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504 Pages·1971·27.914 MB·English
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TO MY PARENTS WHO MADE EVERYTHING POSSIBLE 1 DALE'S AN INTRODUCTION TO SOCIAL BIOLOGY Edited by SUSAN DALE, B.Sc, Head of Biology, Strode's School, Egham FOURTH EDITION LONDON WILLIAM HEINEMANN · MEDICAL BOOKS · LTD. First published September 1946 Reprinted October 1947 Second Edition, March 1949 Third Edition, January 1953 Reprinted with revisions, February 1959 Revised reprint, May 1964 Fourth Edition April 1971 PHYLLIS DALE and SUSAN DALE TUNNICLIFFE, 1971 SBN: 433 07060 9 Printed in Great Britain by The Whitefriars Press Ltd., London and Tonbridge INTRODUCTION TO THE THIRD EDITION THERE is an increasing tendency in teaching modern science to try, as it were, to take it out of its laboratory context and set it against the background of human activity, where it properly belongs. Biology, the science of life, particularly lends itself to this treatment. There is a growing belief that every pupil who leaves school should take with him some knowledge of certain biological facts and principles. And this for two reasons : (1) a knowledge of these facts is essential in order to live a satisfactory physical life, and (2) appreciation of certain broad biological principles enables the individual to use rightly that raw material from which he gradually constructs a personal philosophy of living. In so far as these two issues may be separable, the first is dealt with mainly in Chapters IV to IX inclusive, while the second is more the concern of the remainder of the book. This book is the outcome of a series of discussions held with the combined Sixth Forms (Arts, Science, and Modern) of a large boys' secondary school, and devised with this twofold object. As the discussions proceeded it became apparent that boys of this age had their interests both in the biological aspects of existence, and in man as an animal, already broadly conceived. Consequently the scope of the discussions tended to become ever wider ; and, though the primary object of the course was not lost sight of, much material not strictly biological was discussed. Aspects of politics, theology, morality, spiritualism, propaganda, philosophy, and kindred subjects, came under review. The author soon realised that a degree course in biology was a not altogether adequate preparation for the task he had undertaken, and a considerable amount of supplementary reading had to be done to enable some measure of authoritativeness to be brought to the discussions. It is hoped that this book will help to satisfy a definite need for Sixth Form " General Courses " in schools, and cultural courses in adult education. Obviously a work of this nature must necessarily be experimental. Criticisms and suggestions from teachers and others as to how the book may be improved will therefore be most welcome. A. D. BAKE WELL, 1958. ν INTRODUCTION TO THE FOURTH EDITION WHEN my father wrote this book it was at the beginning of the movement to break the straight jacket of specialisation in Sixth Form teaching. Now such an ideal is widely accepted. It gives me particular personal satisfaction that I am able to continue his aims and to help further his work which was cut short by his tragically early death in 1960. I hope that, with the appearance of a new edition, this book will continue to be found useful for general courses in the Sixth Form as my father envisaged, such as the courses followed for the N.U.J.M.B. "A" Level paper in General Studies. It is hoped that this book will contribute to foundation courses in psychology and sociology which are being introduced into Sixth Form curricula; to the Social Biology option for Teachers' Certificates and B.Ed, degrees; Health Education in postgraduate education studies; and also in introductory courses of interdisciplinary degrees. Not only will students of biological sciences find much in this text which is useful, but so also will those who are interested in the biology of their own species and its interaction with other living organisms. I have attempted to update the contents of this book within the structure of the original text, and added a short appendix concerned with inheritance and DNA. S. D. T. WINDSOR, 1971. vi ACKNOWLEDGMENTS TO THE THIRD EDITION THE author wishes to acknowledge the generous help he has received from many friends during the preparation of this book. In particular I would like especially to thank the following :— My friend and colleague, C. B. Shore, for preparing Figs. 56, 122, 198, 200, 202, 204, 205, 206, and 210 ; my former pupil, H. Wade, for making the drawings for Figs. 11, 12, 13, 17, 22 and 62 ; my pupil, A. J. Grunau for making the drawing for Fig. 73 ; Dr. C. F. Robinow for taking photomicrographs specially for Figs. 59, 68, 75 and 128 ; Dr. Miles for providing the original photographs from which Fig. 105 was made ; the Director of the Whitechapel Clinic for the photo- micrographs for Figs. 131 and 133 ; Mr. L. R. Broster for Fig. 53 ; Dr. Darlington for providing the original photograph for Fig. 77 ; Messrs. Cadbury Ltd. for supplying photographs for Figs. 135 and 175 ; Messrs. Lyons and Co. for supplying material from which Fig. 174 was drawn ; The Liebel-Flarsheim Co. for supplying a photograph of the Kettering Hypertherm in Fig. 132 ; the Kodak Medical Dept. for the original of Fig. 147 ; the Director of the Wellcome Research Institute for the original photographs from which Figs. 154 and 161 were made ; Dr. C. E. Keeler for original photos of a piebald negro for Fig. 96 ; Mr. M. C. Merrill, Chief of Publications, U.S. Dept. of Agriculture, for material to enable Fig. 180 to be made ; Prof. J. Trefouel, Director of the Pasteur Institute, Paris, for the original photograph for Fig. 148 ; the Controller of H.M. Stationery Office for permission to reproduce Figs. 157, 158 and 193, also Tables IX, XX, XXI, XXII and XXIV ; the Editor of" Post-Graduate Medicine " for Fig. 165 ; Parke-Davis & Co. for Fig. 162 ; the Central Office of Information for Figs. 156 and 190 ; Mr. Giles, Registrar of the British Optical Association, for assistance in connection with Fig. 145 ; and Dr. F. Sherwood Taylor for advice concerning the invention of the thermometer and the ophthalmoscope. I wish also to thank those authors and publishers who have permitted the reproduction of illus- trations from their books, details of which are given in the text or after the illustrations concerned. I should like to thank A. J. Low, M.Sc, for reading through the MSS. and making valuable criticisms, and S. F. Marshall, M.A., B.Sc, whose constructive comments materially affected the final form of the book ; also Prof. Charles Singer for reading and criticising Chapter VIII, and Dr. Frank Wokes for reading and criticising Chapter IX. I owe a great debt of gratitude to my friends Miss R. Askew, B.Sc, and Messrs. R. B. Whellock, B.Sc, H. L. Hoyce, M.Sc, D. Riley, B.Sc, vii viii A C K N O W L E D G M E N TS and L. P. Ibbotson, M.A., for the manner in which they performed the very onerous task of correcting the proofs. My wife has been a constant source of encouragement while I have been preparing the book, and she has helped me in a hundred and one ways. For this, and for her patient forbearance, I give her my sincere thanks. Finally, I would pay tribute to my publishers, with whom my relations have been most cordial. Especially must I thank Dr. Johnston Abraham who has read through the book at all stages and has made many valuable suggestions. ACKNOWLEDGMENTS TO THE FOURTH EDITION I WISH to thank the many people who gave me both help and encouragement whilst I was preparing this new edition. In particular I would like to thank the following:— Professor Michael Abercrombie ; my friend and my father's former pupil Dr. Brian Cole for assisting with Chapter VII ; Mr. W. J. Garnett for providing his photographs from which Figs. 31 and 65A were made and for making the black and white prints for Figs. 54 from my father's colour transparencies which are in Mr. Garnett's collection ; Mr. C. Green of the Thoracic Surgery Unit, Guy's Hospital, for advice on heart-lung machines ; Dr. John Napier for advice on Fig. 19 ; Dr. Τ. A. M. Nash for kindly supplying recent information about the situation at Anchau ; Dr. Kenneth Oakely for advising on aspects of Chapter II ; Mr. John Phillip for taking the electronmicrographs from which Figs. 29 and 213 were made ; Miss Margaret Sweeting for helping with information for Chapter X ; Dr. Waddy for discussing aspects of Chapter VII ; Dr. Weidman for his invaluable criticism of Chapter V ; The Director of the Office of Health Economics ; The Director of the Wellcome Medical Museum for helping me obtain photographs from which Figs. 155, 164 and 173 were made. To the publishers, individuals and institutions who have given me their permission to use certain figures I want to express my sincere thanks ; the sources of these illustrations are acknowledged in the appropriate caption. I would like to record my gratitude to Professor J. E. Webb and the Zoology Department of Westfield College, University of London, for their counsel and tolerance. Just as she was to my father, my mother has been a constant source of encouragement during the years whilst I have been preparing this edition. Finally, I must thank my husband, Denis Tunnicliffe, for his advice and forebearance, viii A C K N O W L E D G M E N TS and L. P. Ibbotson, M.A., for the manner in which they performed the very onerous task of correcting the proofs. My wife has been a constant source of encouragement while I have been preparing the book, and she has helped me in a hundred and one ways. For this, and for her patient forbearance, I give her my sincere thanks. Finally, I would pay tribute to my publishers, with whom my relations have been most cordial. Especially must I thank Dr. Johnston Abraham who has read through the book at all stages and has made many valuable suggestions. ACKNOWLEDGMENTS TO THE FOURTH EDITION I WISH to thank the many people who gave me both help and encouragement whilst I was preparing this new edition. In particular I would like to thank the following:— Professor Michael Abercrombie ; my friend and my father's former pupil Dr. Brian Cole for assisting with Chapter VII ; Mr. W. J. Garnett for providing his photographs from which Figs. 31 and 65A were made and for making the black and white prints for Figs. 54 from my father's colour transparencies which are in Mr. Garnett's collection ; Mr. C. Green of the Thoracic Surgery Unit, Guy's Hospital, for advice on heart-lung machines ; Dr. John Napier for advice on Fig. 19 ; Dr. Τ. A. M. Nash for kindly supplying recent information about the situation at Anchau ; Dr. Kenneth Oakely for advising on aspects of Chapter II ; Mr. John Phillip for taking the electronmicrographs from which Figs. 29 and 213 were made ; Miss Margaret Sweeting for helping with information for Chapter X ; Dr. Waddy for discussing aspects of Chapter VII ; Dr. Weidman for his invaluable criticism of Chapter V ; The Director of the Office of Health Economics ; The Director of the Wellcome Medical Museum for helping me obtain photographs from which Figs. 155, 164 and 173 were made. To the publishers, individuals and institutions who have given me their permission to use certain figures I want to express my sincere thanks ; the sources of these illustrations are acknowledged in the appropriate caption. I would like to record my gratitude to Professor J. E. Webb and the Zoology Department of Westfield College, University of London, for their counsel and tolerance. Just as she was to my father, my mother has been a constant source of encouragement during the years whilst I have been preparing this edition. Finally, I must thank my husband, Denis Tunnicliffe, for his advice and forebearance, AN INTRODUCTION TO SOCIAL BIOLOGY CHAPTER I LIFE IN SPACE AND TIME IT is a peculiarity of the human mind that familiar everyday things are taken for granted and are rarely the subject of speculation. Fami- liarity breeds indifference and mental blindness. For instance, when the water is run out from a bath a whirlpool forms over the plug-hole. Everyone has baths, yet few people will have pondered on the whirl- pool and wondered what causes it, or thought whether the direction in which the water rotates can easily be changed, or what factors determine the direction of swirl. Similarly, gardeners who sow parsnip seed expect parsnip plants to come up, and when the parsnips do come up the gardeners do not bother to think why this should always be so, because they find nothing remarkable in the fact that parsnips always beget parsnips. If seeds gathered from a broad bean plant gave rise to a crop of onions, or if a doe rabbit gave birth to a litter of kittens, it would be considered wonderful, yet the fact that plants and animals always reproduce their own kind is equally wonderful. Again, the passage of time is a familiar experience for everyone ; so familiar, indeed, that its nature is rarely considered, except by the most pene- trating thinkers. That is how it is with the phenomenon of Life. Everyone is born alive into a living world, and soon recognises that there are two kinds of matter: the living and the non-living. The difference is simply explained by the assumption that the former possesses a property called Life which the latter lacks. Here, for most people, speculation stops. Life for them is a fundamental, and they are not concerned with an exact definition or a close examination of its attributes. They accept it as they find it. It can be stated at once that precise ideas on the nature of life have not yet been arrived at. Neither the experi- mental biologist nor the philosopher has obtained a satisfactory answer to the question : " What is Life ? " ; nor can either explain exactly wherein lies the difference between a sparrow chirping under the eaves and that same sparrow dead on the garden path. It may be that there is no answer beyond the simple statement that the one has life and the other has not ; in other words, Life may be a fundamental incapable of explanation in terms of other phenomena. Here all that can profitably be said is that Life is an attribute possessed by certain things (living organisms) for a certain period (from the time of their origin until death), and that it appears to be ι 2 SOCIAL BIOLOGY intangible, massless, and invisible, and can only manifest itself by utilising matter. It would thus seem to be a kind of abstraction, like Beauty. The matter imbued with life, which is, of course, a different thing from Life itself, is still susceptible to the ordinary laws of physics and chemistry, but in addition seems to have certain attributes of its own. It is not impossible that Life may exist " neat " so to speak, without permeating matter at all. The idea of a centralised pool of Life, to which all Life returns after the death of the host-matter, is attractive in this connection. In this context we may also ask, " What is Death? ". It is thought popularly that death, in man, is the combina- tion of the effects of the cessation of the heart beat and breathing. Death is regarded as the departure of the Life Essence. Scientifically the death of the body as a whole, called somatic death, and the death of individual body cells are recognised as two different phenomena. However, such definitions are arbitrary because different cells of the body die " individually " faster than others in different locations in the body. The cells of the central nervous system are far more vulner- able to oxygen lack than are any other cells. They become function- less, that is they die, whilst other cells in the body are still alive bio- logically. Thus, respiration and circulation may be restarted in a conventionally dead body so that it appears to be alive again but the body will remain unconscious. It will be just a respiring mass of organic matter. The medical techniques which restore and the machines which maintain circulation and respiration pose vast problems which previously have been confined to the realms of science fiction. What is the legal position? Now laws are passed defining the conditions to be met for a " legal " death to be deemed to have occurred. The mind boggles at the possible results of a situation where a man is able to " revive " the dead. At the touch of a switch the working of a life- maintaining machine could cease. Would this be homicide on the part of the switch operator? This is a frightening ethical situation revolving around the reversibility of the revival process created by man's technology. All living bodies are derived from other living bodies by some form of reproduction, which is essentially the separating off from a parent organism of a bit of material with its contained Life. It is significant that, as far as we know, Life never enters into possession of previously non-living matter and makes it " come alive ", although living material may grow by incorporating into itself non-living food. Whether the Life in a grown man is the same Life that was in him when he was a child it is impossible to say ; and whether it is necessary for the " Life essence " in a child to " grow " to keep pace with the growth of body substance also cannot be determined, because there is no means of measuring " quantity " of Life. All that is known for certain is that the manifestations of Life certainly go on at different rates both in

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