Culicipedia: Species-group, Genus-group and Family-group Names in Culicidae (Diptera)

400 Pages·2018·2.972 MB·English

by  Ralph E. Harbach

Culicipedia: Species-group, genus-group
and family-group names in
Culicidae (Diptera)

Culicipedia: Species-group,
genus-group and family-group
names in Culicidae (Diptera)
Ralph E. Harbach
Department of Life Sciences
Natural History Museum
Cromwell Road
London
SW7 5BD
UK
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© Ralph E. Harbach 2018. All rights reserved. No part of this publication may be re-
produced in any form or by any means, electronically, mechanically, by photocopying,
recording or otherwise, without the prior permission of the copyright owners.
A catalogue record for this book is available from the British Library, London, UK.
Library of Congress Cataloging-in-Publication Data
Names: Harbach, Ralph E., author.
Title: Culicipedia : species-group, genus-group and family-group names in
Culicidae (Diptera) / Ralph E. Harbach.
Description: Boston, MA : CABI, [2018] | Includes bibliographical references.
Identifiers: LCCN 2018023336 (print) | LCCN 2018028612 (ebook) | ISBN
9781786399069 (ePDF) | ISBN 9781786399076 (ePub) | ISBN 9781786399052
(hbk : alk. paper)
Subjects: LCSH: Mosquitoes--Nomenclature. | Mosquitoes--Classification.
Classification: LCC QL536 (ebook) | LCC QL536 .H347 2018 (print) | DDC
595.77/2--dc23
LC record available at https://lccn.loc.gov/2018023336
ISBN-13: 978 1 78639 905 2
Commissioning editor: Ward Cooper
Editorial assistant: Emma McCann
Production editor: Marta Patiño
Typeset by SPi, Pondicherry, India
Printed and bound in the UK by Bell & Bain Ltd, Glasgow
You, whose name has not been formed by anyone,
Whose proof not even intellect can grasp,
To label you would be a loathsome act,
For you are far removed from genus and species.
Nosiri Khisrav
Persian Poet
(1004–1088)

Contents
Preface ix
Introduction xi
Arrangement, Format and Definitions xiii
Symbols, Abbreviations, Words and Phrases in this Tome xiv
Declension (Inflection) and the Formation of Names xiv
Classification xvii
Section 1: Species-group Names 1
Names beginning with letter A 3
Names beginning with letter B 19
Names beginning with letter C 26
Names beginning with letter D 38
Names beginning with letter E 43
Names beginning with letter F 46
Names beginning with letter G 52
Names beginning with letter H 57
Names beginning with letter I 61
Names beginning with letter J 65
Names beginning with letter K 67
Names beginning with letter L 70
Names beginning with letter M 77
Names beginning with letter N 89
Names beginning with letter O 95
Names beginning with letter P 97
Names beginning with letter Q 110
Names beginning with letter R 111
Names beginning with letter S 115
Names beginning with letter T 127
Names beginning with letter U 133
Names beginning with letter V 135
Names beginning with letter W 138
Names beginning with letter X 139
Names beginning with letters Y and Z 140
Section 2: Genus-group Names 141
Section 3: Family-group and Other Suprageneric Names 157
Section 4: Species-group Names Formed from Personal Names 161
Section 5: Species-group Names Formed from Geographical Names 186
Section 6: Genus-group Names Formed from Personal Names 223
Section 7: Some Interesting Species- and Genus-group Names 229
Some Interesting Species-group Names 231
Some Interesting Genus-group Names 238

vii
viii Contents
Section 8: Phylogenetic Classification of Composite Genus Aedes 240
Species-group names 242
Names beginning with letter A 242
Names beginning with letter B 246
Names beginning with letter C 248
Names beginning with letter D 251
Names beginning with letter E 253
Names beginning with letter F 254
Names beginning with letter G 256
Names beginning with letter H 257
Names beginning with letter I 259
Names beginning with letter J 260
Names beginning with letter K 260
Names beginning with letter L 261
Names beginning with letter M 263
Names beginning with letter N 266
Names beginning with letter O 268
Names beginning with letter P 269
Names beginning with letter Q 273
Names beginning with letter R 273
Names beginning with letter S 275
Names beginning with letter T 278
Names beginning with letter U 280
Names beginning with letter V 281
Names beginning with letters W, Y and Z 282
Genus-group names 283
Section 9: Changed Spellings of Species-group Names 288
Some Thoughts about Mosquito Taxonomy and Nomenclature 289
Literature Cited 291
Appendix 1 Current Higher-level Classification of the Culicidae 365
Appendix 2 Multi-generic and Traditional Classifications of the Genus Aedes 369
Appendix 3 Table of Changed Spellings of Species-group Names 373
Preface
The idea for this volume began with the development of 1905e; Edwards, 1932a; Stone et al., 1959; Knight &
an ‘index’ for the online Mosquito Taxonomic Inventory Stone, 1977), comprise a practical amalgamation of taxo-
(http://mosquito-taxonomic-inventory.info/). This involved nomic history, rules that govern the formation of names,
the acquisition of all articles in which the names that mosquito nomenclature and classification. The tome pro-
comprise the nomenclature of the family Culicidae were vides an up-to-date resource for linking names with one
originally published. As the work progressed, it become another and higher taxonomic groups while providing a
apparent that the catalogue of world Culicidae (Knight & concise account of all names (nearly 6000) used in the
Stone, 1977), its supplements (Ward, 1984; Gaffigan & Culicidae, their origins and taxonomic status.
Ward, 1985; Ward, 1992), and consequently the online The need for this tome coincides with a time when
catalogue of the Walter Reed Biosystematics Unit (http:// few professional traditional taxonomists are left to assist
www.mosquitocatalog.org/), which was copied verbatim and advise the new generation of molecular taxonomists
from the world catalogue and its supplements, contain in the basic principles and practice of mosquito tax-
many errors and include only species-group, genus-group onomy. For this reason, this compilation is intended to be
and currently valid family-group names. The tome pre- a valuable resource for students, researchers, entomolo-
sented here is a comprehensive compilation of all scientific gists, librarians and anyone interested in mosquito tax-
names introduced at all levels of classification for Culicidae onomy, systematics and nomenclature.
from the official start of zoological nomenclature on 1 Many colleagues have contributed directly or indir-
January 1758 until 30 June 2018, with corrections to ectly to the consummation of this work. I am grateful to
errors contained in the world catalogue and its supple- all persons who aided in any way, but I would like to par-
ments. Corrections and additions made to information con- ticularly thank the following colleagues. First of all, this
tained in those publications are too numerous to list tome owes its existence to the exhortations of Lorna
individually. In general, however, corrections have been Culverwell and Simon Rycroft. Their suggestions and en-
made to literature citations, page numbers of taxon citations, couragement provided the impetus to transform an in-
and the names of authors, species and type localities. tended ‘index’ into this monograph. I am equally grateful
Culicid nomenclature may be defined as a lexicon of to the Library and Archives staff of the Natural History
generally latinized names applied to groups of mosquitoes Museum, especially Harriet Campbell Longley, for assist-
(actual or perceived monophyletic taxa) that constitute the ance in locating articles and obtaining numerous elusive
hierarchical classification of the family. The groups (taxo- publications through the inter-library loan service that
nomic ranks) whose names are regulated by the International aided the preparation of this tome. A huge thank you to
Code of Zoological Nomenclature (International my revered colleagues who took time from busy schedules
Commission on Zoological Nomenclature, 1999) are the to review and provide advice and comments on various
species, genus and family groups. The species group in- parts of this work: Art Borkent, Lorna Culverwell, Bruce
cludes names given to subspecies and species, the genus Harrison and Christian Kammerer. Thanks also to David
group includes names of subgenera and genera, and the Notten for discussing provisions of the International Code
family group includes the names of tribes, subfamilies and of Zoological Nomenclature, and very special thanks to
families. Emphasis is given to accurately reproducing the Christian Kammerer for his critical appraisal and schol-
names given to members of these three groups and cor- arly comments on text dealing with the application of
rectly associating and listing the original literature sources Latin and Greek in the formation of names. Chen Bin and
in which they were published. Ichiro Miyagi are gratefully acknowledged for providing
Much of what has happened in mosquito taxonomy information about names proposed in Chinese and
since the start of zoological nomenclature has no value be- Japanese works, respectively. My particular thanks to free-
sides historical interest: notwithstanding, the present work lance copy editor Val Porter for diligently and thoroughly
is a unique compendium of mosquito taxonomy. The con- reviewing the manuscript and meticulously checking the
tents and composition, unlike the four catalogues of the references cited. I hasten to add that any lapses and errors
Culicidae published during the 20th century (Theobald, are strictly my responsibility.

ix