Contributions Toward a Bryoflora of Nevada: Bryophytes New for the Silver State. Part III

2014·8 MB·English

by  John C Brinda

Madrono, VoL 61, No. 3, pp. 253-258, 2014
CONTRIBUTIONS TOWARD A BRYOFLORA OF NEVADA: BRYOPHYTES NEW
FOR THE SILVER STATE. PART
III
John C. Brinda
MO
Missouri Botanical Garden, P.O. Box 299, St. Louis, 63166-0299
Lloyd R. Stark
NV
School of Life Sciences, University of Nevada, Las Vegas, 89154-4004
James R. Shevock
Department of Botany, California Academy of Sciences, 55 Concourse Drive,
CA
Golden Gate Park, San Francisco, 94118=4503
shevock@calacademy org
j .
John R. Spence
National Park Service, Glen Canyon National Recreation Area, Page, AZ 86040=1507
Abstract
Forty-two mosses and three liverworts are reported as new for Nevada. The genera Amblyodon,
Campyliadelphus, Isopterygiopsis, Mesoptychia, Myurella, Plagiobryaides, Plagiobryum, Platydictya,
PseudocampyHum, and Preissia are new for the state of Nevada. Splachnobryum obtusum and
Pseudoleskeella rupestris are now confirmed for Nevada based on recently collected material.
Grimmia texicana is excluded from the Nevada checklist based on a herbarium label locality error.
The bryoflora of Nevada with 355 taxa is comprised of two hornworts, 49 liverworts, and 304
mosses.
Key Words: bryophyte inventory.
The most recent published checklist ofNevada Clark Co.: Toiyable National Forest, Mt.
bryophytes listed 310 taxa (Brinda et al. 2007). Charleston Wilderness, South Loop Trail, 28
Other species new to science occurring in Nevada Sep 2008, 10,040 ft, Brinda 2361b (MO, SIU)
have subsequently been published (Greven 2010; [confirmed by Vadim Bakalin].
Medina et al. 2011; Spence and Shevock 2012). In
Preissia quadrata(Scopoli) Nees [Marchantiaceae]
the third installment ofthis series we add 45 taxa
Clark Co.: Toiyabe National Forest, Mt.
as new for Nevada based on voucher specimens.
lEoaccahtiont,axhoenrbahraisa caitasthioornt, aenndtrnyotfeosronspheacbiimteant CBShreaiprnld2ea00s28t3,o6n110aW,i0l(4dC0eArSfnt,,esBMsr,Oin,SdoaUuN2t3hL5V9L)o.(oUpNLTrVai)l,an2d8
and substrate. Field collecting and inventory
efforts continue to document new taxa for the
Mosses
Nevada bryoflora (Shevock et al. 2005; Spence et
al. 2006). Many of the species reported herein Aloina ambigua (Bruch & W.P. Schimper) Lim-
have been reviewed, examined, or determined by
pricht [Pottiaceae]
pFelrosroansofinvNoolrvtehd iAnmtehreibcrayoPprhoyjteectvo(lFumNeAs o2f00t7h,e ClaTrrakilC,o.4:MVaalrle2y00o6f,F2ir0e30Stfatt,eBPrairnkd,aM7o2u4se(’MsETXaUn,k
2014). UNLV) and Brinda 726 (CAS, MEXU,
UNLV) [confirmed by Claudio Delgadillo].
Liverworts
Amblyodon dealbatus (Swartz ex Hedwig) Bruch
Lophozia heterocolpos (Thedenius) M. Howe & W.P. Schimper [Meesiaceae]
[Scapaniaceae] Elko Co.: Humboldt National Forest, East
Elko Co.: Humboldt National Forest. East Humboldt Wilderness, canyon wall above
Humboldt Wilderness, base of cirque, head- beaver ponds near East Humboldt Highland
waters ofFourth Boulder Creek, 16 Aug 2007, Trail, Third Boulder Creek Basin, 15 Aug
9Bi4l7l5Dofty,leS]h,evOonckse3e0p2y98me(tCaAmSo)rp[hdiecterromciknewdalblys 2U0N07L,V)92[7d5eteftr,miSnhedevboyckBil3l03B9u4ck](.CAOSn, moNiYs,t
and ledges in a subalpine conifer forest. rock soil and litter over rock terraces and cliff
face with Primula Linnaeus, Saxifraga Lin-
Mesoptychia badensis (Gottsche) L. S5derstr5m naeus, and Veratrum Linnaeus within a subal-
& Vaha [Jungermanniaceae] pine forest of Pinusflexilis E. James.
MADROto
254 [VoL 61
Anoectangiiim handelii Schiffner [Pottiaceae] Wilderness, Humboldt National Forest, slopes
Clark CO'.: Spring Mountains: cliff above Pine south ofThird Boulder Creek toward Birdeye
Creek, 4Apr2005, 4500 ft, Brinda andShevock Lake, 14 Aug 2007, 9200 ft, Shevock 30303 (C
112 (CAS, UNLV); hill east ofBlue Diamond, AS); Humboldt National Forest, Ruby Moun-
26 Feb 2006, 3350 ft, Brinda 695a (UNLV); tains, Lamoille Canyon along Ruby CrestTrail
White Rock Spring, 26 Feb 2006, 4825 ft, to Dollar Lakes, 20 Jul 2002, 8975 ft, Shevock |j
Brinda 703 (UNLV); Gateway Canyon, 26 Feb andGlazer22483a(CAS) [determinedbyDavid t
2006, 3865 ft, Brinda 721 (UNLV); south fork Toren]. Open subalpine metamorphic or gra-
of Pine Creek, 27 Apr 2006, 4265 ft, Brinda nitic soils over rock terraces.
1012 (UNLV); Virgin Mountains, Sandstone
Bluffs above the Corral North ofQuail Point, Didymodon eckeliae Zander [Pottiaceae]
28 Jan 2006, 4265 ft, Brinda 587 (MO, UNLV) Carson City.: Clear Creek Range just south of
[confirmed by Richard Zander]; Valley ofFire, highway 50 and 2.3 miles west ofhighway 395,
Mouse’s Tank, 4 Mar 2006, 2040 ft, Brinda 5 Apr 2002, 5200 ft, Shevock 21965 (CAS).
727b (UNLV); Valley of Fire, Mouse’s Tank Douglas Co.: Red Canyon, northern edge of
Trail, 10 Mar 2007, 1965 ft, Brinda 1284 BurbankCanyon ScenicArea, easternslopes of I
(UNLV); Frenchman Mountain, Great Un- Pine Mountain, 7 Apr2002, 6000 ft, Shevocket
I
c(oUnNfLoVrm)i;tyM,u5ddMyarMo2u0n0t6a,in2s1,70Bofwt,lBorfinFidrae,78127 aFlor2es2t0,20Ki(nCgAsSt)o.nLaCnadneyroCno.:RoTaoidyanbeearNatthieonoalld •|
MFoeubnt20a0i7n,s,20B9o5wlft,oBfrFiinrde,a 11726F5eb(C2A0S0)7;, 2M0u9d5dfyt, BKriinngdsatoanndGuBaarnndeSrta3t1i5o4n,(5CASSe,pM2O0)11.,G7r5a3n0itfitc, ;
Brinda 1269 (UNLV). and marble rocks within pinyon pine wood- |
lands and Jeffrey pine forests. ;
Brachythecium turgidum (C.J. Hartman) Kind-
berg [Brachytheciaceae] Drepanodadus polygaeius (Bruch & W.P. Schim- j
Elko Co.: East Humboldt Wilderness, Humboldt per) Hedenas [Amblystegiaceae] j
National Forest, slopes south ofThird Boulder Carson City.: Sierra Nevada, Lake Tahoe Basin,
Creek toward Birdeye Lake, 11 Aug 2007, highway 28 at Bliss Creek between Skunk
9250 ft, Shevock 30324 (CAS, UNLV). This Harbor and Secret Harbor, Shevock 22429
rather glossy appearing species displays a (CAS, UNLV). Lander Co.: Kingston Creek,
distribution pattern of widely disjunct occur- Toiyabe Range, Toiyabe National Forest, 30 j
rences across the Intermountain West. The May 2002, 7400 ft, Shevock and Glazer 22338
Nevada occurrence is from vertical rock walls (CAS, UNLV) [determined by David Toren].
with dripping water in a subalpine forest. In wet meadow and fen habitats.
Bryum calobryoides J.R. Spence [Bryaceae] Encalypta alpina Smith [Encalyptaceae]
Clark Co,: Spring Mountains, Rainbow Moun- Elko Co,: East Humboldt Wilderness, East Hum-
tain Wilderness, south fork of Pine Creek, 29 boldt Mts, Humboldt National Forest, wall of
Apr 2008, 4690 ft, Brinda2263 (CAS, UNLV); cirquejust above headwaters ofFourth Boulder
Spring Mountains, Rainbow Mountain Wil- Creek, 16 Aug 2007, 9575 ft, Shevock 30434
derness, southfork ofPine Creek, 29Apr2008, (CAS, UNLV) [determined by David Toren;
4690 ft, Brinda 2279 (MO, NY). Elko Co.: confirmed by Diana Horton]. On moist rock
along Jarbidge River at Lower Bluster Camp- ledges and terraces mixed among Blepharos-
ground, Humboldt National Forest, 25 Aug toma trichophyllum (Linnaeus) Dumortier, Dis-
2002, 6600 ft, Shevock and Glazer 22740a tichium capiiiaceum (Hedwig) Bruch & Schim-
(CAS). Small plants can occur among other per, and Piatydictyajungermannioides (Bridel)
mosses so it can be easily overlooked. The H.A. Crum, in a subalpine conifer forest. A
orbicular-shaped leaves are, nonetheless, high- highly unexpected addition to the bryoflora of
ly diagnostic. Nevada considering the distribution pattern of
widely disjunct occurrences coupled with spe-
Campyliadelphus chrysophyllus (Bridel) Chopra cific microhabitat requirements.
[Amblystegiaceae]
Lander Co.: Toiyabe Range, Toiyabe National Entosthodon bolanderi Lesquereux [Funariaceae]
Forest, headwaters ofBig Creek, 30 May2002, Clark Co.: Spring Mountains, Red Rock Canyon
8100 ft, Shevock and Glazer 22352 (CAS, National Conservation Area. South Fork Pine
UNLV) [determined by David Toren], Creek just above mouth of canyon, 4 Apr
Didiodontium olympicum Renauld & Cardot 2005, 4200 ft, ShevockandBrinda.26535 (CAS)
[determined by David Toren].
[Dicranaceae]
Elko Co.: East Humboldt Wilderness, Humboldt Entosthodon tucsoni (E.B. Bartram) Grout [Fu-
{
National Forest, slopes south of Third Boulder nariaceae]
CreektowardBirdeyeLake, 14Aug2007,9200ft, Clark Co,: Spring Mountains, Red Rock Canyon I
Shevock30342 (CAS, UNLV); East Humboldt National Conservation Area, South Fork Pine j
|
2014] BRINDA ET AL.: BRYOPHYTES NEW FOR THE SILVER STATE. PART III 255
Creek, 4 Apr2005, 4200 ft, BrindaandShevock south ofthe Muddy Mountains, 11 Mar 2007,
100 (CAS, UNLV); Spring Mountains, Red 1615 ft, Brinda and Rosentreter 1301 (UNLV)
Rock Canyon National Conservation Area, [confirmed by Ken Kellman]; South Virgin
South Fork Pine Creek, 27 Apr 2006, 4200 ft, Mountains, along Gold Butte Road east of
Brinda and Shevock 999 (UNLV); Spring Tramp Ridge, 21 Apr2007, 2935 ft, Brinda and
Mountains, Red Rock Canyon National Con- Shevock 1490 (UNLV) [confirmed by Ken
servationArea, South ForkPineCreek, 27Apr Kellman]; Virgin Mountains, WhitneyPockets,
2006, 4200 ft, Shevock and Brinda 26535a 21 Apr 2007, 3060 ft, Brinda and Shevock
(CAS) [determined by David Toren], 1510b (UNLV); Lake Mead NRA, sandstone
outcrops along the south side of Northshore
Fabronia ciliaris (Bridel) Bridel [Fabroniaceae] Road, about 1 km east ofWest End Wash, 31
Clark Co.: South Virgin Mountains, Gold Butte, 21 Mar 2013, 1730 ft, Brinda and Stark 4448
Apr2007, 3970 ft, Brinda 1477 (CAS, UNLV). (CAS, MO, NY, UNLV). This recently de-
FissMeos bryoides Hedwig [Fissidentaceae] scribed species (Kellman 2011) was known
White Pine Co,: Egan Range, Sawmill Creek only from the type locality in coastal central
California. The occurrences for Nevada are
Canyon 5.4 mi west of Cave Valley, 31 Jul
1979, 8000ft, Lavin, Mozingo andRyser M-220 tdihsetrreifbourteioan raenmdarkhaabbilteatexftoernsithoins isnpecbioetsh.
(NY) [confirmed by Ronald Pursell]. Grimmia vaginulata is not uncommon on the
Gemmabryum demaretianum (Arts) J.R. Spence red sandstone outcrops of southern Nevada.
[Bryaceae] Gymnostomum calcareum Nees & Hornschuch
Clark Co.: McCullough Mountains, pass on
[Pottiaceae]
highway 164, 16 Oct 2005, 4815 ft, Brinda Clark Co.: BLM Red Rock Canyon National
394, 397(UNLV); El Dorado Mountains, near RecreationArea. NorthForkPineCreek,4Apr
h(iUgNhLwVa)y.16E5s,m2e3raFledba 2C0o0.6:, 3W3h0i0teft,MBoruinntdaain6s4,7 2(0C0A5S,,43U0N0LftV,)She[vdoectke,rmBirnienddab,yanDdavSitadrkTo2r6e5n0];7
road to Middle Canyon, 3 Jul 2005, 7035 ft, BLM Red Rock Canyon National Recreation
Brinda 225 (UNLV). Area. North Fork Pine Creek, 4 Apr 2005,
Gemmabryum klinggraeffii (W.P. Schimper) J.R. 4300ft, Brinda, Shevock, andStark 78(UNLV).
Spence & H.P. Ramsay [Bryaceae] Elko Co.: Ruby Mountains, Humboldt Na-
Clark Co,: University of Nevada, Las Vegas, tional Forest, between Liberty Lake and
Main Campus, lawn Between Physics and Liberty Pass, 31 Jul 2004, 10,325 ft, Shevock
Health Sciences Buildings, 27 Jan 2011, 26039a (CAS) [determined by David Toren].
2030 ft, Brinda and Greenwood2615 (UNLV). Gymnostomum viridulum Bridel [Pottiaceae]
Gemmabryum violaceum (Crundwell & Nyholm) Clark Co.: Spring Mountains, Gateway Canyon,
J.R. Spence [Bryaceae] 26 Feb 2006, 3865 ft, Brinda 720 (CAS, MO,
Churchill Co.: Stillwater Range between Sham- UNLV) [determined by Richard Zander]. This
rock Canyon and Fondaway Canyon, 3 May species resembles and occupies the same
2006, 1500 m, Tonenna 38 (CAS); Stillwater microhabitats as G. calcareum but the axillary
Range between Shamrock Canyon and Fond- gemmae in G. viridulum readily distinguish this
away Canyon, 3 May 2006, 1500 m, Tonenna species.
43 (CAS). Clark Co.: Spring Mountains, HomomalMum mexicanum Cardot [Hypnaceae]
Bonanza PeakTrail, 4Jul 2005, 8140 ft, Brinda Clark Co.: Red Rock Canyon National Recrea-
296 (UNLV); Virgin Mountains, Cabin Creek, tion Area, Willow Spring Picnic Area, 2 Apr
21 Apr 2006, 4560 ft, Brinda 916a (UNLV). 2003, 4825 ft, Shevock and Stark 23970 (CAS,
Lincoln Co.: Hiko Range, highway 93 about UNLV); Red Rock Canyon National Recrea-
3 miles E ofhighway 375, 3 Apr 2005, 4050 ft, tion Area, Willow Spring Picnic Area, 2 Apr
BrindaandShevock12a (UNLV); Hiko Range, 2003, 4825 ft, Shevock andStark23972 (CAS)
highway 93 about 3 miles E ofhighway 375, 3 [confirmed by David Toren], On sandstone
Apr 2005, 4050 ft, Brinda and Shevock 15b boulders. White Pine Co.: Snake Range, Great
(UNLV). Nye Co.: Death Valley National Basin National Park, Grey Cliffs off of Baker
Park, Grapevine Mountains, Amargosa Range Creek Road toward Grey Cliffs Group camp-
above Strozzi Ranch site south of Wahguyhe ground, 27 Jul 2003, 7200 ft, Shevock, Glazer
Peak, 11 Jan 2002, 6800 ft, Shevock and York and Clifton 24343a (CAS). On lime-
21687 (CAS, UNLV). stone walls mixed with Hypnum vaucheri
Lesquereux.
Grimmia vaginulata K. Kellman [Grimmiaceae]
Clark Co.: Frenchman Mountain, below north Hygroamblystegium noterophilum (Sullivant &
summit, 30 Jan 2007, 3745 ft, Brinda 1251 Lesquereux ex Sullivant) Warnstorf [Amblys-
(UNLV); Lake Mead NRA, Northshore Drive tegiaceae]
MADRONO
256 [VoL 61
Elko Co,: Ruby Valley, 20 May 1935, 6000 ft, fV. Myurella tenerrima (Bridel) Lindberg [Pterigy-
W. Bennett s,n. (NY 714904) [confirmed by nandraceae] li
Ryszard Ochyra]. Elko Co.: East Humboldt Mountains, East
Hygrohypnum bestii (Renauld & Cardot) Broth- HFourmesbto,ldatlonWgilEdaesrtneHssu,mbHoulmdbtolHidgthliNnaetiTornaaill ji|i:j
erus [Amblystegiaceae] i
just below saddle, Fourth Boulder Creek
ElkNoatCioo,n:alEaFsotreHsut,mbsololpdets WbieltdweerennesBso,uHlduemrboLladkte (BaCsAiSn,, U16NLAVu)g [2d0e07t,erm9i6n3e5d fbt,ySDhaevviodckTo3r0e4n0].6 ||
and fens with beaver ponds above East
Humboldt Highline Trail, 15 Aug 2007, Crevice of metamorphic rock outcrops in a j|j'
9490 ft, Shevock 30389 (CAS). Esmeralda subalpine forest ofPinusflexilis.
Co,: White Mountains, Inyo National Forest, Orthotrichum pilosissimum Medina, Lara &
Morris Creek, 28 May 2002, 6800 ft, Shevock Garilleti [Orthotrichaceae] |
and Glazer 22292 (CAS). Washoe Co.: Lake Esmeralda Co.: Grapevine Mountains, Amargosa
Tahoe Basin, shaded streamlet above Incline Range, Death ValleyNational Park,just south
Village, 29 Jul 2002, 7500 ft, Shevock 22607 ofStrozzi Ranch site and Brier Spring, 14 Dec
(CAS) [determined by Dave Jamieson]. On 2001, 6400 ft, Shevock, Davis andDavis21581a j
rocks in splash zone ofstream. (CAS); White Mountains, Inyo National For- \
I
Hy[gArmobhlyysptneugmiacsetayer]iacum (Limpricht) Brotherus ePseta.k,Mo3r1riMsaCyre2e0k0,2,no6r8t0h0bfat,seShoefvoBcokun22d1ary- 'f
(CAS). Eureka Co.: Toiyabe National Forest
Elko Co.: East Humboldt Wilderness, Humboldt
along road 004, 9.4 mi east ofMonitor Valley,
NFaotuirotnhalBoFuolredsetr,Cbraeseekofofcfiroqfue,EahsetadHwuamtbeorsldotf May 2002, 7500 ft, Shevock and Glazer 22368 i|
3H0ig4h1l8in(eCATSra,ilM,O1,6NAYu,gU2N00L7,V)94[7d5etfet,rmSihneevdobcyk (HCuAmSb)o.lHdutmNbaotlidotnaClo.F:orSeasntt,aBRuofsfaaloMoCurneteaki,ns,* |j|
David Toren]. On seepy metamorphic rock Oct 2008, 5200 ft, Garilleti et al s.n. (CAS, j
walls and ledges with Primula and Saxifraga NY, VAL). Mineral Co.: Anchorite Hills,
Toiyabe National Forest, Box Canyon, 28
within a subalpine Pinusflexilis forest.
Oct 2008, 7000 ft, Lara et al s.n. (CAS, UC).
Imbribryum mildeanum (Juratzka) J.R. Spence Nye Co.: Alta Toquima Wilderness, Toiyabe j
[Bryaceae] National Forest, Pine Creek above Pine Creek ;
Humboldt Co,: Santa Rosa Range, Humboldt Campground, May 2002, 7700 ft, Shevock and |
National Forest, Lye Creek about 2 mi west Glazer 22397 (CAS). This species was recently ^
ofMartin Creek Ranger Station, 23 June 2004, described for Nevada (Medina et al. 2011). We
7915 ft, NachlingerandTiehm2640(CAS, MO, have added it here in an effort to track all taxa ,
NY, UNLV, WTU). Lander Co.: Sheep Creek new for Nevada since the published checklist j
j
Range, Plateau north of Battle Creek, 12 (Brinda et al. 2007). '|
June 2002, 6595 ft, Nachlinger and Tiehm
2446 (CAS, MO, NY, UBC, UNLV, WTU). Philonotis caespitosa Juratzka [Bartramiaceae]
White Pine Co.: Antelope Range, Lookout Elko Co,: East Humboldt Mountains, East |I
Humboldt Wilderness, Humboldt National
STpireihnmg,214041Ju(nCeAS20,02M,O6,680NYf,t, NUaBcChl,inUgeNrLaVn,d Forest, cirque below Greys Peak west ofAngel !'
WTU). Lake, 21 Jul 2002, 8800 ft, Shevock and Glazer
22582 (CAS, UC, UNLV) [determined by Dan \
Isopterygiopsis pulchella (Hedwig) Z. Iwatsuki Norris; confirmed by David Toren]. On moist |
[Hypnaceae] to wet soil, high elevations.
Elko Co.: Jarbidge Mountains, Humboldt Na-
Plagiobryoides viuosula (Cardot) J.R. Spence
tional Forest, road paralleling Jarbidge River
jaSubhneocvvtoeicoknPiwaninetdhCGFrloaexzeekCrre2Ce2ak7m,6p22g5r(AoCuuAgnS)d20[0jdu2es,tte6r7am0bi0onvefted, Chur[rBocrchykialcCleaaCneo]y.o:nStainldlwaFtoerndRaawnagyeCbaentywoene,n 3ShMaamy- ij
by David Toren], 2006, 1500-1700 m, Tonenna 28 (CAS); Still- j
water Range between Shamrock Canyon and
Jaffueliobryum raui (Austin) Theriot [Grimmia- Fondaway Canyon, 3 May2006, 1500-1700m,
i
ceae] Tonenna 37 (CAS). Lander Co.: Toiyabe
Clark Co.: Virgin Mountains, near Red Bluff Range, Toiyabe National Forest, along forest
Spring, 19 Feb 2006, 2100 ft, Brinda 645b road 002, 3.5 miles above Kingston Guard ;
(UNLV); Gold Butte Backcountry Byway Station, 30 May 2002, 30 May 2002, Shevock i
Road at Whiskey Pocket west of Whiskey and Glazer 22345 (CAS). This species has also
Pass, 21 Apr 2007, 3025 ft, Shevock andBrinda recently been documented for Death Valley i
29783 (CAS, UNLV) [determined by David National Park at Grapevine Springs in adja- i
i
Toren]. cent California and therefore likely to be
j
2014] BRINDA ET AL.: BRYOPHYTES NEW FOR THE SILVER STATE. PART III 257
discovered in adjacent Esmeralda and Clark 6250 ft, Shevock and Glazer 22678 (CAS,
counties. Plants occur in wet riparian areas or UNLV); Freighters Defeat along road 752
seeps with high salt concentrations generally north of town of Jarbidge, 26 Aug 2002,
associated with either Didymodon tophaceus 5650 ft, Shevock and Glazer 22803 (CAS, NY,
(Bridel) Lisa or Eudadium verticillatum (Hed- UNLV). Humboldt Co.: Pine Forest Range,
wig) Bruch & Schimper. BLM Winnemucca Dist., upper slopes of
Plagiobryumzierii (Dickson ex Hedwig) Lindberg aWnodoNdacChalninygoenr,2575A0u1g(C2A00S5,,U7N1L50V)ft,[Sdehteevromcik-
[Bryaceae] nations by David Toren]. Along stream cours-
Elko Co.: East Humboldt Mountains, Humboldt es. This plant can readily be separated from
National Forest, cirque above falls west of Amblystegium serpens by the presence of
Angel Lake, 3 Aug 2004, 8500 ft, Robertson paraphyllia on the stems. An earlier name for
8669 (CAS, UC); East Humboldt Wilderness,
East Humboldt Mts., base of cirque, headwa- this species is Amblystegium radicale.
ters of Fourth Boulder Creek, 16 Aug 2007, Pseudoleskeella rupestris (Berggren) Hedenas &
9475 ft, Shevock 30426 (CAS, NY, UC, Sdderstrom [Leskeaceae]
UNLV) [confirmed by Spence]. Wet soil over Clark Co.: Red Rock Canyon National Recrea-
metamorphic walls and ledges with Salix, tion Area, North Fork Pine Creek, 4Apr2005,
Primula, and Saxifraga in a subalpine Pinus 4500 ft, Shevock, Brinda and Stark 26526
fiexiiis forest. (CAS). In the Nevada checklist (Brinda et al.
Platydictya jungermannioides (Bridel) H. Crum 2007) this species was reported from Nevada
but we had not previously seen a specimen to
[Hypnaceae]
Clark Co.: Toiyabe National Forest, Spring validate its occurrence in the state.
Mountains National Recreation Area, Mary Ptychostomum pacMcum J.R. Spence & Shevock
Jane Falls, 3 Oct 2004, 9045 ft, Brinda 410 [Bryaceae]
(UNLV); Toiyabe National Forest, Mt. Washoe Co.: Sierra Nevada, Tahoe Meadows,
Charleston Wilderness, wash below Big Falls, Toiyabe National Forest, 13 Sep 2009, 8740 ft,
3 Oct 2004, 9055 ft, Brinda 414 (CAS). Elko Wishner 9512 (CAS, UC). This species was
Co.: East Humboldt Mountains, East Hum- recentlydescribed asnewto science (Spence and
boldt Wilderness, Humboldt National Forest, Shevock 2012). We have included it here in an
avalanche chute with steep rocky canyon, effort to track all taxa new for Nevada since the
Third Boulder Creek Basin, 15 Aug 2007, published checklist (Brinda et al. 2007). This
9275 ft, Shevock 30396 (CAS, MO, NY, species is likely to remain a very rare species in
UNLV); wall of cirque just above headwaters Nevada although it is widespread in fens in the
ofFourth BoulderCreek, 16Aug2007, 9575 ft, Californian portion ofthe Sierra Nevada.
Shevock 30434b (CAS). Mixed with Encalypta
alpina, Blepharostoma trichophyllum, and Dis- Splachnobryum obtusum (Bridel) C. Muller Hal.
tichium capillaceum. White Pine Co.: Snake [Splachnobryaceae]
Range, Great Basin National Park, Snake Clark Co.: Lake Mead National Recreation
Creek near Shoshone Campground, 26 Jul Area, Gold Strike Canyon Hot Springs, 4
2003, 8200 ft, Shevock, Glazer and Clifton Apr 2010, 985 ft, Brinda2613 (CAS, MO, NY,
24294a (CAS); on soil bank of creek mixed UNLV) and Webb s.n. [Mar 2010] (UNLV);
with Mnium marginatum (Dickson ex Wither- Virgin Basin, Lake Mead at water’s edge,
ing) Palisot de Beauvois; upper slopes of Blue 5 miles north-northeast of Middle Point, 22
Canyon, 27 Jul 2003, 9300 ft, Shevock, Glazer Apr 1941, Clover 6269 (NY!). In Brinda et al.
and Clifton 24318a (CAS); on soil bank of (2007) we had doubts about this taxon actually
creek mixed with Mnium marginatum; upper occurring in Nevada and it was assumed the
slopes of Blue Canyon, 27 Jul 2003, 9300 ft, record was likely to be a herbarium processing
Shevock, Glazer and Clifton 24325 (CAS, MO, error. However, with this collection weconfirm
NY, UNLV) [confirmed byDavidToren]. This this taxon for the Nevada bryoflora.
is a very small moss that can easily be
overlooked. It frequently occurs in mixed Tortella alpicola Dixon [Pottiaceae]
populations among other larger mosses and Clark Co.: Spring Mountains, north-facing Ra-
lrievceerswsoerstsp.riMmiacrriloyhaibnitsahtadiesddaplmapcess.oil and rock v2i0n0e6,a4b2o5v0efSt,oBurtihndFaor1k01o8f(PCiAnSe,CMreOe,k,U2N7LVA)p;r
Sheep Range, near the end of(dosed) Sawmill
Pseudocampylium radicale (P. Beauvois) Vander- Road, 6 Aug 2006, 7578 ft, Brinda 1213 (MO,
poorten & Hedenas [Amblystegiaceae] UNLV). Lincoln Co.: Mormon Mountains,
Elko Co.: Jarbidge Mountains, Humboldt Na- Upper South Fork Toquop Wash, 14 May
tional Forest, along forest road 748 at Seven- 2005, 6860 ft, Brinda 132 (CAS, MO, UNLV)
tysix Creek, Major Gulch, 24 Aug 2002, [confirmed by Patricia Eckel].
MADRONO
258 [Vol. 61
Warnstorfia fluitans (Hedwig) Loeske [Calliergo- publication. According to Greven (2010), this
naceae] species is related to Grimmiaarizonae. We have
Washoe Co.: Sierra Nevada, Carson Range, Big determined that the collection cited by Greven
Meadows South of Verdi, north end of (2010, and personal communication) actually
Meadows, 25 Sep 1998, 8715 ft, Tiehm 12764 was collected near Las Vegas, New Mexico
(NY) [determined by David Toren]. rather than Las Vegas, Nevada. Therefore, the
literature reference of Grimmia texicana occur-
Warnstorfia pseudostraminea (C. Muller Hal.)
Tuomikoski & T. Koponen [Calliergonaceae] ring in Nevada is erroneous and this species is
hereby excluded from the Nevada bryoflora.
White Pine Co.: Snake Range, Great Basin
National Park, Alpine Lakes Trail from Acknowledgments
Wheeler Peak trailhead to Stella Lake, 27
Aug 2005, 10,070 ft, Shevock and Giazer27619 We thank the various land management agencies
(CAS, UNLV) [determined by David Toren]. throughout Nevadaforprovidingcollectingpermitsfor
this long-term inventory project. Identifications or
Weissia ligulifolia (E.B. Bartram) Grout [Pottia- confirmations provided by Vadim Bakalin, Bill Buck,
ceae] Claudio Delgadillo, Bill Doyle, Patricia Eckel, Diana
Clark Co.: Bureau of Land Management, Las Horton, Dave Jamieson, Ken Kellman, Dan Norris,
Vegas Dist., Virgin Mountains, White Rock Ryszard Ochyra, Ronald Pursell, David Toren, and
site at mouth ofCabin Canyon, 15 Nov 2002, Richard Zander are greatly appreciated. We are also
U39N5L0Vf)t, [Sdheetveorcmkineetd alb.y2R36i3c2har(dCASZa,ndMerO],; rapepvrieecwieartsivtehaotfecnohmamnceendtsthperofivniadlevderbsyiotnw.o anonymous
‘Nay’ Canyon, 28 Feb 2006, 4720 ft, Brinda Literature Cited
583 (UNLV); near Red Bluff Spring, 19 Feb
2M0o0u6n,ta2i0n3s5,ftn,eBarrinhdiag6h1w8ay(U1N65L,V)2;3 EFleDbo2r0a0d6o, BrinSdpae,ncJ.e.C.,2L0.07R.. SAtnarka,nnJo.tRa.teSdhecvhoecckkl,isatndofJ.tRh.e
t3a3i0n0s,fGt,atBerwinadyaC6a4n9yo(nU,N2L6V)F;ebS2p0r0i6n,g3M8o65unft-, FlorhbirasytooopfrhyyNtinoesrthtoehfstNAatemev.earTdiahc,eaBwriEytdohilotngooirtsietasl11o0Cn:o6mc7o3ml-il7et0c5tt.ieneg
Brinda 715 (UNLV); Valley ofFire State Park, (eds.). 2007. Flora of North America. North of
Mouse’s Tank Area, 4 Mar 2006, 2095 ft, Mexico, Vol. 27. Mosses Part 1. Oxford University
Brinda 743B (UNLV). Lincoln Co.: Hiko Press, New York and Oxford.
Range, highway 9 about 3 mi east ofjunction Flora of North America Editorial Committee
withhighways375 and318, 3 Apr2005,4050ft, (eds.). 2014. Flora of North America. North of
Brinda and Shevock 3 (UNLV); Hiko Range, Mexico, Vol. 28. Mosses Part 2. Oxford University
highway 9 about 3 mi east of junction with Press, New York and Oxford.
highways 375 and 318, 3 Apr 2005, 4050 ft, Greven, H. 2010. Grimmia texicana sp. nov. (Grim-
Brinda and Shevock 7 (UNLV); Hiko Range, miaceae) from Texas and its separation from
highway 9 about 3 mi east of junction with Grimmia arizonae. The Bryologist 113:360-364.
hSihgehvwoacyksa3n7d5 Barnidnd3a18,26346A8pr(C2A0S0,5, U4N0L50V)f;t, KellGmriamnm,iaKc.eae20)1a1.neGwrismpemciiaesvfaogritnhuelacteant(rBalrycoopassitdao,f
California. Madrono 58:190-198.
Hiko Range, highway 9 about 3 mi east of Medina, R., F. Lara, V. Mazimpaka, J. R. She-
junction with highways 375 and 318, 3 Apr vock, and R. Garilletl 2011. Orthotrichum
2005, 4050 ft, Shevock and Brinda 26476 pilosissimum (Orthotrichaceae), a new moss from
(CAS); South Pahroc Range, Pahroc Summit, arid areas of Nevada with unique axillary hairs.
3 Apr 2005, 5000 ft, Shevock andBrinda26498 The Bryologist 114:316-324.
(CAS, NY, UNLV) [determined by David Shevock, J. R.,J. R. Spence, and L. R. Stark. 2005.
Toren]. Contributions toward a bryoflora of Nevada:
bryophytes new for the Silver State. Part I.
Madrono 52:66-71.
Excluded Species Spence, J. R., L. R. Stark, andJ. R. Shevock. 2006.
Contributions toward a bryoflora of Nevada:
Grimmia texicana Greven [Grimmiaceae]
W bryophytes new for the Silver State. Part II.
Clark Co.: 2 mi of Las Vegas. This record is Madrono 53:400M03.
based on a single herbarium specimen at FH, AND J. R. Shevock. 2012. Ptychostomum
examined and subsequentlycited as this species pacificum (Bryaceae) a new fen species from
in Greven (2010). The collector’s name and California, Oregon, and western Nevada, USA.
date of collection were omitted in the 2010 Madrono 59:156-162.