Contributions to the Knowledge of the "Staphylinus-complex" (Coleoptera, Staphylinidae, Staphylinini) of China. Part 16. The Genus Ocypus LEACH, 1819, Subgenera Ocypus s. str. and Matidus MOTSCHULSKY, 1860

2007·19.4 MB·English

by  Smetana, A

Elytra,Tokyo,35(2):441-459,November3,2007
Contributionsto the Knowledgeof the“Staphylinus-complex”
(Coleoptera,Staphylinidae,Staphylinini) of China
Part i6. TheGenus coypusLEACH, 1819,Subgeneracoypuss.str.
andMatidus MoTscHULsKY, 1860
AlesSMETANA
AgricultureandAgri-FoodCanada,Biodiversity,Cent1-alExperimentalFarm,
K.W.NeatbyBldg.,Ottawa,OntarioKIAOC6,Canada
Abstract The paper dealswith the species of the subgenera coypus and
Matidusofthegenuscoypusknowntooccuratpresentintheterritoryof thePeople's
RepublicofChina. Fourspecies,allnew,aretreatedinthesubgenuscoypuss.str:0.
umbro (Sichuan), 0. rhoetus (Sichuan:JiajinShan), 0.zopyrus (Sichuan: Gongga
Shan)and0.thericles(Sichuan:Jintiang). coypuscoreanusJ.MOLLER,1925and0.
nitensnitens (ScHRANK, 1781) are recorded for the first time from thePeople's
RepublicofChina. Eachspeciesisdescribed,illustratedandallavailabledistributional
andbionomicdataaregiven. AkeytotheChinesespeciesof thesubgenus0、〔ypuss.
str.isgiven.
Keywords: Coleoptera,Staphylinidae,Staphylininae, coypus,subgeneraOypus
s.str andMatidus,Palaearctic,mainlandChina, taxonomy,newspecies,distribution.
Introduction
This isthesixteenthofaseriesofpapersdealingwiththegeneraof the“St11phy-
1inus-complex”(seeSMETANA&DAVIES,2000)ofthePeople'sRepublicofChina. It
treats the fewspeciesof thepredominantlywest Palaearcticsubgenera coypuss.str.
(fourspecies)andMatidusMoTscHULsKY,1860(twospecies).
Bothsubgeneraare treated in therevised concept presentedbySMETANAand
DAVIES,2000.
Material and Methods
In thispaper, allspeciesof thesubgenera coypuss.str and Matiduspresently
known to occur in the territory of thePeople'sRepublicof Chinaare treated. The
specimensthispaperisbaseduponcomefromseveralsourcesandthematerial,although
notverynumerous (150specimens) isbelieved,asthatofpreviouspapers,toinclude
mostofthespecimensrecentlycollectedinChina. Therelativepaucityofthematerial
isduetothefactthatthetwosubgeneracontainmainlywestPalaearcticspeciesandare
representedinmainlandChinaonlybyafewspecies.Somespecimenswerecollectedby
442 AlesSMETANA
theauthor,othersweretakenbyEuropeanandJapanesecoleopterists, thenamesof
whichappear in“Typematerial”or“Additionalmaterialstudied”undereachspecies.
Thematerial isdeposited invariouscollections, both institutionalandprivate. The
ho1otypes that aresaid tobehousedin thecollectionoftheauthor will beeventually
depositedintheMuseumd'Histoirenature11e,Geneve,Switzerland.
Noneofthefourspeciesofthesubgenuscoypuswerepreviouslydescribed. Onthe
other hand,bothspeciesofthesubgenusMatiduswerepreviouslydescribed.
Most featureson theaedoeagusaredescribedasseeninventralview. Theventral
viewoftheaedoeagusisinterpretedasthefaceoftheaedoeagustowhichtheparamere
isattached. Asinpreviouspapers, it isabsolutelynecessarytoviewtheaedoeagusin
straightventralview,inwhichthebasalbulbus,andparticularlythebasalportionofthe
paramere,areexactlyhorizontal. Ifthebaseoftheparamereisslantedtoeitherside,the
shapesoftheapicalportionsofboththemedianlobeandtheparamerewillbepossibly
quitedifferent fromthewaytheyaredescribedandillustrated. Whendescribingthe
aedoeaga1featuresandreferringtotheleftorrightside,itistheleftorrightsideasthey
appear intheaccompanyingillustrationtheuser isreferredto.
Thekeytospecieswasconstructedhavingastrictlypractical purposeinmind,
thereforetheappearanceof thespeciesinthekey isat random. Since the key relies
heavily onthecharactersof theaedoeagus, it isrecommendedthat theaedoeagus is
routinely removedandmountedwith thespecimens.
Thesymbolsusedinthetext,whenreferringtothedepositionsofspecimens,areas
follows:
APC CollectionofAndreasPuTz,Eisenhiittenstadt,Germany
ASC Collection of AlesSMETANA, Ottawa,Canada
FMNH CollectionoftheFieldMuseumofNaturalHistory,Chicago,Illinois
MSC CollectionofMichaelScHOLKE, Berlin,Germany
NMW CollectionoftheNaturhistorischesMuseum,Wien,Austria
YSC CollectionofYasutoshiSHIBATA,Tokyo,Japan
Thenumber of paratypes is given for each locality behind the locality data,
followedbytheacronymofthecollectioninwhichtheparatype(s) is(are) deposited.
Alldataarepresentedinfullforho1otypesandallotypes. Inpreviouslydescribedspecies
thenumber ofspecimens (unsexed) isgiveninbracketsbehindlocalitydata, followed
againbytheacronymofthecollection inwhichthespecimensaredeposited.
Taxonomic Part
Genus coypusLEACH, l819
Subgenus coypusLEACH, 1819
coypusLEACH,1819,172
Contributionsto theKnowledgeofthe“Staphylinus-compIex,”16 443
GoeriusWESTWOOD, 1827,58.
IsopterumGIsTEL,1856,388.
NudabemusCoIFFAIT,1982,74.
TypespeciesofOypus:Staphy11ntlscya11etlsPAYKULL,1789(=Staphy1111tlsophthalmlcusScoPoLI,1763).
TypespeciesofGoerius:Staphy11n1lsolens0.MJtLLER,1764.
Typespeciesof isopterum:Staphy11nuscyane1ls PAYKULL, 1789 (=Staphy11nusophtha1'nlcusScoPoLI,
1763).
TypespeciesofNudabemils:Nudabe1nuscaerulescensCoIFFAIT,1982.
Des〔:riptivenotes. Thebasictaxonomicinformationonthesubgenuscoypuss.str.
_
wasprovidedbySMETANA&DAVIES(2000), thereforeonlyadditional informationis
presentedhere. XanthoypusJ.MULLER,1925, listedasajuniorsynonymof 0〔:ypusin
theabovepaper,isremovedfromthissynonymyandtentativelysetasideasapossible
junior synonymof Age1osus SHARP, 1889, pendingaconfirmation when theChinese
speciesofAge1osusarerevised.
Thesubgenericcharactersonmaxillaryand labial palpi weregiven in theabove
paper (SMETANA&DAVIES,2000),butitshouldbementionedherethattheshapeof
thelastsegment of themaxillarypalpusvariestosomeextent, frombeingelongate-
fusiformtoshort-fusiformwithmoreorlesstruncateapex.
Thepubescenceoftheabdominaltergitesisuniformlydark,piceous-blacktoblack
inall Chinesespecies.
Theshapeofsternite9ofthemalegenitalsegmentissimilarinallChinesespecies,
thebasalportionisnarrowandnomorethanmoderatelylong,apicalportionisvariably
emarginatemedioapica11y(Figs.1,10,14). Tergite10ofmalegenitalsegment issimple,
ofvariablytriangularshape,withsimplesetation(Figs 2,6). Theaedoeagusislarge
and elongate in all Chinesespecies, with both median lobe and paramere variably
asymmetrical. Theparamereissituatedonmedianlobemoreorlessasymmetrically,the
undersideofparamerebearsblacksensorypegsetae,numberofwhichvariesconsidera-
bly (Figs 4,8,13,17).
Thetergite10of femalegenital segment of theChinesespecies (femaleofone
speciesnot known)isingeneralshort,markedlynarrowedtowardsubacuteapex(Figs.
18,23),ormoderatelynarrowedtowardsubtruncateapex (Fig 9). Theapicalportion
of thetergiteisstronglysclerotizedinallspecies.
AllChinesespeciesofthissubgenusareof largesize,somebeingamongthelargest
Chinesemembersof thefamilyStaphylinidae.
Key toChineseSpeciesofSubgenus coypus
Legsuniformlyrufobrunneous. Apicalportionsofbothmedianlobeandparamere
in situ (ventral view) of characteristic, broadly lanzet-1ikeshape (Fig 3).
AedoeagusasinFigs3,4. Length19.0-23.0mm.CentralandnorthernSichuan.
0. umbrosp nov
- Legspredominantlydark.Apicalportionsofbothmedianlobeandparamereinsitu
444 AlesSMETANA
(ventralview)ofdifferentshapes 2
2 Sensorypegsetaeonundersideofparameremoderatelynumerous,situatedmostly
nearapexofparamere(Fig. l3).AedoeagusasinFigs.11-13.Length18.0-22.0
mm.Sichuan:GonggaShan 0.zopyrussp
nov
Sensory pegsetae on underside of paramere numerous, extended considerably
posteriadalongeach lateralmarginofparamere(Figs.8,17)
3 Paramereelongate,narrow,withnarrowlyarcuateapex(Fig.17),apicalportionof
median lobelong,narrow,withsubacuteapex (Fig. 16). Aedoeagusasin Figs.
15-17. Length20.0-24.0mm. Sichuan:Jintiang.・・・・・・・・・・0. thericlessp nov.
Parameremoderately longandwide,withobtuseapex (Fig 8), apicalportionof
medianlobeshort,wide,withsubacuteapex.AedoeagusasinFigs 7,8.Length
17.0-23.0mm.Sichuan:ErlangShan 0. rhoetussp nov
DescriptionsofSpecies
coypus(coypus)umbrosp
nov
(Figs.1-5)
Diagnosis. Largespecieswith rufobrunneouslegs,andwithquitecharacteristi-
callyshapedaedoeagus.
_Description. Piceous-blacktoblack,moderatelydull,apexofabdomenindistinctly
paler in somespecimens; maxillary and labial palpi rufobrunneous, antennaedark
brunneous to brunneopiceouswith first segment usually paler; legs rufobrunneous;
pubescenceofdorsalsideofbodyuniformlypiceous. Headof roundedquadrangular
shape, with roundedposterior angles, wider than long (ratio 127), eyes small, flat,
temperaconsiderablylongerthaneyesfromabove(ratio2.77),dorsalsurfaceofhead
finelyanddenselypunctateandpubescent,interspacesbetweenpuncturesondiscabout
aslargeasdiametersof punctures, punctationgraduallybecomingslightlydenserand
coarsertowardposteriorandlateralmargins;noimpunctatemidlinepresent;interspaces
between punctures with relatively coarsesubmeshedmicrosculpture. Gular sutures
narrowlyseparated;postgenaefinelyandratherdenselypunctate, interspacesbetween
puncturesnear gularsutureslarger thandiametersofpunctures. Dorsalsideofneck
withpunctationsomewhatfinerthanthatonhead. Antennamoderatelylong,segment
3 longer thansegment 2 (ratio128),segments4 to7longer thanwide, becoming
graduallyshorter,outersegmentsaboutaslongaswidetovaguelywider than long, last
segmentshorter than twoprecedingsegmentscombined. Pronotum longer thanwide
(ratiosbelow1.13), almost parallel-sided,narrowmarginalgroovedisappearingdown-
Figs.1-8. - 1-5.coypusumbro:1,apicalportionofsternite9ofmalegenitalsegment;2,apical
portionof tergite10ofmalegenitalsegment;3,aedoeagus,ventralview;4,undersideofapical
portionofparamere;5, tergite10of femalegenital segment. - 6-8. 0ypusrhoetus:6, apical
portionof tergite10ofmalegenitalsegment;7,aedoeagus,ventralview;8,undersideofapical
portionofparamere.
ContributionstotheKnowledgeof the“Staphylinus-complex,”16 445
446 AlesSMETANA
wardsat about middleof pronota11ength; disc with fine, entire impunctatemidline;
punctationondiscindistinctlyfinerandlessdensethanthatondiscofhead,pubescence
andmicrosculptureoninterspacesbetweenpuncturessimilartothatonhead. Pronota1
hypomeronwithafewmicrosetae. Scutellumfinelyanddenselypunctateandsetoseon
entiresurface, surfacewithvery fine, rudimentarysubmeshedmicrosculpture. Elytra
short,slightlydepressedatbase,vaguelydilatedposteriad,atsutureconsiderably(ratio
0.58),atsidesdistinctly(ratio0.78)shorterthanpronotumatmidline;punctationvery
fineanddense,difficult toobserveamongdensegranulosemicrosculpture. Wingseach
reduced tominute, non-functionalstump. Abdomen with fifthvisibletergitelacking
paleapicalseamof palisadesetae; tergite2 (in front of first visibletergite) entirely,
densely and finely punctate and pubescent; all tergites evenly, finely and densely
punctate, punctation gradually becomingslightlysparser towardapex of abdomen;
interspaceswithveryfine,densesubmeshedmicrosculpture.
M ale. Sternite8withrathernarrowandshallow,obtuselytriangularmedioapi-
ca1emargination. Genitalsegmentwithsternite9withnarrow,moderatelylongbasal
portion,apicalportionwithdistinct,obtuselytriangularmedioapica1emargination(Fig.
1). Tergite10rathernarrow,evenlynarrowedtowardnarrowlyarcuateapex,setoseas
in Fig 2. Aedoeaguslarge,ofcharacteristicshape(Figs 3,4);median lobegradually
dilated intowide,triangularapical portionwithsubacuteapex; paramerelarge,mark-
edly dilated into widelanzet-shapedapicalportion, withshort longitudinalcarina
on
faceawayfrommedianlobe(Fig 3), apexofparameredistinctlynot reachingapexof
median lobe (Fig 3);sensory pegsetaeonundersideofparamerequitenumerous,
forming two dense lateral groups joined at apex of paramere; apical setae minute,
situatedat left lateralmargin(undersideup) belowapexofparamere(Fig 4).
Fema1e. Tergite10of genitalsegment wide,markedlynarrowed towardsub-
acuteapex (Fig 5).
Length19.0-23.0mm.
Typematerial. Holotype (male): China:“CHINAC-SichuanV2004 Kangding
cityenv. bushes & fields, 2600-2800mM. Hacke1&R.Sehna1” Allotype(female):
China:“Chinac.,Sichuanocc. Kanding30.6.-8.7.1999Dr.Vlad.Boneslegit” Holo-
typeandallotypein theSMETANAcollection,Ottawa.
Paratypes: [Sichuan]:samedataasholotype, 1 (MSC);samedataasallotype,
5早早(ASC); Kangdingenv., 2500-3000m, 6.-9.VII i995 30°05'N 101-55'E M.
Tryzna& 0.Safranek lgt.,3 ,2早早(ASC); (Sichuan)Umg.Kangding10.-12.VI.
l995, 2800m, Heinz leg., 3 ,2早早(ASC);WestSichuan,Kanding,12.-21.7.94,
2600m, Benes lgt., l (ASC); W-Sichuan, Kangdingenv., 2691m, 30°02.502'N
101°57.264'E,14.VI2006,R.Sehna1&M.Tryzna,21 ,12早早(ASC,MSC);Umg.
Kangding, 3000-3100m,21.-22.VII i994,Heinzleg., 1 (ASC); Kanding, Mogecuo
NP, 3000m, 20.VII i998, leg. S. Murzin, 3 (MSC); W-Sichuan, Sa'doenv., V.
2004,primaryforest,3540m,29°36,4'N101°22,9'c leg.Hackel&Sehna1,1 (MSC);
SBarkambetweenLianghekou-Fubian,3459-3650m,Quercusshrubsandmixedwood,
10.-30.VI 2004,1gt.R. Fabbri,2 ,2早早(ASC,MSC);Sichuan,Ganzipref., Daxue
ContributionstotheKnowledgeof the“Staphy1l川Is-complex,”16 447
ShanN Kangding, 101.57E30.03N,22/24.V.1997,1 log.A. Piitz (APC).
Additionalthreespecimensinbadshapewerenotlabelledasparatypes.
Geographicaldistribution. Oypusumbroisatpresentknown fromlocalitiesinand
aroundDaxueShan,southtoSabdearea,aswellasfromareaaroundBarkamincentral
andnorthernSichuan. MostavailablespecimenscomefromthevicinityofKangding.
Bionomics. Littleisknownaboutthehabitatrequirementsofthespecies;however,
itseemstofrequent bothforestandthemoreorlessopenhabitatsofshrubsandfields.
Specimenswerecollectedatelevationsfromabout 2,500m to3,650m.
Recognition. Theshapeoftheaedoeagusof0.umbroisquitecharacteristicand
allows immediate recognition of thespecies. Externally 0. umbro differs from the
remainingthreespeciesofthesubgenusby therufobrunneouslegs.
Etymology. Thespecificepithet isthenameofUmbro, -onis,m, thepriestand
leaderoftheMarses, inapposition.
coypus(coypus) rhoetussp
nov
(Figs 6-9)
Diagnosis. coypusrhoetussharesmost ofthecharacterstateswith0. umbro, but
differs from it by theentirely differentlyshaped aedoeagus, and by a few external
characters,asgiven in thedescription.
Description. In all character statessimilar to 0. umbro, but different as follows:
appendagesdarker,maxillaryandlabialpalpioftendarkbrunneoustobrunneopiceous,
antennaepiceous,becominggraduallymoreorlesspalertowardapex; legspiceous,with
dorsal faceof fronttibiaeand front tarsipaler, rufobrunneous.
M ale. Medioapica1emargination of sternite 8 somewhat wider and deeper.
Genitalsegmentwithsternite9similar,butmedioapica1emarginationslightlydeeper;
tergite l0similar tothat of 0. umbro,butwithapexslightlydifferentiated,setoseasin
Fig6. Aedoeagus(Figs 7,8) large,elongate;medianlobesubpara11e1-sidedinmiddle
portion, narrowed intoslightlyasymmetrical apicalportionwithsubacuteapex;para-
merelongandwide,asymmetrical,situatedonmedianlobeasymmetrically,withlong
longitudinal carinaon faceawayfrommedianlobe;undersideofparamerewithvery
numerous sensory pegsetae, forming wide lateral group alongeach lateral margin,
joinedbelowapexofparamere;sensorypegsetaeminute,situatedasinFig 8.
Fem a1e. Tergite10 of genitalsegment similar to that of 0. umbro, but less
markedlynarrowed towardsubtruncateapexwithmiddleportion narrowly different-
iated (Fig 9).
Length17.0-23.0mm.
Typematerial. Holotype(male) and allotype (female): China: “Ch. C-Sichuan
(LUDING) ERLANGSHANMt. R., W-slope29:51:24N/102:l5:24E-2850m
mixed forest/meadow, 19.VII 2000K &B.Brezina, M. Brychcin, M. Hacke1. In the
SMETANAcollection,Ottawa.
Paratypes: [Sichuan]:samedataasholotype, 8 ,6早早(ASC,MSC);Sichuan
448 AlesSMETANA
ErlangShanPass3200mVII2000M.Hacke11eg.,4 (ASC);C-SichuanJiajinShan
3400m30°22.45'N102°l6.644'El5.VI2002M.Janataleg.,2 (ASC);W.Sichuan
(GanziTibet.Aut.Pref.,LudingCo.),WErlangShanPass,2600m,7kmSSELuding
29°51'N/102°15'E,20.-29.VI.1999D.W.Wrase,2 ,1早(MSC);samebutKiefer,
Hase1Bliiten, 21.VI.1999, leg. M.Schiilke, 1 (MSC);Ya'anPrefecture, Tianquan
Co., WErlangShanPass,2780m,29.51.27N102.15.47E,21. or 29.VI.1999, leg.A.
Piitz,sifted, 5 ,1早(APC,ASC).
Geographicaldistribut1or1. OypusrhoetusisknownonlyfromJiajinShan,particu-
larlyfromtheErlangShansectionnear Luding. ItmaybeendemictoJiajinShan.
Bionomics. Thespecimensof theoriginalseriescomefrompitfall traps,someof
whichwereset in“mixed forest/meadow” formation. Specimenscomefromelevations
between 2,600-3,200m.
Recognition. 0〔、ypus rhoetusmaybeeasilydistinguished from0. umbro by the
darkappendages,andbytheshapeof theaedoeagus. Itmaybedistinguishedfrom0.
zopyrusonlybythedifferentshapeoftheaedoeagus.
Etymology. Thespecificepithet isthenameofRhoetus,-i,m,akingofMarsians,
in apposition.
0〔、ypus (coypus)zopyrussp nov
(Figs.10-13)
Diagnosis. O◆cypuszopyrussharesmostofthecharacterstateswithboth 0. umbro
and0. rhoetus,butdiffersfrombothbytheshapeoftheaedoeagus,andfrom0.umbro
by thedarkercoloredappendages.
I)escriptiorz. Inallcharacterstatessimilar to0. umbro anddifferentonlybythe
differentlyshapedaedoeagusandthedarkercoloredappendages.
M ale. Medioapica1emarginationofsternite8not appreciably different from
that of 0. umbro. Genital segment with sternite9similar, but with medioapica1
emargination wider and deeper (Fig. 10). Tergite 10similar to that of 0. umbro.
Aedoeagus(Figs.11-13) large,elongate;medianlobenarrowedintoslightlyasymmet-
rical apicalportionwithsomewhat differentiated,knob-likeapex;paramere1ongand
wide,asymmetrical,situatedonmedianlobeasymmetrically,with long,slightlyarcuate
longitudinalcarinaon faceawayfrommedianlobe;sensorypegsetaeonundersideof
paramere moderately numerous, forming a solid, small apical field below apex of
paramere,extendedmoderatelyposteriadalongeachlateralmargin;apicalsetaeminute,
Figs 9-17. - 9.0 ypusrhoet1ls:tergite10offemalegenitalsegment. - 10-14. 0 ypuszopyrils:
10,apicalportionofsternite9ofmalegenitalsegment;11,aedoeagus,ventral view;12,apical
portionofmedian lobeof aedoeagus,lateral view; 13, undersideofapical portionof paramere.
- 14-17. coypus therieles: 14, apical portion ofsternite9 of male genital segment; 15,
aedoeagus, ventral view; 16, apical portion of median lobe of aedoeagus, ventral view; 17,
undersideofapicalportionofparamere.
ContributionstotheKnowledgeofthe“Staphylinus-complex,”16 449
10
14
11 16 17
450 AlesSMETANA
situatedasin Fig. l3.
Fem aleunknown.
Length18.0-22.0mm.
Type material. Holotype (male): China: “CHINA Sichuan, Gongga Shan,
Hailuogou,aboveCamp3,3,000m6.VII9629°35N102°00EC53”/“collectedbyA.
Smetana,J.FarkacandJ. Kabatek”. IntheSMETANAcollection,Ottawa.
Paratypes: [Sichuan]: W Sichuan 3-6.VII i994, 29.35N 102.00E 2900-3200m
Gonggashan-HAILUOGOU, 1gt. D. Kral & J. Farkac, 2 (ASC); C-Sichuan
(Kangding)GONGGASHANmassive,3100m,upperHAILUOGOUvail. (Camp3)
29°35'N/101°59'E 26.V.-6.VI.1993, B. Brezina, 1 (ASC); Sichuan (LUDING)
GONGGASHAN,2700-3200m,upperHAILUOGOUvalley29:35N/101:59E,con
forest 4.VII i994 K. & B. Brezina, 1 (ASC);GonggaShan, Hailuogou, in front of
Glacier 1, 2850m, 29°35N 102°00E, 7. VII i998, A.Smetana [C76], 1 (ASC);
Sichuanc. LudingCo.,Moxienv.,22.5-10.6.1993,Dr.Vlad. Beneslegit,2 (ASC,
MSC).
Geographicaldistribution. OcypuszopyrusisknownonlyfromtheGonggaShan;it
islikelyendemictothisrange(butseeunderRecognitionandcomments).
Bionomics. Theholotypewastakenfromunderapieceofrottingwoodonthefloor
of an original Ables forest with rich undergrowth of rhododendrons and various
deciduousshrubs. Theparatypebearingthecode[C76] wastakenrunningonthefloor
ofasimilar forest. Infact,mostspecimensweretakenintheconiferousforestaround
andaboveCamp3.
Recognitionandcomments. Ocypuszopyrusmaybedistinguished from0. rhoetus,
inaddition to thegeographical isolation, onlybythedifferentshapeof theaedoeagus.
Ihaveseenonemalespecimenof0.zopyruswithlocalitylabel“China,Sichuanocc.
Emei-Shan,10.7.1993LeshanCo.,1500mDr.M.Hacke11egit”(ASC). Iconsider the
occurrenceof 0.zopyrusonEmeiShanunlikely,unlessit isconfirmedbycollectionof
additionalspecimens. Thespecimenwasnot designatedasaparatype.
Etymology. Thespecific epithet is the name of Zopyrus, -i, m, a celebrated
physiognomist, inapposition.
coypus(coypus)thericlessp nov
(Figs.14-18)
Diagnosis. coypusthericlessharesmostof thecharacterstateswith 0. umbro,but
differs from it by the entirely differently shaped aedoeagus, and by a few external
Figs. 18-24. - 18. coypus thericles: tergite10of femalegenital segment. - 19-23. coypus
coreanus: 19, apical portionofsternite9ofmalegenitalsegment; 20,apical portionoftergite10
of malegenital segment; 21,aedoeagus,ventral view;22, apicalportionof aedoeagus, lateral
view;23, tergite10of femalegenital segment. - 24. 0ypus nitens nltens: sternite9ofmale
genitalsegment.