Corrélation of the early Albian-late Turonian radiolarian biozonation with planktonic and agglutinated foraminifera zonations in the Pieniny Klippen Belt (Polish Carpathians) Marta B/^K Institute of Geological Sciences, Jagiellonian University, Oleandry 2a, 30-063 Krakôw (Roland) [email protected] Krzysztof BP^K Institute of Geography, Cracow Pedagogical University, Podchor^zych 2, 30-084 Krakôw (Roland) [email protected] B^k M. & B^k K. 1999. — Corrélation of the early Albian-late Turonian radiolarian biozona¬ tion with planktonic and agglutinated foraminifera zonations in the Pieniny Klippen Belt (Polish Carpathians), in De Wever P. & Caulel J.-P. (eds), InterRad VIII, Paris/Bierville 8- 13 septembre 1997. Geodiversitas 2^ (4) : 527-537. ABSTRACT Marine pelagic deposits of the lower AJbian to upper Turonian inter\'al in the Polish part of the Pieniny Klippen Belt are relativcly rich in radiolarian fauna as well as in planktonic and agglutinated foraminifera. The proposcd KEYWORDS radiolarian zones (Holoayptocanium barbui^ Hemiayptocapsa prepolyhedra Cretaceous, Radiolaria, and Hemicryptocapsa polyhedra) hâve been correlated with planktonic forami¬ Foraminifera, nifera biozonarion (from TidnelUt primula to Dicarinella prhnitiva zone.s), integrated biostraiigraphy, and with agglutinated foraminifera biozonation (from Haplophragmoides Pieniny Klippen Belt, Carpathians. nonioninoides to Vvigerinarnmma ex ^r.jankoi zones). RÉSUMÉ Corrélation des biozones à radiolaires de l'Albien inférieur- furonien supérieur avec celles à foraminifères planctonicjues et agglutinés dans la région des klippes piénines (Carpathes polonaises). Les dépôts marins pélagiques de PAlbien inférieur-Turonien supérieur de la zone dc.s klippc.s piénines sont relativement riches en faune de radiolaires, ainsi que de foraminifères planctoniques et agglutinés. Les zones de radio¬ MOTS CLÉS laires proposées (Holocryptocanium barbais Ifvniicryptoctipsa prepolyhedra cr Crétacé, Radiolaria, Hemiciypwcapsa polyhedra) sont en corrélation directe avec les biozones de Foraminifera, foraminifere.s planctoniques (de la zone Ticinelln primula à la zone biostratieraphie intégrée, zone des klippes piénines, Dicarineüa primitiva) et celles des foraminif^res agglutinés (de la zone Carpathes. Haplophragmoides nonioninoides à la zone Uvigerinammina ex jankoï). GEODIVERSITAS • 1999 • 21 (4) 527 B^kM. &B^k K. INTRODUCTION lyses. A detailed description of profiles, sample location, and lithology is given in papers by Detailed micropalaeontological studies of marine HL (1992, 1993» 1998) and M. B^k (1995, pelagic Cretaceous deposirs hâve been carried out 1996a, b, 1997). In 18 of these sections, radiola- in rhe Picniay Klippen Belt during the last 15 rian skelctons hâve been foimd together with years. A part of thcse concerncd bioscracigraphy, forarninifera. and thc best sections (15) were based on differejic groups oFmîcrofauna; forami- used lierc for corrélation purposes (Figs 1,2). nifera (Birkenmajcr & Jednorowska ]987i In thc sections examined, the microfauna has Gasihski 1988i Kostka 1993; K. Bi^k 1998), nan- been recovered from rhe deposirs of the noplankton (Dudziak 1985)» dinoq'srs (Jamihski Kapusnica and Jaworki formations. Liihological 1990) and radiolaria (M. B^k 1995, 1996a, b). features, thickness and âge of these lithostratigra- However, only very few studies comprise an intc- phical unies, distinguished by Birkenmajer grated biostraiigraphy of rhese deposirs (1977, 1987), and Birkenmajer 6c Jednorowska (Birkenmajer Z*/1979; K. BtjkfY^ïZ. 1995). (1984) and presented below, concern only the The Crccaccous dcposits ol thc Picniny Klippen studied sections. Bclt providc ihc bcst marctial lor iTc intcgratcd biostraiigraphical studies in the whole KaPIISNTCA l-OrU'tATKlN Carparhians, because rhey represenr pelagic and This formation is subdivided in two members, hemipelagic calcareous-marly faciès, and contain the Brodno Member and the Rudina Member, abundant micmfauna. These depos’us hâve been both represented in the sections investigated. formed în the Pieniny Klippen Basin The Brodno Member. Ir has been identified (Birkcnmajer 1977), a pari of the Penninic only in the one section as a 40 cm thick layer of Ocean-northcrn part of Western Tethys black maris, ? upper Aprian in âge. The radiola- (Fourcade étal. 1993) (Fig. lA). The sédimenta¬ rian launa is présent burver)’^ rare in these depo- tion during Albian thtough Campanian rook sits. place iindcr deep-water conditions, ar outer shelf The Rudina Member. (1-20 m thick; lower- through Jower bathyal/abyssal depths, above or upper Albian) It is represented by green-grey and near CCD (Birkenmajcr & Gasihski 1992; black maris and inarly shales with niarly liniesto- K. B^k 1995a, b). l'hcy crop oui in a highiy tcc- ne imercalaiions, with red maris appearing in the tonically deformed zone, known as the Pienîny upper part. Abundant and well-prcserved radio- Klippen Belt, rhat séparâtes the Innct ftom rhe larian skeletons are présent in green and black Outer Carparhians (Fig. IB, C)'. maris and niarly shales. The présent aurhors propose here an integrated local biostrarigraphy based on planktonic Fora- JAWOKKI POKMAI'ION minifera, agglutinaicd l'uraminifera and Radio- Deposits of the Jaworki Formation hâve been laria for early Albian rhrough laie Turonian lime examined in sections belonging to ail Klippen span. rhe corrélation is based on studies of successions (Fig. 2). microfauna from the sanie samples wirhin the The Brynczkowa Mari Member. (2-22 m thick; same sections. Foraminifera and Radiolaria are Vraconian-middle Cenomanian) it consists of relativcly abundant in the studied deposirs. green or grey-green maris with thin scarce red mari inrercalation. Moderarely to poorly preser- ved radiolaria are rare. GEOLOGIC^\LSETTING AND The Skalski Mari Member. (4-15 m thick; RADIOLARIAN HORIZONS middie-upper C^cuonvanian) It consisis of varie- gaced maris, iniercalated with green, red and grey Twenty five sections locaced at Niedzica, maris. Hxcept of thc Altana Shale Bed, thc radio- Kroscienko, Szczawnica, Jaworki, Sromowce, larians are présent in this member but are rare Dursztyn, Szaflary and State Byscre hâve been and mosily poorly preserved. chosen for integrated micropalaeontological ana¬ The Altana Shale Bed. (0.4-3 ni thick; 528 GEODIVERSITAS • 1999 • 21 (4) Radiolarian and foraminiferal biozonation in the Pieniny Klippen Belt Fig. 1. — Location of the study ârea: A, In the Western Tethys {rectangle B) (map of late Cenomanlan palaeoenvironments after Fourcade et ai 19&3; simplîfied); B. C. in the Carpathians (Pieniny Klippen Bell in black). D. in lhe Pieniny Klippen Belt. Legend. 1. investigated outcrops; 2. symbols ot outcrops; 3. more important dislocations. List ot outcrops (from west to east): Sb, Stare Bystre; Ki. Kietowy stream: Sz, Szaflary quarry, Lor Lorencowe Klippes: Csk, Czerwona Skaia, Zt, Ztobowy Creek; Nd. Niedziczanka slream; Kos, Kosarz.yska Valley; McO Macelowa-Osice; Pg. Podskalnia Gôra ML Mag. Magierowa Klippe; Or. Orlica Hlll: Bw. Bukowiny Valley: Bd. Bukowiny Hill. Cenomanian/Turoniaiî boundary) h consists of danc: wcll-preserved siliceous specimens are pré¬ black-blue, grecnish and black shales and marly sent in green and grcy shales and marly shales. shales. Radiolarians are abundant bur low-diver- The radiolarians arc also abundant in black sified and poorly preservcd. Their pyritized ske- shales but are often diftlcult to déterminé, becau- Ictons are mostly found within the Czorsztyn sc of Fe-oxide coating and pyritization. Succession deposits. The Trawnc Menibcr. (5 m, lowcr-middle The Magierowa Mari Member. (9 m; Cenomanian) It consists of gre>'-grccn maris and Cenomanian/Turonian boundar)^ Il is reprcscn- shaly maris with sandstonc intercalations. icd by shales, marly shales, maris, and thin-bcd- Radiolarian fauna is scarce: only a few poorly ded marly ümestonc of grey to green colour with preserved specimens bave been found. black alternations. Radiolarian fauna is abun- The Sneznîca Siltstone Member. (3-? 40 m; GEODIVERSITAS • 1999 - 21 (4) 529 I B^k M. & B^k K. Fig, 2. — Lithological columns of the studied sections in relalion lo the âge of deposits (dashed Itne cortesponds \o the Cenomanian/Turonian boundary) and tbeir atlnbutton to Klippen successions and sedimenlary zones (Pieniny Klippen Basin recons¬ truction after Birkenmajer (1977)]. Lithological units invesligated |llthoslratlgraphy aller Birkenmajer (1977, 1987), Birkenmajer & Jednorowska (1984)]: 1, Lorencowe Chert Bed; 2. Osice Siltslone Member; 3, Macelowa and Pusteinia Mari rnenibers 4, Trawne Member; 5, Sneznica Siltslone Member, 6, Altana Shale Bed; 7, Magierowa Member; 8, Skalski Mari Member; 9, Brynczkowa Mari Member; 10. Rudina Member: 11, Brodno Member; 12, Pieniny Limestone Formation. Scale bars: 5 m. except of Branisko Succession sections, 1 m. GEODIVERSITAS • 1999 - 21 (4) 530 Radiolarian and foraminiferal biozonation in the Pieniny Klippen Belt middlc Cenomanian-middle Turoiiian) It is rcpre- the familles Actinommidac, Pracconocar)'om- scntcd by altcrnating bright blue-grey and green midac, Xiphostylidac (gênera Staurosphaeretta shaly maris widi thin-bcddcd slltstone and sand- and Tviaciomii) y and Dactyliosphaeridae stone, and bright-grccn pclitic Ümcstonc intercala¬ {Diictyiîodisciis^ Godia and Dactyliosphaera). tions. Radiohiriims arc raie in this nicmbcr, but Other Spumellaria bclong to the généra the upper Cenomanian deposirs arc cnriched in Ilexapyrûniis^ Cavdspongla., Pseudoaulophacus, wcll-preser\’cd, pyi iri/.ed radiolruian skclcron. Patellulû^ Paroniiella and CrucelLi. The Macelowa Mari Member. (15-70 m; lower Within the Klippen successions investigated, the Cenomanian-iSantonian) Ir corisists nnjstly of red radiolarians are most abundant and diversificd in maris and marly Jimesronc vvith thin intercala¬ the Czorsztyn Succession deposits (ourer shelf- tions of greenish or bltiish siltstones and sand- upper barhyal environment), becoming generally stones which appcar in its lower and upper parts. scarce in the deposits of the Pieniny Succession Poorly preserved and mainly calcified radiola- (lower bathyal environments, close to Calcium rians are présent in this member. They are diver- Compensation Depth). sified, especially those lound in a horizon The radiolarian specimens are most abundant corresponding to the lower-middle Turonian. and diverse in green and black marly shalcs, The Pustelnia Mari Member. (6-45 m; middle maris and limestones ol the Kapusnica and Cenomanian-Santonian) Ir consists of pure Pomied/nik formations, rhe Magierowa Member brick-red, strongly lectonized maris wïthout clas- and ihe Aliana Shale Bed of rhe Jaworki ric intercalations. Radiolarian skcletons are vciy Eorination. Thcy arc scarce in red and variegated scarcct poorly preservedt and maixily calcified. maris and shales of the Macdowa Mari, The Lorencowe Chéri Bed. (3 m; upper Brync/kowa Mari and the Skalski Mari members Santonian-lower Campanian) h is represented by ol the Jaworki Formation. This itilght he dépen¬ a two-meter-thick complex consistiiig of altema- dent on lirhology and iliagenesis factors respon- ting variegated maris (mostly red) wirh rhin sible for préservation of siliceous microfossils; intercalations of lighr-green limestone. however, radiolarian fauiias radiarion and turn¬ Radiolarian skeletons are also scarce, poorly pre- overs could be also taken inro account. Both ser\’'ed, mostly calcified. causes might reflecr an ecological response of marine biota to global cUmace and sea level changes during mid-Crctaceous rimes. RADIOLARIAN ASSEMBLAGES Two great changes in the radiolarian assemblages Seventy seven radiolarian species hâve been hâve been retorxied within the srudied deposirs: determined from the Cretaceous deposics ran- 1. Beginning of an important first radiation of ging from lower Albian to Santonian of the Radiolaria occurred during the middle Albian Pieniny Klippen Bclt. 20 gênera and 53 spedes {Ticmella priwula foraminiferal zone» K. Bijk of Nassellaria, and 11 généra and 24 species of 1992)' as maiiy new species had rheir first appea- Spumellaria were recognised (M. Bqk 1997). rance in the middlc and rhe lare Albian, The The assemblages, in ail Klippen successions maximum of différentiation in the radiolarian mvestigated, are dominated by cryptoccphalic assemblage is obsers^ed in the Vraconian deposits and cryptothoracic Nassellaria belonging ro the (Rotalipara tidnensis through the Rotalipora gênera Holocryptoaviiuni^ Hepiicryptocapsa^ appennhiica-Plevwm<tlina huxtorfi foraminiferal Cryptainphorella. Dorypyle. Hiscocapsa^ zones, K. Bijk 1998). Starting from the upper Trisyringium and Squirtabollum, Multiscgmcntcd part of the It appennimat-P. hiixtorfi foraminitc- Nassellaria arc also common being represented ral zone towards the Cenomanian/Turonian by the gênera Dictyomitra, Thanarla, boundary, a relative decrease in the number of Pseudodictyomitm^ Stichomitm, Xrtus, Obeliscoites species occurred. As a rcsult, the radiolarian làu- and Torculum. Spumellaria are less common; the nas in the Czorsztyn, Branisko and Niedzica suc¬ most abundant are the specimens belonging to cessions show great similarities. The same GEODIVERSITAS • 1999 • 21 (4) 531 B^k M. & B^k K. characteristic taxa from rhe Riborder Nassellaria the Cenomanian deposits; planktonic Foramini- (both cryptorhoracic and cryptocep halle forms) fera prcvail there. with dominating rotaliporids, represented by the généra Holocryplocanium, associated with représentatives of the généra Hemicryptocapsa, Squinclbollum and Cryptam- Pmeglo ho tru Haï fua Hedbergelht^ Wbile in ella, phorella arc the most abundant, svhilc the gênera Glvbiga inellûidtSy HcteroheUx and Shuckoina. Dictyornitray Pseudodictyomiiray Thanarla, Ratio of calcareous to agglutinated bcnihic forms Stichomitrity Toradumy atul Xitm arc also com- is different in pelagic and hemipelagic deposits of mon. this stage. Agglutinated taxa prcvail (more than 2. The nexr change in radiolarlan assemblages 80% of benthos) in the scaglia rossa-type took place around ihe Cenomanian/Turonlan (Macelowa Mari Member) and fivschoid deposits boundary, Ir surted during the latc Cenomanian (Trawne Member, Sneznica Siltstonc Mentber). (Rotalipora aishniani loraminiferal 7one, K. B;^k being dominated by Plectorecurvoides tiheruanSy 1998). At that time, dic radiolarlan assemblages Recurvoides sp., Bidbohitcidites sp.. Trochamniina from tire Picniny Succession becamc similar to sp., Haplophragtnoides sp.. Spiroplecuimminn those from rhe Czorsztyn, Niedzica and Branisko naparroanriy Arcmlndimina predii, Dorodva sp., successions. Moreover, ir was enriched in formvS Triittda sp., Glomospira sp., and by tubular fi^rms. described so iar only from the Silesian and the Calcareous forms are more frequent in variegated Skole units of the Outer Flysch Carpathians and red pelagic maris (Brynczkowa Mark Skalski {Praeconnearyomma lipmanae^ Godia sp., Mari and Fustelnia Mari membets). However, Diacanthocapsa sp. — sec M. B^k 1994; Corka they are represented by single specimens only. 1996). Agglutinated foraminifers indude the généra Dentalindy Nodosariay Triitix, Lenticulina, MarginulinopnSy Mnrginulinay SaYUcenariUy FORAMINIFERAL ASSEMBLAGES AsîiUùlus. Planulariity Globulinay OolinUy Pyridimu Rantfdimy huebulimmay P/eurostorneP The early and middle VVlbian Foraminifera in the la y ValvH U n e ria, Pu lU n ta » (ha ng u la riuy Pieniny KJippcn Bcit represent a well-diversified GyroidinoideSy Gavelinellay Lingulognvelindla and assemblage. Planktonk forms prcvail, being Fpnnides. represented by hedbergellids (//. dtdrioensisy A quilc different foraminiteral assemblage H. planispira and H. mnplcx)^ Glohgerinelloides appears in black shale faciès close to the bentoneusis^ and ticincllids (Tidnella primnliiy Cenomatiian/Turonian boundary. Planktonic T robeni) (k. Bak 1992). Benthos fs dominated Foraminifera are still frequent in the lower part by cakâreous forms with the most frequent of the shales, being represented predominantly Gyroidimida inJhureMceay Gavcl/nella cenomam- by Rotalipora cushmanh they are accompanied by ca and G, interniedia^ and accompanied by whiteinellids, praegiobotruncanids, hedbergeilids Denîalina sp., Nodosuria sp. and Latticidhja sp. and single form.s from the généra Globigerinel- Agglutinated foraminifoa {predominantly infau- loules and Heterohflix. In a higher pari ol the nal forms) are infrequent in rhese deposirs, with shales, only single forms’ of Praeglohotruncana Dorothia gradata, Xpiroplecfinaîa annectemy delrioensis and Hedbergella detrioensis bave heen Tritaxüt gatdtma and Textularia foeda prevaillng. found. Benthonic forms are dominated by very The upper Albian planktonlc Foraminifera are frequent Lenticulina gaultiria. Tliis form is very numerous and more div'crsified. Besicles the accompanied by single specimens of agglutinated most frequent hedbetgellids, Bitidnella hreggien- foraminifera from the généra BulhobaculiteSy sisy PlanornitUna huxtfjrfiy Glabigerinelloides bento- '['rochamminta HarmosinUy Glomospira, Ammo- nesisy G, ultramicrciy praegiobotruncanids and disais and Rhizammhia. rotaliporids occur there. Bctuho.s i.s very rare, The middlc-laïc Turoniaii foraminifcral assem¬ dominated by calca^eou^s(cid:9632) forms, similar to those blage considcrably difters from the Cenomanian présent in the older assemblage. one. Middle furonian was the acme of Helveto- A very similar assemblage is also characteristic of glohotruncana helvetica and H. praehelvetica 532 GEODIVERSITAS • 1999 • 21 (4) Radiolarian and foraminiferal biozonation in the Pieniny Klippen Belt which characterise this assemblage, while ben- ihonic forms are almost absent. Durlng che late Turonian, bcnihonic forms bccorae more fre¬ quent, and they prcvail in the latest Turonian deposiis. rhc bcnthos is dominatcd by aggluti- nated taxa (more than 90%) with vcty characte* ri.stic Uvigertnammina jankoi^ Recurvoides gnduleytsis. GerochitJnmina Cütivena, Kiirrerul'tna canijonnis, Huplophragmoides c£ bulloides and Bulhohamlites problematims. Calcarcous benthos is represented predominantly by the généra Eponides^ Stensioema and Gyroidinoides. RADIOLARIAN BIOSTRATIGRAPHY Radiolarian spccics wcrc idenufied in more than 200 samples from 18 sections in the deposits of the Pieniny, Branisko, Nicdzica and Czorsznm successions of the Pieniny Klippen Belt (M. B^k 1997). Scvcntccn horizons containing abundanr and well-prcscrvcd Radiolaria hâve been chosen for doser analysis (M. B^k 1999); the Bio- Graph 2.02 computer program (Savary & Guex 1991) based on the Unitary A-Ssociations Method was applied. This program produced a scqucnce of 1 1 U. A. (Hig. 3), which wcrc used for construcring a radiolarian zonal standard (M. Bqlc 1999). Three radiolarian zones and five subzones were proposed for the early Albian dtrough late Turonian rime span (Fig. 3). These arc: (1) the Holoayptocttnium barbui zone wilh Stichornitra Tosilen.sis^ Squinahollum fossile, Thauitrlü veneUi^ Torcubon dengoi and Oheliscoises maximus subzones; (2) the Hemicryptoatpsa pre- polyhedra zone; (3) die Hemwryptoaipsa palyhedm zone, The top of each zone and subzone is defi- ned by the FAD of ihe index taxon and aiso by the base of the overlying zone. Fig, 3. — Occurrence range chan ot index radiolarian apecies recorded In the Albian to Coniacian deposits in the Pieniny Klippen Bell successions, based on their first and the last appearances. and their co-occurrence based on the Unitary FORAMINIFERAL BIOSTRATIGRAPHY Associations. Abbreviations ot zones: S. tosaensis. Stichomitra fosaensrs; S. fos., Squinabollum fossile; T. pulch., Thanajia pulchra, O. maxim,, Obeliscoites rnaximus. 1, local planktonic Extensive biostratigraphical studies based on foraminiferal zonation atter Robaszynski & Caron (1985) modi- fied by K. t3a> (1992,1998), planktonic foraminifera were carried out in Cretaceous deposirs of the Polish pan of the Pieniny Klippen Belt (Alexandrowicz 1966; Jednorowska 1987; Kostka 1993). The local bio¬ Alexandrowicz er 1968a, 1968b; Jednorowska zonations hâve been tied to the zonal standards 1979; Gasihski 1983, 1988; Birkenmajer & of the Western Tethys. GEODIVERSITAS • 1999 • 21 (4) 533 B^kM. &B^k K. The local planktonic biozonation hcre présentée! d'he définition ol the Holocryplocanium barhui (K. B^îk 1992, 1998), is based on studies of zone used in this zonation lollows that of 18 field secrion.s (more than 270 samples) repre- Schaaf’s (1985)., with some modification of its senting deposits from ail sedirnentary zones ol upper limit. The lower boundary of H. hnrbui the Pieniny KÜppen Basin in Roland. The zones zone {- lowcr boundary of the Stichornitra distingnished and their chmnostratigraphic cali¬ tûsaensh subzone) is not wcil defined, because the bration, correspond very well with the Medi- eailicst iAlbian and larest Aptian ( Ticinella wbeni terranean zona! standard as pioposcd by planktonic loraminifetal zone and Haplo- Robaszynski Sc Caron (1995), with only minor pbritgnioldes nonioninoides agglutinated loramini- révision (for more details, see K. B^k 1998). leral zone) deposits arc not well represented in The biozonation based on agglutinaced foramini- the sections investigated, their radiolarian fuuna fera foUows that proposed for Outer Carparhians is rare and poorly preserved. d'he upper bounda- by Geroch & Nowalc (1984), and applied aiso to t)'- ot the H. barhui zone hdls wirhin a lower part the Pieniny Klippen Bclt (K, B^k ei al. 1995). of the Rotalipora reicheli planktonic foraminiferal zone and tbc Bulbohaadite^ problrmaticus agglu- tinafed foraminiteraJ zone (middle Ceno¬ manian). The Sipdnahollum fossile radiolarian CORRELATION OF RADIOLARAN subzone coiucîdes with the Biticinella bfrggiensis ZONATION WITH PLANKTONIC AND foraminiferal zone (Late Albian). The Thanarla AGGLUTINATED FORAMINIFERA ZONES pulthra subzone coincide.s with the Rotalipora tkinemis-R. subtidnertsis forajninifcral zone (laie Varions radiolarian zonations for the latc Cieta- Albian). ITc lower limit of the Toradnw deiigoi ceoLis deposits were proposed depending on the subzone correlates with the upper boundary of area concerned. Tw'O proposed standards calibra- the Ratalipora tiànvnsiS'R. suhtiçinemis foramini¬ ted with chronostrâtigraphy in the western part feral zone. Its upper limit correlates with the of the Tethys deserve spécial aaention. d’he first upper boundary ot the Rotalipora appenaimea standard was proposed by Llumitricâ (1975) for foraminiferal zone (Albian/Cenoinanian bounda¬ the Romanian C'arpachians. Hc rccognised two ry), and also coïncides with the lower limit of the radiolarian assemblages, the oldei assemblage Obeliscoües maximus subzone. and the younger one correlatcd with planktonic The lower boundarv' of ihe Hemicryptocapsa pre~ foraminifcr-a. The Holoçryptocaniam barbui- polyhedra zone corresponds to a lower part of the H. tuberadiitum assemblage corresponds, accor- Rotalipora reicheli planktonic foraminiteral zone ding to L'iiimitrica (I975}> to the Rotalipora (middle Cenomanian). Its upper boundary reicheli-R. ci^diwaai7.ont (latest Cenomanian). (= lower boundary of tlîc Pîtrnscrypîovapsa poly¬ O’Dogherty’s (1994) standard was more dctailcd hedra zone) corresponds to an upper pan of the and compriscd a calibration with the Northern R, cushmani planktonic foraminiferal zone (latc Apeninnes and Beric Cordillera. l'hLs calibration Cenomanian). I be upper boundary of the bas been based on planktonic Foraminifera par- H. polyhedra zone bas not been defined within tially following the zonal standard as proposed by the deposits itwestigated because the Radiolaria Caron (1985), with some modifications. from the latest Turonian and the Coniacian are O’Dogherry (1994) recognised five radiolarian very poorly preserved (mosrly calcified). This zones correlated with the HedbergeUa zone correlates with the Rotalipora cn^hrnani (Barremiari) through the Marginommeana sigali through Dicarinellay>r;w/oW planktonic fbrami- (late Turonian) foraminiferal zones. niferal zones ((cid:9632)= upper part ol Bulhohaculites pro- In this study, wc propose a corrélation of radiola¬ hlematicus and lowcr parc of Uvigerinarmnsna ex rian zones with planktonic and agglutinaced fora¬ gf. jankoi agglutinated foraminiferal zones; late miniferal zones for the early Albian rhrough late Cenomanian through late Turonian). Turonian time span, for the PoHsh part of the It îs hoped that the proposed radiolarian zonal Pieniny Klippen Belc (Fig. 4). standard will bccome a calibration tool to whicb 534 GEODIVERSITAS • 1999 • 21 (4) Radiolarian and foraminiferal biozonation in the Pieniny Klippen Bell LOCAL PLANKTONIC LOCAL RADIOLARIAN STAGES FORAMINIFERAL BENTHONIC ZONES AND SÜBZONES ZONES (1) FORAMINIFERAL M. Bak, 1999 ZONES (2) !9oH Dicarinella primitiva Marginotruncana sigali Turonian Uvigerinammina Helvetogtobotruncana ex gr. jankoi Hemicryptocapsa polyhedra helvetica zone Praeglobotrvncana delrioensis Rotalipora cushmani Bulbobaculites Hemicryptocapsa Roi.ilipora giùü!ihr4fr,eii^s problematicus prepolybedra zone Rotalipora reichell Cenomanian Rotalipora Obeliscoites maximus globotruncanoides R appenninica — -R.appen.^ P.buxt. Torculum dengoi iooH R buxtorfi-R. ticinensis_ R.ticinensis-R.subticinensis Thanarla pulchra Biticinella breggiensis Plectorecun/oides Squinabollum fossile alternans Ticinella primula Stichomitra tosaensis Albian Haplophragmoides nonioninoides (cid:9632) 110^ Ticinella H roberti 4 Fig. 4. — Corrélation of radiolarian biozones and subzones with planktonic and agglutinated Foraminifera zonations in the Pieniny Klippen Belt. PoHsh Carpalhians (chronologicat scale after Robaszynski & Caron (1995)]. 1, local planktonic foraminiferal zonation after Robaszynski & Caron (1985) modified by K. B^k (1992. 1998): 2. local benthonic foraminiferal zonation after Geroch & Nowak (1984) modified by K. Bak étal. (1995). K Bak (1995a). local zonations from orher Carpathian régions vainable éditorial commcnts. Particular thanks could bc correlated. are due to Prof M. Caron (Institut de Géologie de rUniversité, Fribourg) and Dr. P. Dunikrica (Institut de Géologie et Paléontologie^ Université Acknowledgements de Lausanne) who tcvised the manuscript. Mrs The authors are greatly indebted to Prof. K. 1. Chodyri (Institute of Geological Sciences, Birkenmajer (Institute of Geological Sciences, Jagiellonian University, Krakow) helped us in the Polish Academy of Sciences, Krakow) for laboratory part of the work. This research is a GEODIVERSITAS * 1999 • 21 (4) 535 B^kM. &B^k K. contribution of the IGCP 362 Project, Tethyan Btik M. 1993. — l.ute Albian-carly Cenomanian and Boréal Cretaceous. Radiolaria from rhe Czonszryn Succession, Pieniny Klippen Belt, Carpathians. Studia Geologica Polonica 102: 177-20'^. — 1994- — Radiolaria from Cenomanian deposits of REFERENCES the Silesian Nappe near Sanok, Polish Carp.ithians. Bulletin of îhe Polish Academy of Sciences, Eanh Alexandruwicz S. 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