-June. NewBeninrecord.Certainlythemostsignificantrecord forthedam.ThespeciesisuncommoninBurkinaFaso21 White-facedWhistling-DuckDendrocygnaviduata Niger16andNigeria9 andhasnotbeenrecordedineither, Commonbreedingresident.Thedamoftenhas100+birds Ghana15orTogo3 al,thoughithasbeenrecordedinIvory present.ManyyoungbirdspresentinDecembersuggest- Coast19.Itisrepor,tedlylocalinMaliandChad11.ItWhas, ingprobablebreedinginthearea.LayingdatesinNigeria however,beenrecordedontheMekrouRiverinPark 16 . areJuly-October2’10. At 18.20hron7May19981observedwhatatfirst m African PygmyGoose Nettapusauritus appearedtobeasmallTringawalkingontheliliesc70 Probablebreedingresident,whichisuncommontorare, aheadofme.Iquicklyrealisedthatthistinybirdwasvery throughoutthecountry.Threemalesonthedamon23 probablyMicroparracapensisandIhadexcellentviewsofthe February1998remaineduntilMarch13.Thisisasignificant birdthroughatelescopeoverthenext45minutesand exploitationofasmallareaofsuitablehabitat211 confirmed itas an adult ofthis species. It had a rich . chestnutcrown,awhitesuperciliumandachestnutbrown GarganeyAnasquerquedula eyestripe.Thechestnutonthecrownextendedontothe NotuncommonPalearcticmigranttotheextreWmenorth, napeandwidenedacrosstheneck-sides.Arichgolden- particularlyalongtheNigerRiverandinPark 12,17,20 A . yellowwashwasapparentontheneckandupperbreast. pairatthedamon20Februarytoatleast14March1998 Purewhiteunderparts.Slightlygreenishlegs.Orangeor werefurthersouththanpreviousrecords,althoughithas 118-BullABC Vol6No2 Damsas newhabitatforWestAfricansavannah: Claffey .. brownishbill. Itstinysizemadeconfusionevenwith eral observations at a muddy end ofthe dam. a juvenile Actophilomis africanus unlikely. I ob- Wood Sandpiper Tringaglareola served the bird again on the evening of9 Maywith CommonPalearcticmigrantthroughoutthe country. twocompanionsandonseveraloccasionsthereafter AfewbirdspresentduringOctober-April,with20on until at least 17 May. 15April 1998andsmallernumbersonfollowingdays Greater Painted-Snipe Rostratala benghalensis indicating northward migration at that time. Uncommonresidentbutprobablyoverlookedbecauseof Common SandpiperActitishypoleucos itsdiscreetbehaviourandpreferenceforofteninaccessible VerycommonPalearcticmigrantthroughoutthecountiy habitat.Singlefemalerecordedon17May1998.Several OneortwobirdspresentonthedamduringOctober-May previousrecordsfrommostareasofthecountry. Four-banded Sandgrouse Pterocles quadricinctus Ringed Plover Charadriushiaticula Probablynotuncommoninthedrynorth13andinthisarea CommonPalearcticmigrantinthesouth.Singlebirdon15 duringthedryseason.RecordedinDecemberandApril. April1998isthefirstinlandrecord. Kittlitz’s Plover Charadriuspecuarius Conclusions NewBeninrecord.Rareintra-Africanmigrant.Anadultin Thisbriefstudyisinterestingforanumberofreasons.The breedingplumageatthedamon8April1998,notseenon appearance of Little Grebe, African Pygmy Goose, subsequentdays.Severalbirds,includingimmatures,at GarganeyandmostparticularlyLesserJacanaareallasso- GrandPopoon19-21November1995,inanareaofdrying ciatedwithtirepresenceofNymphaewhichistobeexpected swampinfrontoftheAubergeatGrandPopo5 andat onthistypeofstill-water,andallarenewtothisarea. , nearbyAnechoinTogoon14January1995.Thespecies GreaterPaintedSnipe,Kittlitz’sPlover,Forbes’Plover doesnotappearonDowsett&Forbes-Watson’slist9and andSpur-wingedLapwingVanellusspinosusareallresident israreinTogo3.ReportedtooccuronlyincoastalBenin11, waders.Kittlitz’sandForbes’Ploversarerelativelyrarein althoughthereareseveral recordsfarinland inWest thisareaandthusofinteresthere.Severalmorespecieswill Africa16 inevitablyoccurandthesitewillrepaymoreconstant . Forbes’ Plover Charadriusforhesi observation.AfricanCrakeCrexegregiahas,forexample, NewBeninrecord.Uncommontorare.Twobirdsob- alreadybeenrecordedinthearea3andalmostcertainly servedonthedamon4April1998.Twobirdsrecordedon occursinthevicinityofthedam. ThedamsupportsanumberofPalearcticmigrants, 17January1992nearapoolontheroadfromOuidahtothe Togoborder.ThespecieswasnotmentionedbyDowsett particularlyduringspringmigrationinMarch-April,as &Forbes-Watson9 indicatedbytheoccurrenceofSquaccoHeroninsome . numbers,PurpleHeron,RingedPlover,CommonSnipe, Curlew Sandpiper Calidrisferruginea Greenshank,andGreen,Wood,CommonandCurlew NewBeninrecord.UncommonPalearcticwintervisitoror Sandpipers.LittleStintCalidrisminutushasalreadybeen passagemigrantprincipallyinthesouthofthecountry5. recordedintheareaandappearslikelytoberecordedatthe Singlerecordedonthedamon4April1998.Previously daminthefuture. recordedon21January1994and21November1995atthe Severalspeciesprobablybreedatthedamornearby, southendofLacAheme.Twenty-threebirdsrecorded includingLittleGrebe,Long-tailedCormorant,Darter, duringthe1996WaterfowlCensus8.NotlistedbyDowsett White-facedWhistling-Duck,BlackCrakeAmaurornis &Forbes-Watson9. flavirostris,Allen’sGallinule,CommonMoorhenandAfri- Common Snipe Gallinagogallinago canJacana. UncommonPalearcticmigrantbutperhapsoverlooked.A Finally,GarganeyandRingedPloverarenewtosouth- recordon23February1998wasonlythesecondforthe ernBorgou,andLittleGrebe,CommonMoorhen,Lesser area. Jacana,Kittlitz’sPlover,Forbes’PloverandCurlewSand- Greenshank Tringa nehularia piperarenewtotheBeninlist8, (g? CommonPalearcticmigrant,widelyrecordedbutonlyone References previousrecordforthisarea.Singleon15April1998when 1. Blundell, M. 1992. Wild Flowers ofEast Africa. therewerealsoseveralothermigratingwaderspresent London, UK: HarperCollins. suggestingthatthedammaybeanimportantsiteforspring 2. Brown,L.H.,Urban,E.K.andNewman,K. 1982. The migrants. BirdsofAfrica.Vol 1.London,UK:AcademicPress. Green Sandpiper Tringa ochropus 3. Cheke,R.A. andWalsh,J.F. 1996. TheBirdsofTogo: NotuncommonPalearcticmigrantinOctober-April.Sev- an annotated check-list. B.O.U Check-list No 14. Tring: British Ornithologists’ Union. DamsasnewhabitatforWestAfricansavannah: Claffey BullABC Vol6No2-119 i 4. Claffey,P.M. 1995.BirdsoftheBeterouarea,Borgou LittleEgretEgrettagarzetta Province, Republic ofBenin. Malimbus 17: 63-84. GreatEgretEgrettaalba 5. Claffey, P.M. in prep. Review of the sources, new PurpleHeronArdeapurpurea GreyHeronArdeacinerea r(eDcoowrsdesttan&dDroewvsiestiotn-LoefmatihreeR1e9p9u3b)l.ic of Benin list WHhaimteer-kfaocpedScWohpisutsliunmgb-rDeutctkaDendrocygna viduata 6. De Sousa, S. 1988. FloreduBenin. Vol 3- Cotonou: AfricanPygmyGooseNettapusauritus Universite Nationale du Benin. GarganeyAnasquerquedula 7. Dennis,N.andTarboton,W. 1993- Waterbirds.Cape Black-shoulderedKiteElanuscaeruleus Town: Struik. VulturineFishEagleGypohieraxangolensis 8. Dodman T. and Taylor, V. (eds) 1996. African EgyptianVultureNeophronpercnopterus WaterfowlCensus1996. Wageningen: Wetlands In- MarshHarrierCircusaeruginosus AfricanLittleSparrowhawkAccipiterminullus ternational. LizardBuzzard Kaupifalcomonogrammicus 9. Dowsett, R.J. andForbes-Watson,A.D. 1993- Check- Red-neckedBuzzard Buteoauguralis listofBirdsoftheAfrotropicalandMalagasyRegions. BlackCrake Limnocoraxflavivostra Liege: Tauraco Press. Allen’sGallinulePorphyrioalien 10. ElgoodJ.H. 1994.TheBirdsofNigeria:anannotated CommonMoorhen Gallinulachloropus check-list. B.O.U Check-list No 4. Second edition. Denham'sBustard Neotisdenhami Tring: British Ornithologists’ Union. AfricanJacanaActophilomisafricanus 11. Girard, O. 1998.Echassiers, Canards, etLimicolesde LesserJacana Microparracapensis GreaterPainted-Snipe Rostratulabenghalensis I’OuestAfricain. Chateau d’Olonne: Castel Editions. RingedPloverCharadriushiaticula 12. Giraudoux,P., Degauquier, R.,Jones, P.J.,Weigel,J. Kittlitz’sPloverCharadriuspecuarius and Isenmann P. 1988. Avifaune du Niger: etat des Forbes' PloverCharadriusforbesi connaissances en 1986.Malimbus 10: 1-140. Spur-wingedLapwing Vanellusspinosus 13. Green,A.A.andSayerJ.A. 1977.LaConservationdes CurlewSandpiperCalidrisferruginea EcosystemesForestiersdelaRegiondesMontsKouffes. CommonSnipeGallinagogallinago Unpubl. report. Cotonou: FAO/PNUD. Greenshank Tringanebularia 14. Green, A.A. and Sayer, J.A. 1979. The birds of the GWroeoednSSaannddppiippeerrTTrriinnggaagolcarheroolpaus PendjariandArliNationalParks.Malimbus1: 14-19. CommonSandpiperActitishypoleucos 15. Grimes,L.G. 1987. TheBirdsofGhana-anannotated Four-banded Sandgrouse Pteroclesquadricinctus check-list.B.O.UCheck-listNo9. Tring: BritishOrni- AfricanGreenPigeon Treroncalva thologists’ Union. Black-billedWood Dove Turturabysinnicus 16. Hayman, P., Marchant, J. and Prater, T. 1986. VinaceousDove Streptopeliavinacea Shorebirds. London, UK: Helm. VioletTuracoMusophagaviolacea 17. Koster, S.H. andGrettenberger,J.F. 1983- Aprelimi- WesternGreyPlantain-eaterCriniferpiscator narysurveryofbirdsinParkW,Niger.Malimbus19: African PalmSwiftCypsiurusparvus LittleSwiftApusaffinis 62-73. Woodland KingfisherHalyconsenegalensis 18. SzaniawskyWski, A. 1982. Plan Directew:- Parc Na- StripedKingifisherHalyconchelicuti tionaldu duNiger. Rapport Technique 3- Rome: PiedKingfisherCerylerudis FAO. CarmineBee-eaterMeropsnubicus 19- Thiollay, J.M. 1985. The birds ofIvory Coast:status Abyssinian RollerCoraciasabysinnica and distribution.Malimbus7: 1-59- AfricanGreyHornbill Tockusnasutus 20. Thonnerieux, Y. 1985. Notes complementaires sur SingingBushLarkMirafracantillans SunLarkGaleridamodesta l’avifaunedesParcsNationauxdel’Arli(Burkina) et MosqueSwallowHirundosenegala de la Pendjari.Malimbus7: 137-139. HouseMartinDelichonurbica 21. Thonnerieux, Y., Walsh, J.F. and Bortoli, L. 1989. YellowWagtailMotacillaflava L’avifaune de la ville de Ouagadougou et ses envi- CommonBulbul Pynonotusbarbatus rons (Burkina Faso). Malimbus 11: 7-40. Green-backed EremomelaEremomelapusilla BleatingWarblerCamaropterabrachyura B. P. 302, Parakou, Benin. White-shouldered BlackTitParusleucomelas Scarlet-chestedSunbird Nectariniasenegalensis Long-tailedShrike Corvinellacorvina Appendix: List of 72 species observed Fork-tailed DrongoDicrurusadsimilis PurpleGlossyStarlingLamprotornispurpureus within the perimeters of the dam. Northern Red Bishop Euplectesfranciscanus LittleGrebe Tachybaptusruficollis Yellow-mantledWhydah Euplectesmacrourus Long-tailedCormorantPhalacrocoraxafricanus Red-cheeked Cordon-Bleu Uraeginthusbengalus DarterAnhingamelanogaster Orange-cheekedWaxbill Estrildamelpoda LittleBitternIxobrychusminutus Pin-tailedWhydah Viduamacroura SquaccoHeronArdeolaralloides Yellow-frontedCanarySerinusmozambicus CattleEgretBubulcusibis GreenHeronButoridesstriatus 120-BullABC Vol6No2 Damsas newhabitatfor WestAfricansavannah: Claffey

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